Chalcionellus hauseri (Schmidt, 1894)

Penati, Fabio & Vienna, Pierpaolo, 2006, An updated catalogue of the Histeridae (Insecta: Coleoptera) of the Arabian Peninsula, with biogeographical remarks, Zootaxa 1157 (1), pp. 1-74 : 45-51

47. Chalcionellus hauseri (Schmidt in Hauser)

Distribution: Turkmenistan, Iran, Kazakhstan, Mongolia, Saudi Arabia.

Chorology: Centralasiatic.

Material. Saudi Arabia: “Saudi Arabia”, sine data ( CHTY).

Notes. New to the Arabian Peninsula.

48. Chalcionellus tunisius (Marseul, 1876) ( Fig. 17 )

Distribution: Tunisia, North Africa, Syria, Saudi Arabia, Yemen.

Chorology: N­African.

Literature. Saudi Arabia: Sharoura [= Sharurah, 17°34’31”N 47°09’42”E], 29.III.1979, 1 ex. ( Mazur 1994) .

Material. Saudi Arabia: Daharan , Al Jubail [= Dhahran, al Jubayl, 27°02’50”N 49°39’41”E], 4.IV.2001, 6 ex. (4 ex. MSNTC, 1 ♂ CHFP, 1 ex. CHPV) . Yemen: near Shiakh Othman (Wadi) [=Ash Shaykh`Uthman, 12°52’53”N 44°59’42”E], III.1987, 1 ♀ ( MNHUB) .

Notes. New to Yemen.

49. Pholioxenus kodymi Olexa ( Fig. 18 )

Distribution: Iraq, Syria, Saudi Arabia.

Chorology: SW­Asiatic.

Literature. Saudi Arabia: 90 km N of Riyadh, III–IV.1985, in pitfall traps on sandy soils in several wadis and in sand dune massif Irk­Banban [24°58’33”N 46°34’15”E], more than 80 ex. ( Kryzhanovskij 1987) .

50. Pholioxenus mesopotamicus Olexa ( Fig. 18 )

Distribution: Iraq, Saudi Arabia, Kuwait.

Chorology: SW­Asiatic.

Literature. Saudi Arabia: Wadi Hurayamala [= Wadi Huraymala, 25°07’29”N 46°06’53”E], 770 m, 24.III.1988, 4 ex. ( Tishechkin 2002) .

Material. Kuwait: Al Abdaliyah , 29°03’N 47°43’E, 24.II.1996, 3 ♀ and 1 ♂ ( CMNH) .

Notes. New to Kuwait.

51. Pholioxenus olexai Kryzhanovskij ( Fig. 18 )

Distribution: Saudi Arabia.

Chorology: Endemic.

Literature. Saudi Arabia: 90 km N of Riyadh [= Riyad, 24°38’27”N 46°46’22”E], 27.III.1985, holotype and ibidem, 26.III–14.IV1985, 11 paratypes ( Kryzhanovskij 1987) .

52. Hypocacculus (Colpellus) praecox (Erichson) ( Fig. 19 )

Distribution: Egypt, Mediterranean Subregion, Canary Is., Cape Verde, Afrotropical (= Ethiopian) Region, Saudi Arabia, Afghanistan, Oman.

Chorology: Afrotropico­Mediterranean.

Literature. Saudi Arabia: “El Hahaz” [= El Hejaz] ( Reichardt 1932: 97; 2 ex. in MNHUB seen by us) .

Material. Oman: Dhofar, Al Mughsayl env. W, 16°53’74”N 53°48’97”E, 40 m, 5.IX.2000, 1 ♀ ( MSNTC) .

Notes. New to Oman.

53. Hypocacculus (Hypocacculus) harmonicus (Marseul) ( Fig. 20 )

Distribution: Republic of South Africa (Capland), East Africa, Sudan, Saudi Arabia, Oman.

Chorology: Afrotropical (spread in E and S Africa).

Literature. Saudi Arabia: “El Hahaz” [= El Hejaz], 8 ex. ( Reichardt 1932: 99; 5 ex. in MNHUB examined by us) .

Material. Oman: Dhofar, Salalah env. W [17°01’03”N 54°04’58”E], near Veterinary Hospital, 27.VIII.2000, 1 ex. ( MSNTC) ; Dhofar, Rd. Taqah to Madnat al Haq , 2 km after jct., 5.IX.2002, 3 ex. ( MSNTC) ; Dhofar, Rd. 41 tra Raysut e Adwnab , 16°57’72”N 53°52’73”E, 90 m, 23.IX.2001, 2 ex. (1 ex. MSNTC, 1 ex. CHPV) ; Dhofar, Khor Ron , 17°02’27”N 54°26’16”E, 35 m, 24.VIII.2000, 3 ex. ( MSNTC) ;

ibidem, 12.IX.2000, 2 ex. ( CHFP); Dhofar, Rd. 49 near Ain Sahanawt , 17°07’40”N 54°09’44”E , 24.IX.2001, 2 ex. ( MSNTC); Dhofar, Ain Razat , 17°07’75”N 54°14’42”E, 11.IX.2002, 2 ex. ( MSNTC); Dhofar, Hajaif env., 17°14’53”N 54°01’67”E, 900 m , 4.X.2001, 2 ex. ( MSNTC); Dhofar, Rd. 47 ante Al Mughsayl , 16°53’45”N 53°48’57”E, 30 m , 1.III.2004, 1 ♀ ( MSNTC); Dhofar, Ain Garziz env., 17°06’29”N 54°04’35”E, 110 m , 16.III.2004, 1 ♂ ( MSNTC) .

Notes. New to Oman.

