Brocchinia subanodosa Sacco, 1894

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 15

Brocchinia subanodosa Sacco, 1894

Figs 3D, 4H 1–H 3

Brocchinia mitraeformis (Br.) — Sacco 1894: 68, pl. 3, fig. 81 [non Voluta mitraeformis Brocchi, 1814 ].

Brocchinia mitraeformis var. subanodosa Sacco 1894: 69 , pl. 3, fig. 83.

Cancellaria (f. Narona ) mitraeformis Brocc. — Hoernes & Auinger 1890: 280, pl. 33, figs 17a–c [non Voluta mitraeformis Brocchi, 1814 ].

Brocchinia mitraeformis (Brocc.) — Boettger 1902: 40 [non Voluta mitraeformis Brocchi, 1814 ].

Cancellaria (Brocchinia) mitraeformis Brocc. — Sieber 1936: 92 [non Voluta mitraeformis Brocchi, 1814 ].

Narona (Brocchinia) mitraeformis (Brocc.) — Sieber 1956: 245 [non Voluta mitraeformis Brocchi, 1814 ].

Narona (Brocchinia) mitraeformis parvula (Beyr.) — Hölzl 1973: 194 [non Cancellaria parvula Beyrich, 1856 ].

Narona (Brocchinia) mitraeformis parvula ( Beyrich, 1856) — Steininger 1973: 435 [non Cancellaria parvula Beyrich, 1856 ].

Brocchinia tauroparva Sacco, 1894 — Landau et al. 2006: 81, pl. 7, figs 5–6 [non Brocchinia tauroparva Sacco, 1894 ].

Brocchinia subanodosa Sacco, 1894 — Brunetti et al. 2011: 121, figs 20A–H, 22C–D [cum syn.].

Type material. lectotype BS.045.16.003 designated by Brunetti et al. (2011), Museo Regionale di Scienci Naturali , Torino ( Italy); Bordighera ( Italy), Piacenzian (Late Pliocene) .

Illustrated material. Figs 3D, 4H 1–H 3: specimen illustrated in Hoernes and Auinger (1890, pl. 33, figs 17a– c), Bad Vöslau ( Austria), height: 13.4 mm, width: 5.3 mm, NHMW 1872/0030/0106.

Studied material. 7 spec. Niederleis ( Austria) NHMW 1860 /0015/0701; 1 spec. Bad Vöslau ( Austria) NHMW 1872 /0030/0106; 1 spec. Forchtenau ( Austria) NHMW 1866 /0001/1080; 1 spec. Lăpugiu de Sus ( Romania) NHMW 1859 /0037/0030.

Discussion. Recently, Brunetti et al. (2011) clarified the status of this species, which was frequently intermingled with Brocchinia mitraeformis ( Brocchi, 1814) or B. tauroparva ( Sacco, 1894) in the literature ( Chirli 2002, Landau et al. 2006 and references therein). Brunetti et al. (2011) state that the protoconch of B. subanodosa has 2.75 whorls instead of 2 in B. depressiplicata Sacco, 1894 [= Voluta mitraeformis Brocchi, 1814 , which is a primary homonym of Voluta mitraeformis Lamarck, 1811 ]. Differences in the teleoconchs are the reduced axial sculpture and the broader spiral cords of B. subanodosa . The Austrian shells agree fully in protoconch morphology and teleoconch outline and sculpture with B. subanodosa as defined by Brunetti et al. (2011).

Although Voluta mitraeformis Brocchi, 1814 is a primary homonym of Voluta mitraeformis Lamarck, 1811 , a replacement name is not obligatory according to ICZN Art. 23.9.5, as both taxa have been considered as separate genera after 1899. Therefore, the proposal of Brunetti et al. (2011) is disputable and Brocchinia mitraeformis ( Brocchi, 1814) is a valid name.

Distribution. This species occurs from the Burdigalian to Langhian in the Paratethys and appears during the Late Miocene in the Proto –Mediterranean Sea where it becomes a ubiquitous species during the Pliocene. Its last occurrence is mentioned by Brunetti et al. (2011) from the Gelasian of Italy.

Paratethys —Ottnangian: North Alpine Foreland Basin (Bavaria: Gernergraben, Kaltenbachgraben); Badenian: Vienna Basin ( Austria: Niederleis, Bad Vöslau); Eisenstadt–Sopron Basin ( Austria: Forchtenau); Transylvanian Basin ( Romania: Lăpugiu de Sus, CoŞteiu de Sus). Proto –Mediterranean Sea —Tortonian: Po Basin ( Italy: Montegibbio); Early Pliocene: Italy; Spain (Estepona), Early Pleistocene: Italy.