Acanthocercus cyanocephalus ( Falk, 1925 ) Angolan Tree

Marques, Mariana P., Ceríaco, Luis M. P., Blackburn, David C. & Bauer, Aaron M., 2018, Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola Atlas of Historical and Bibliographic Records (1840 - 2017), Proceedings of the California Academy of Sciences 65, pp. 1-501 : 284-285

Acanthocercus cyanocephalus ( Falk, 1925) Angolan Tree Agama

Agama cyanocephala Falk 1925:83 . Neotype: ZFMK 88492 (collector P. Wagner) designated by Wagner et al. (2018:27) . Neotype locality: “garden in Ikelenge (-11.241592, 24.273256), northern Mwinilunga District,” Zambia. Original type locality: “ Angola ” without further precision .

Stellio nigricollis : Bocage (1866a:43).

Stellio atricollis : Bocage (1879b:95, 1895a:22), Peters (1881:147), Boulenger (1885:358, 1905:110), Ferreira (1900a:49), Angel (1923:158), Schmidt (1933:9), Parker (1936:132), Monard (1937b:60), Themido (1941:7).

Agama atricollis : Boulenger (1885:356, 1905:110), Laurent (1950a:12, 1964a:38), Ferreira (1903:15), Schmidt (1933:9), Monard (1937b:58), Parker (1936:132), Themido (1941:7), Frade (1963: 253), Manaças (1963:228).

Agama atricollis atricollis : Klausewitz (1957:161).

Agama cyanogaster : Loveridge (1957:195).

Acanthocercus atricollis : Spawls (2010).

Acanthocercus cyanocephalus : Ceríaco et al. (2014b:670), Ceríaco et al. (2016b:75), Wagner et al. (2018:27) .

Acanthocercus cyanogaster : Branch and Conradie (2015:200).

Global conservation status (IUCN): Least Concern.

Global distribution: The species is known from the southern Democratic Republic of Congo, through Angola and western Zambia to far northern Namibia.

Occurrences in Angola (Map 245): The species occurs in the entire country with exception of the southwest and the northwestern regions.

Malanje: “Bange N’gola” [-8.43333,

16.56667] ( Boulenger 1905:110); “Duque de

Bragança ” [-9.10000, 15.95000] ( Bocage

1866a:43, 1895a:22; Boulenger 1885:359,

1905:110; Ferreira 1903:15; Monard

1937b:57; Loveridge 1957:196); “ Malanje ”

[-9.55000, 16.35000] ( Peters 1881:147; Bocage 1895a:22; Monard 1937b:57); “PungoAndongo” [-9.66667, 15.58333] ( Boulenger

1905:110); “Capanda” [-9.72841, 15.34585]

(Ceríaco et al. 2014b:670); “Cangandala

National Park” [-9.84606, 16.72233] (Ceríaco et al. 2016b:75). Lunda Norte: “Dundo”

[-7.36667, 20.83333] ( Laurent 1950a:12);

“Cassange” [-9.58333, 17.86667] ( Bocage

1895a:22; Monard 1937b:57; Wagner et al.

2018:47); “Carumbo, Lucapa” [-8.42278, MAP 245. Distribution of Acanthocercus cyanocephalus in Angola.

20.73917] (Branch and Conradie 2015:200);

“Cacolo (Minungo)” [-10.15000, 19.28333] ( Manaças 1963:228). Lunda Sul: “Alto Cuílo” [-10.01667, 19.55000] ( Laurent 1964a:38); “Village Sá-Tchisseke, près des sources du Cuílo, Alto Chicapa” [-10.86667, 19.38333] ( Laurent 1964a:39); “Alto Chicapa” [-10.88333, 19.23333] ( Laurent 1964a:38); “Lunda” [-10.96667, 20.06667] ( Monard 1937b:57, 58; Wagner et al. 2018:47).

Moxico: “Fazenda Santa Cruz, Luso” [-11.78333, 19.91667] ( Manaças 1963:228); “environs du lac Calundo “ [-11.80000, 20.86667] ( Laurent 1964a:39); “Calombe, Luso” [-11.83333, 19.93333] ( Manaças 1963:228); “Cazombo, Haut Zambèze ” [-11.88333, 22.91667] ( Laurent 1964a:39); “Calunda, Haut Zambèze ” [-12.11667, 23.46667] ( Laurent 1964a:39); “Cassamba” [-13.10000, 20.35000] ( Manaças 1963:228); “Sessa (Luchazes)” [-13.91667, 20.80000] ( Manaças 1963:228); “Vila Gago Coutinho (Bundas)” [-14.10000, 21.43333] ( Manaças 1963:228). Luanda: “Dande” [-8.46667, 13.38333] ( Wagner et al. 2018:47). Kwanza Sul: “Mombolo” [-12.16667, 14.83333] ( Wagner et al. 2018:47). Bié: “Chitau “ [-11.43333, 17.15000] ( Schmidt 1933:9; Wagner et al. 2018:47); “Silva Porto ” [-12.38333, 16.95000] ( Manaças 1963:228); “Cachingues” [-13.06667, 16.75000] ( Manaças 1963:228). Huambo: “Bimbi” [-11.81667, 15.83333] ( Monard 1937b:57, 58); “Galanga (Galange)” [-12.06667, 15.15000] ( Bocage 1895a:22; Monard 1937b:57); “ Mt. Moco ” [-12.41667, 15.18333] ( Parker 1936:132); “ Huambo ” [-12.76667, 15.73333] ( Wagner et al. 2018:47); “Cuma” [-12.86667, 13.06667] ( Wagner et al. 2018:47). Benguela: “Quissange” [-12.43333, 14.05000] ( Bocage 1895a:22; Monard 1937b:57); “Quindumbo” [-12.46667, 14.93333] ( Bocage 1895a:22; Monard 1937b:57). Huíla: “ Rio Cuce” [-13.51667, 15.20000] ( Bocage 1895a:22; Monard 1937b:57); “Kalukembé” [-13.78333, 14.68333] ( Monard 1937b:57, 58); “Caconda” [-13.73333, 15.06667] ( Bocage 1895a:22; Monard 1937b:57; Themido 1941:7).

Undetermined Locality: “with no precise location” ( Bocage 1879b:95), “Quando” ( Monard 1937b:57); “Cuanza r.” ( Boulenger 1905:110).

Taxonomic and distributional notes: Acanthocercus species outside of Arabia and the Horn of Africa, including those in Angola, have long been referred to A. atricollis (Smith, 1849) . A reference to Stellio nigricolis ( Bocage 1866a) appears to be a lapsus for A. atricollis as well (Ceríaco et al. 2014b; Wagner et al. 2018). Klausewitz (1957) divided this species into several subspecies, but Angolan material fell into the nominotypical form ( Branch 1998, Ceríaco et al. 2014b). Subsequent revisionary work on the genus ( Wagner et al., 2018) revealed that some Angolan Acanthocercus are referrable to A. cyanocephalus , a long-forgotten name proposed by Falk (1925) based on Angolan material, but without a precise type locality. Southern Angolan and northern Namibian specimens referred to A. cyanocephalus by Wagner et al. (2018), however, represent an undescribed lineage, treated here as Acanthocercus sp. (see below). Loveridge (1957) erroneously synonymized Stellio nigricollis cited by Bocage (1866a) from Angola with Agama cyanogaster (Rüppell, 1835) and more recently Branch and Conradie (2015) also cited this species from Angola. This certainly represents a misidentification or a lapsus for A. cyanocephalus , since A. cyanogaster is restricted to the Horn of Africa and adjacent parts of East Africa (Uetz and Hošek 2017).