54. Hypocacculus (Hypocacculus) metallescens (Erichson) ( Fig. 20 )

Distribution: Italy (Sardinia), Hungary, Romania, Mediterranean Subregion, South Russia, Central Asia, Africa, Mascarene Is., Saudi Arabia, Oman.

Chorology: undefined.

Literature. Saudi Arabia: “El Hahaz” [= El Hejaz] ( Reichardt 1932: 99; 2 ex. in MNHUB seen by us); Khulais valley [= Khulays, 22°09’14”N 39°19’06”E], 80 km NE of Jeddah, 1997–1998 ( Al­Ghamdi 1999).

Material. Saudi Arabia: “ Saudi Arabia ”, sine data ( CHTY) . Oman: Dhofar, Al Mughsayl env. E, 16°52’53”N 53°43’51”E, 8.III.2004, 1 ♂ ( CHFP) ; Dhofar, 7 km after Tawi Atayr , 17°03’46”N 54°36’15”E, 600 m, 9.III.2004, 1 ex. ( MSNTC) ; Dhofar, Ain Razat env., 17°07’N 54°14’E, 16.III.2004, 1 ex. ( MSNTC) .

Notes. New to Oman. The chorology of this species is undefined, because documentation of its supposed wide distribution preceded the description of the two almost identical afrotropical species H. simillimus Vienna and H. simulans Vienna with which H. metallescens could have been confused in the past.

55. Hypocacculus (Hypocacculus) simulans Vienna in Penati et Vienna ( Fig. 20 )

Distribution: Republic of South Africa, Malawi, Yemen ( Gomy, 2004).

Chorology: Afrotropical (spread in E and S Africa).

Literature. Yemen: Wadi Sharez [a few km east of Hajjah, 15°41’36”N 43°36’11”E], II.1985, 1 ♀ ( Gomy 2004); ibidem, 24.III.1985, cow dung, 6 ♂ and 4 ♀ ( Gomy 2004).

56. Hypocacculus (Hypocacculus) spretulus (Erichson) ( Fig. 20 )

Distribution: Portugal, South Europe, Syria, Crimea, Caucasus, Central Asia, Iran, Saudi Arabia, Sri Lanka, Burma, Vietnam.

Chorology: undefined.

Literature. Saudi Arabia: Fifa near Gizan [= Faifa, 17°08’38”N 43°07’09”E], 1240 m, 27–31.III.1983, 1 ex. ( Mazur 1994) .

57. Hypocacculus (Nessus) controversus Müller ( Fig. 21 )

Distribution: Balkans, Spain, Greece, Turkey, Jordan, Saudi Arabia.

Chorology: Mediterranean.

Literature. Saudi Arabia: Wadi Hurayamala [= Wadi Huraymala, 25°07’29”N 46°06’53”E], 770 m, 10.III.1988, 1 ex. ( Tishechkin 2002) .

58. Hypocacculus (Nessus) hosseinius (Théry) ( Fig. 21 )

Distribution: Morocco, Egypt, Iran, Saudi Arabia, Oman.

Chorology: Saharo­Sahelo­Sindian.

Material. Saudi Arabia: Kushm Dibi [?], 20.IV.1978, 1 ex. ( CHSM) . Oman: Dhofar, Al Mughsayl env., 16°53’90”N 53°46’30”E, 9.IX.2002, 1 ♂ ( MSNTC) .

Notes. New to the Arabian Peninsula. This species seems to be closely related to a few other Palearctic Nessus characterized by striking psammophilous features (body convex, pronotal hypomeron setose, pronotal and elytral surface almost entirely and coarsely punctate, mid and hind tibiae enlarged bearing long spinulae, etc.): H. tigris (Marseul) , H. orbus Reichardt , H. occator Reichardt , H. eremobius Reichardt , H. balux Reichardt and H. neftensis Olexa. Due to their relative rarity, these species are poorly known and for this reason a future comprehensive revision of this group might

ascribe the two specimens of H. hosseinius (Théry) from the Arabian Peninsula to another species.

59. Hypocacculus (Nessus) rubripes (Erichson) ( Fig. 21 )

Distribution: Portugal, Mediterranean Subregion, Netherlands, Central Europe, Crimea, Caucasus, Iran, Central Asia, Mongolia, Russian Far East: Ussuriyskiy Kray, India, tropical Africa.

Chorology: undefined.

Literature. Saudi Arabia: Khulais valley [= Khulays, 22°09’14”N 39°19’06”E], 80 km NE of Jeddah, 1997–1998 ( Al­Ghamdi 1999) .

Material. Saudi Arabia: “Saudi Arabia”, sine data ( CHTY).

Notes. The chorology of this species is undefined, because in the past H. rubripes has been confused with some very similar species.

Figures

FIGURE 17. Distribution of the Histeridae in the Arabian Peninsula. Chalcionellus geminus, Chalcionellus tunisius, Chalcionellus sp..

FIGURE 18. Distribution of the Histeridae in the Arabian Peninsula. Pholioxenus kodymi, Pholioxenus mesopotamicus, Pholioxenus olexai.

FIGURE 19. Distribution of the Histeridae in the Arabian Peninsula. Hypocacculus (C.) praecox, Hypocacculus (C.) sp., Hypocacculus (C.) sp..

FIGURE 20. Distribution of the Histeridae in the Arabian Peninsula. Hypocacculus (s. str.) harmonicus, H. (s. str.) metallescens, H. (s. str.) simulans, H. (s. str.) spretulus.

FIGURE 21. Distribution of the Histeridae in the Arabian Peninsula. Hypocacculus (N.) controversus, H. (N.) hosseinius, H. (N.) rubripes.