Bothriurus, MONOPHYLY,

Vrech, David E., Peretti, Alfredo V. & Mattoni, Camilo I., 2011, Sperm package morphology in scorpions and its relation to phylogeny, Zoological Journal of the Linnean Society 161 (3), pp. 463-483: 477

publication ID 10.1111/j.1096-3642.2010.00647.x

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Our results support a nonmonophyletic genus Bothriurus  as suggested by Prendini (2000, 2003), Mattoni (2003), and Mattoni & Prendini (2007). Four different groups can be recognized in the analysed species of this genus. The first is composed of members of the Bothriurus  subgenus ( Bo. araguayae  , Bo. bonariensis  , and Bo. chacoensis  ) with packages that are either thick and straight or slim and coiled anteriorly. Another group is formed by Bo. bocki  and Bo. inermis  , with packages similar to those of Timogenes  and Vachonia  . The remaining Bothriurus species  (subgenus Andibothriurus  ) are characterized by cane-like sperm packages. Bothriurus  from the Bo. rochai  group and Bo. asper  (both belonging to the Bothriurus  subgenus) have loosely aggregated spermatozoa and do not show ordered sperm packages.