Nannaria fracta, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

publication ID

http://dx.doi.org/10.3897/zookeys.1030.62544

publication LSID

lsid:zoobank.org:pub:48997D3B-9B4E-4348-8F30-0C8E8672569A

persistent identifier

http://treatment.plazi.org/id/8D8D39A0-7084-4BA9-8F47-9D0C7C99F55B

taxon LSID

lsid:zoobank.org:act:8D8D39A0-7084-4BA9-8F47-9D0C7C99F55B

treatment provided by

ZooKeys by Pensoft

scientific name

Nannaria fracta
status

sp. nov.

Nannaria fracta   sp. nov. Figs 80 View Figure 80 , 81 Vernacular name: "The Breaks Interstate Park Twisted-Claw Millipede" View Figure 81

Material examined.

Holotype: United States - Virginia • ♂; Dickenson County, Haysi, Breaks Interstate Park , Laurel Branch Trail at intersection with Cold Spring Trail ; 37.2897°N, - 82.2999°W; elev. 565 m; 28 Sep. 2017; hand collected; J. Means, D. Hennen leg.; VTEC MPE031781. GoogleMaps  

Paratypes: United States - Virginia • ♂; same collection data as holotype; VTEC, MPE03179 View Materials GoogleMaps   1 ♂; same collection data as holotype; VMNH MPE03756 View Materials GoogleMaps   2 ♂♂; same collection data as holotype; VTEC MPE03183 View Materials GoogleMaps   , 84; 1 ♂; Dickenson County, Breaks Interstate Park , in camping area; 37.2936°N, - 82.3005°W; 16 Apr. 1983; D. Ogle leg.; VMNH NAN0156 • SCAU GoogleMaps   - Kentucky • 2 ♀♀; Pike County, Pikeville, Bob Amos Park , 424 Bob Amos Dr. WW Gearheart Hiking Trail ; 37.4690°N, - 82.5462°W; elev. 359 m; 28 Sep. 2017; hand collected; J. Means, D. Hennen leg.; VTEC MPE03185 View Materials GoogleMaps   , 86.

Other material.

United States - Virginia • 1 ♂; Dickenson County, Breaks Interstate Park ; 37.2936°N, - 82.3005°W; 7 Sep. 1967 GoogleMaps   ; Neff, R. Hoffman leg.; VMNH NAN0154 • 1 ♂; Russell County, 1 mile NW of Lynn Spring ; 37.1153°N, - 81.9411°W; 20 Apr. 1962 GoogleMaps   ; R. Hoffman leg.; VMNH NAN0155 • 1 ♂; Washington County, Clinch Mountain Wildlife Management Area, deciduous forest above Big Tumbling Creek ; 36.9931°N, - 81.7368°W; 21 Sep. 2011 GoogleMaps   ; S. Roble leg.; VMNH NAN0157. For detailed collection data see Suppl. material 7   .

Diagnosis.

Adult males of Nannaria fracta   sp. nov. are distinct from other Nannaria   and the nearby N. aenigma   , based on the following combination of characters: Gonopods. Gonopodal acropodite acicular, slightly curving medially before tip, not bending abruptly medially at 90° at tip as in N. bobmareki   sp. nov. or gently curving medially as in N. aenigma   . Distal zone short, rounded-not quadrate as in N. bobmareki   sp. nov., or long and serpentine as in N. aenigma   . Acropodite with small, shelf-like medial flange just before tip (Fig. 80C View Figure 80 , red arrow). Prefemur with long, acicular prefemoral process with a pronounced, sharp prefemoral spine (Fig. 80C View Figure 80 , red triangle), not fused with prefemoral process as in N. bobmareki   sp. nov. or lacking as in N. aenigma   . Telopodite basal zone > ½ length of acropodite, not ca. 1/2 length of acropodite as in N. bobmareki   sp. nov. or ca. 1/6 length of acropodite as in N. aenigma   . Telopodite basal zone basal zone with pronounced medial swelling (Fig. 80A View Figure 80 , red circle), not with slight medial swelling as in N. bobmareki   sp. nov., or lacking as in N. aenigma   . Color: Tergites with red paranotal spots (Fig. 81 View Figure 81 ). Dark brown background. Dorsum of collum smooth with red margin.

Measurements.

♂ holotype (VTEC, MPE03178): BL = 28.9, CW = 3.7, IW = 1.8, ISW = 0.85, B11W = 4.4, B11H = 2.5; ♀ paratype (VTEC, MPE03185): BL = 35.0, CW = 4.5, IW = 2.7, ISW = 1.2, B11W = 5.6, B11H = 4.2.

Variation.

No known variation.

Distribution.

Nannaria fracta   sp. nov. has a linear distribution extending from eastern Kentucky into western Virginia (Kentucky: Pike County; Virginia: Dickenson, Russell, and Tazewell counties; Suppl. material 7, Fig. 126 View Figure 126 ). Distribution area: 98 km2; status: MRE.

Ecology.

Individuals of N. fracta   sp. nov. have been collected from mesic hardwood forests composed of beech, maple, tuliptree, hemlock, and rhododendron. Specimens taken from Bob Amos Park in Kentucky were found under 1-2 cm of hardpacked, dark soil on the side of a hiking path, while specimens collected from Breaks Interstate Park in Virginia were found in a very moist rhododendron cove along the bank of Laurel Branch Creek, under 1-2 cm of sandy, dark soil.

Etymology.

This species is named for Breaks Interstate Park, where it was originally collected by R. L. Hoffman in 1962. The specific name is an adjective derived from the Latin fractura, meaning break or fracture.

Type locality.

United States, Virginia, Dickenson County, Haysi, Breaks Interstate Park, Laurel Branch Trail at intersection with Cold Spring Trail, 37.2897°N, - 82.2999°W.