Ipomoea stans Cav.

Wood, John R. I., Munoz-Rodriguez, Pablo, Williams, Bethany R. M. & Scotland, Robert W., 2020, A foundation monograph of Ipomoea (Convolvulaceae) in the New World, PhytoKeys 143, pp. 1-823 : 1

276. Ipomoea stans Cav. , Icon 3: 26. 1795. (Cavanilles 1795-96: 26)

Convolvulus stans (Cav.) Kunth , Nov. Gen. Sp. 3: 96. 1818 [pub. 1819]. (Kunth 1819: 96).

Convolvulus firmus Spreng. , Syst. Veg. 1: 613. 1825 [pub. 1824]. (Sprengel 1824: 613), nom. illeg. superfl., based on C. stans (Cav.) Kunth

Convolvulus sinuatus Sesse & Mocino , Pl. Nov. Hisp. 1: 24. 1888. ( Sessé y Lacasta and Moçiño 1887-90: 24), nom. illeg. non Convolvulus sinuatus Petagna ex Steudel (1840) . Type. MEXICO. Guanajuato, Ixtla, Sessé & Moçiño 117 [5023], lectotype MA603854, designated by McDonald 2001: 89).

Ipomoea stans var. hirsuta B.L. Rob. , Proc. Amer. Acad. Arts 29: 319. 1894. (Robinson 1894: 319). Type. MEXICO. Jalisco, E. Palmer 324 (lectotype GH n.v., designated by House (1908b: 187), isolectotypes MO, NDG, US, YU).

Ipomoea jaliscana House , Ann. New York Acad. Sci. 18: 187. 1908. (House 1908b: 187). Type. Based on Ipomoea stans var. hirsuta B.L. Rob.

Type.

Plant cultivated in Madrid, presumably of Mexican origin, lectotype MA475857, designated by McDonald (1994: 133) and re-designated here (see below).

Description.

Perennial herb (reported sometimes to be annual), branched at base with many ascending stems to 1 m forming a small bush, stems woody below, crisped pubescent, rootstock tuberous, reputed to be poisonous. Leaves shortly petiolate 2.3-5.5 × 1.5-2.5 cm, oblong to oblong-ovate, obtuse to truncate, base truncate and then broadly cuneate onto petiole, margin lyrate-dentate, both surfaces glabrous with scabrous veins and margins to pubescent with densely pubescent veins, abaxially with prominent venation; petiole 3-5 mm. Flowers solitary (rarely paired), axillary; peduncles 0.5-5 cm, pubescent; bracteoles variable sometimes linear-oblanceolate c. 5 mm long, sometimes foliose with lyrate margins and reaching 15 mm; pedicels 3-10 mm, thickened upwards, scabrous to pubescent; sepals unequal, glabrous to scabrous, margins scarious, outermost 7-12 × 6 mm, broadly to narrowly oblong-elliptic, obtuse, inner 10-16 × 8 mm; corolla 6-7.5 cm long, flared to funnel-shaped, purple, glabrous, limb c. 4 cm diam. Capsules 14-16 × 11 mm, ovoid, shortly rostrate, glabrous, ± enclosed by sepals; seeds 6-8 × 4-5 mm, minutely puberulent.

Illustration.

Carranza (2007: 61); Figures 10G , 139 .

Distribution.

Locally common in open pine forest, dry scrub and secondary vegetation, 1300-2700 m. Endemic to central Mexico.

MEXICO. Est. México & Dist. Fed.: Y. Mexia 2751 (BM, P); Sierra de Guadelupe, E. Bourgeau 496 (BM, MO, P, S); ibid., E.K. Balls & W.B. Gourlay 4948 (K). Guanajuato: E. Ventura & E. López 6964 (MEXU). Guerrero: P. Tenorio et al. 9566 (MO). Hidalgo: Pachuca, C.G. Pringle 6915 (BM, K, P, S); H. Piug 4797 (P). Jalisco: Guadalajara, C.G. Pringle 4488 (BM) - var. hirsuta ; Río Blanco, E. Palmer 324 (BM, K) - var. hirsuta ; Zapopan, A. Rodríguez & P. Montiel-Moncayo 6340(IEB) - var. hirsuta . Michoacán: Morelia, Punguato, G. Arsène 5967 (BM, S, US); Tiripetio, Morelia, G. Cornejo Tenorio 3475 (K). Oaxaca: Ghiesbreght s.n. (K, P); Tihuacan, C. Conzatti 164 (K). Puebla: P. Tenorio & C. Romero 6828 (MO); San Antonio, E.K. Balls & W.B. Gourlay 4510 (BM, K); Juan N. Méndez, J.I. Calzada 24405 (K, MEXU); San Luis Tultitlanapa, C.A. Purpus 3367 (BM). Querétaro: Colón, Galerías, E. Carranza & E. Pérez 4907 (IEB, MEXU). San Luís Potosí: Parry & Palmer 627 (BM, MO). Veracruz: Totalco, M. Vásquez 2100 (BM, K, MEXU). Zacatecas: Dressler 71 (MO); Hartweg s.n. (K).

Typification.

McDonald (1994: 133) designated the lectotype from MA but did not annotate or specify the sheet. Hence we have redesignated MA475857 as the lectotype.

Note.

Very variable in indumentum and size and shape of sepals and bracteoles. Very hairy plants with relatively short but broad sepals from Jalisco (Pringle 4488, E. Palmer 324, A. Rodríguez & P. Montiel-Moncayo 6340) may be recognised as var. hirsuta .

Figures

Figure 10. Pollen of Ipomoea species. A I. triloba (D'Arcy 317) B I. cryptica (Steinbach 6311) C I. purpurea (Parada & Rojas 2664) D I. alba (Wood et al. 27828) E I. dumosa (Hinton et al. 9479) F I. hederifolia (Queiroz 15975) G I. stans (Y. Mexia 275112) H I. suffulta (Pringle 4755). Photos by Robert Scotland.

Figure 139. Ipomoea stans. A habit (1) B abaxial leaf surface C habit (2) D abaxial leaf surface E outer sepal F inner sepal G calyx and capsule. Drawn by Rosemary Wise A, B from Pringle 4488; C-G from Bourgeau 46.