Coprophanaeus, Olsoufieff, 1924

Edmonds, W. D. & Zidek, J., 2010, A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae), Insecta Mundi 2010 (129), pp. 1-111 : 89-91

Coprophanaeus (C.) ignecinctus ( Felsche, 1909)

Fig. 232, 235 , 238 , 267-272

Phanaeus ignecinctus Felsche, 1909: 755

Coprophanaeus ignecinctus (Felsche) (recomb. by Blackwelder 1944: 209)

Phanaeus telamon ignecinctus Felsche (stat. by Pereira and Martínez 1956: 232)

Type. Lectotype male, here designated, Staatliches Museum für Tierkunde, Dresden.

Diagnosis. General – Metallic coloration ( Fig. 267-271 ), where present, bright red (rarely with green highlights, Fig. 269 ). Clypeus strongly and abruptly angulate lateral to median teeth. Posterior (ventral) surface of protibia completely granulorugose (as in Fig. 231 ). Cap of anterior angle of metasternum deeply bifurcate. Anterior surface of metasternum completely densely punctured. Metasternal disk finely punctured, bearing compound row of larger punctures along anterior margin. Basal pronotal fossae distinct. Elytral interstriae 2-5 raised midlongitudinally ( Fig. 232 ). Length 20-25 mm.

Male – Sides of base of head horn usually slightly convergent, rarely slightly divergent; anterior surface weakly concave, lateral angles acutely tuberculate; central process about same length as base, almost upright and angularly produced on posterior surface (as in Fig. 220 ). Pronotal prominence ( Fig. 267-269 ) with rounded lateral angles, cleat-shaped, flanked by deep oval concavities; width equal to distance between outer margins of eyes.

Female – Cephalic carina trituberculate; length of clypeus greater than that of frons.

Specimen examined – 87.

Distribution. Yungas province in Bolivia and Peru ( Fig. 238 ).

Collection Records. BOLIVIA: Cochabamba – Prov. Chaparé, 17 o 10.47’S 65 o 46.66’W, 1550 m (Feb, Nov) ; Yungas del Palmar , 1200 m (Jan-Feb) ; Limbo, 2000 m (Jan); 109 km E Cochabamba , 17 o 08’52”S 65 o 42’54”W, 1400 m (Feb) ; 124 km E Cochabamba [at Río Espírito Santo ], 17 o 03’45”S 68 o 38’38”W, 700 m (Feb) ; road from Cochabamba to Villa Tunari, 17 o 08’50”S 65 o 42’29”W, 1480 m (Feb) ; Cordillera Mostenes, Isiboro-Securé National Park [Laguna Carachupa], 16 o 13’58”S 66 o 24’54”W, 1310 m (Sep) ; Carmen Pampa, 16 o 37’06”S 66 o 28’50”W, 1820 m. Santa Cruz – near Bermejo [Refugio Los Volcanes], 18 o 06’50”S 63 o 36’15”W, 1080 m . PERU: Cusco – San Pedro, 13 o 03’15”S 71 o 32’54”W ; Megatoni National Sanctuary, 12 o 11’14”S 72 o 28’14”W, 1350-1900 m (May); La Convención, Echarate, 12 o 40’14"S 73 o 10’21"W, 1722 m (Oct). Junin – Puerto Eden [ Río Tambo valley ], 1400 m (Mar-Apr) ; Calabaza, 2000 m (Sep); near Satipo, 1200 m (Dec).

Comments. Olsoufieff (1924) was unable to see specimens of Felsche’s species, which he tentatively regarded as a variety of C. telamon . Janssens (1940) countered Olsoufieff “… cette espèce est parfaitemente valable …”, but was himself countered by Pereira and Martínez (1956; see also 1960), who combined it along with C. corythus as a subspecies of C. telamon . We agree with Janssens and with Arnaud (2002c) that Felsche’s taxon merits species recognition.

The form of the male horn and pronotum of C. ignecinctus is very similar to that of C. telamon (q.v.). The central projection of the horn is strongly bent anteriorly, and the angle of bend is angularly produced but never bituberculate.

Felsche’s (1909) description of this species is brief and includes no mention of the number of specimens he had on hand. The Felsche collection holdings in Dresden include six C. ignecinctus , only two (a pair) of which bear labels specifying the type locality, Marcapata, Peru. These two we consider syntypes, and it is the male which we hereby designate the lectotype. It is a small, worn individual missing the protarsal spurs as well as the left mesotarsus and both metatarsi. It bears five labels: a) handwritten in black on white “Marcapata, Peru ” ; b) printed on green “Coll. C. Felsche, Kauf 20, 1918”; c) handwritten “ Ph. ignecinctus [male symbol] [unintelligible]”; d) printed on white “Staatsl. Museum für Tierkunde Dresden”; and e) our lectotype designation printed on white with red margin.

Figures

Figure 238. Approximate geographic distribution of Coprophanaeus (s. str.) species in the dardanus species group B.

Figure 228-236. Characters of the Coprophanaeus (s. str.) dardanus species group. 228) Oblique lateral view of metasternum (asterisk marks capped anterior angle of shield), C. degallieri. 229) Same, C. parvulus. 230) Ventral view of left protibia of C. degallieri (arrow marks row of coarse punctures). 231) Same, C. telamon. 232) Dorsal view surface of left elytron of C. ignecinctus. 233) Same, C. milon. 234) Same, C. magnoi. 235) Dorsal view of head of female C. ignecinctus. 236) Same, C. magnoi.

Figure 267-272. Coprophanaeus ignecinctus. 267-269) Male habitus. 270) Aedeagus (dorsal view below; lateral view above). 271-272) Female habitus.

Figure 213-227. Characters of the Coprophanaeus (s. str.) dardanus species group. 213-214) C. dardanus, anterior view of cephalic process of large male. 215) Same, small male. 216) Same, female. 217) C. telamon, aedeagus (lateral view above; dorsal view below). 218-219) C. corythus, anterior view of male cephalic process. 220) C. telamon, same. 221-222) C. telamon, lateral view of male cephalic process (arrow in 222 indicates posterior tubercle). 223) C. corythus, lateral view of male cephalic carina. 224) C. degallieri, metasternal shield. 225-226) C. milon, anterior view of male cephalic process. 227) C. magnoi, same.