Cnesterodon raddai Meyer & Etzel, 2001

Lucinda, Paulo H. F., 2005, Systematics of the genus Cnesterodon Garman, 1895 (Cyprinodontiformes: Poeciliidae: Poeciliinae), Neotropical Ichthyology 3 (2), pp. 259-270 : 263

Cnesterodon raddai Meyer & Etzel, 2001 ( Fig. 4 )

Cnesterodon raddai Meyer & Etzel, 2001: 248 , fig. 1 (holotype, male); 2 (paratype, female); 3 (gonopodium) & 4 (aquarium specimen, male). Type - locality: swamp near Resistencia, Rio Paraná basin, Argentina.

Diagnosis. Cnesterodon raddai is diagnosed by the following autapomorphies: (1) the small number (one to four) of patches of dark pigmentation along flanks; (2) minute size of adults; (3) predorsal line absent; (4) three pelvic-fin rays in males [33-3]. Cnesterodon raddai further differs from its congeners by: (1) pleural ribs in adult males not curved forward converging to the same point towards pelvic girdle [59-0]; and (2) anterior and posterior orbital bony plates present [143- 1].

In addition, C. raddai differs from its congeners, but C. omorgmatos , by the circular or irregular dark brown blotches along body sides. Cnesterodon raddai is distinguished from C. omorgmatos by the general size of adults (14.6-17.0 mm [adult males], 17.1-23.2 mm [adult females] vs. 18.7-24.8 mm [adult males], 20.0-30.5 mm [adult females], respectively) and by its color pattern (one to four clearly rounded dark spots or wide and short bars along flanks vs. six to nine dark brown circular to irregular blotches or vertical wide bars along body sides, respectively).

Distribution. Cnesterodon raddai is distributed in the rio Paraguay and lower rio Paraná drainages.

Cnesterodon iguape + C. hypselurus + C. septentrionalis + C. brevirostratus + C. carnegiei + C. omorgmatos Clade [103]

Diagnosis. Members of this clade share the following uniquely derived and unreversed feature: constriction of unpaired appendix of gonopodium [103-1**].

Additionally, this clade is diagnosed by the following not uniquely derived and/or reversed features: (1) medial surface of ascending process of premaxilla approximately straight [11- 0]; (2) teeth on fourth ceratobranchial [27-0]; (3) first proximal radial of dorsal fin located between neural spines of 13 th and 14 th vertebrae in adult males [62-1].

Figures

Fig. 4. Cnesterodon raddai. UMMZ 207503, small stream at bridge on dirt road to Ayolas, Misiones, Paraguay: (a) male, 16.9 mmSL; (b) female, 23.2mm SL.