Liogenys, Guerin-Meneville, 1831

Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. & Almeida, Lúcia M., 2020, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae) from the southern South American Transition Zone and boundaries: taxonomic overview with four new species, Zootaxa 4896 (1), pp. 46-84 : 48-49

publication ID

https://doi.org/10.11646/zootaxa.4896.1.2

publication LSID

lsid:zoobank.org:pub:92DECB9B-36F8-45D8-8F3A-AF91A3B73FE8

DOI

http://doi.org/10.5281/zenodo.4385619

persistent identifier

http://treatment.plazi.org/id/357687CC-FFC9-9A47-FF03-E9F94BACFD03

treatment provided by

Plazi

scientific name

Liogenys
status

 

Identification key to Liogenys   from the southern South American Transition Zone and boundaries *

*Note on dimorphism: Males have protarsi and mesotarsi flat (wide or not) and females have all tarsi cylindrical.

1. Length of antennal club two times longer than the funicle or almost so ( Fig. 10D View FIGURE 10 ); head, pronotum, and scutellum dark purplish brown, elytra yellow ( Fig. 10A View FIGURE 10 ); elytra glabrous; metatibia with inner margin not carinate in both sexes........................................................................................... Liogenys xanthocera Harold, 1869  

- Length of antennal club as long as or slightly longer than funicle; head, pronotum, scutellum, and elytra unicolored, yellow or light brown, elytra sometimes slightly lighter in color than pronotum; elytra glabrous or with bristles; male with metatibia weakly or strongly carinate toward apex....................................................................2

2. Head more coarsely punctate than pronotum ( Fig. 8B View FIGURE 8 ); pronotum wider than elytra at its humeral callus; galea with a single, long, hook-like tooth ( Fig. 13G View FIGURE 13 ); antenna with 9 antennomeres; elytra with bristles............................................................................................... Liogenys maxillaricuspis Cherman   , new species

- Head as punctate as or less coarsely punctate than pronotum; pronotum as wide as or slightly narrower than elytra at its humeral callus; galea with four or five teeth ( Fig. 13E View FIGURE 13 ), the superior tooth slightly larger than the other teeth; antenna with 9–10 antennomeres; elytra glabrous or with bristles................................................................... 3

3. Pygidium longer than wide ( Fig. 1E View FIGURE 1 ) and always with bristles.................................................. 4

- Pygidium as wide as or wider than long, glabrous or with bristles................................................6

4. Antenna with 10 antennomeres; body length up to 8.5 mm; clypeus emargination in general deep, eventually shallow, narrower than the distance between eyes in both sexes; protibia with three teeth in both sexes; males with inner margin of metatibia carinate towards apex or not................................................................................ 5

- Antenna with 9 antennomeres; body length more than 8.5 mm; clypeus emargination noticeably shallow, females with emargination narrower than the distance between eyes, males with emargination as wide as the distance between eyes ( Fig. 4F View FIGURE 4 ); protibia in males with two teeth and in females with three teeth; males with inner margin of metatibia carinate......................................................................................... Liogenys flavida Moser, 1918  

5. Head as wide as or narrower than posterior margin of pronotum ( Fig. 9C View FIGURE 9 ); clypeus as long as frons; vertex finely to coarsely punctate; males with inner margin of metatibia carinate straight towards apex; protarsi noticeably enlarged, protarsomere II wider than long.............................................................. Liogenys rufoflava Moser, 1918  

- Head distinctly wider than posterior margin of pronotum ( Fig. 1B View FIGURE 1 ); clypeus noticeably shorter than frons; vertex smooth ( Fig. 1B View FIGURE 1 ); males with inner margin of metatibia not carinate; protarsi slightly enlarged, protarsomere II longer than wide..................................................................... Liogenys brachyclypeata Cherman   , new species

6. Elytra and pygidium glabrous; males with inner margin of metatibia carinate towards apex; protarsomere II noticeably wider than long; apex of parameres lanceolate in a single plane ( Figs. 3F View FIGURE 3 , 9F View FIGURE 9 )................... Liogenys flaveola Moser, 1924  

- Elytra and pygidium with bristles; males with inner margin of metatibia weakly or not carinate, if carinate, carina never reaching the apex; protarsomere II as long as or longer than wide; apex of parameres lanceolate or not, if lanceolate, never in a single plane ( Figs. 2F View FIGURE 2 , 5F View FIGURE 5 , 7F View FIGURE 7 )................................................................................. 7

7. Elytra dull yellow, never pruinose or if so, not uniformly, pronotum unicolored with elytra; males with metatibial transverse carinae incomplete ( Fig. 6E View FIGURE 6 ).............................................................................8

- Elytra light brown, uniformly pruinose, pronotum dark brown, darker than elytra; metatibia with one or two transverse carinae complete in both sexes ( Fig. 5D View FIGURE 5 .......................................................................... 9 View FIGURE 9

8. Clypeal lateral margin with a blunt projection ( Fig. 2D View FIGURE 2 ), convex; clypeal teeth well developed in both sexes, teeth acute; protibia with three teeth in both sexes; males with protarsomere and mesotarsomere II equally wide; spurs of metatibia uneven in shape ( Fig. 2E View FIGURE 2 )................................................................ Liogenys calcarata Frey, 1970  

- Clypeal lateral margin not produced, straight or concave, clypeal teeth well developed only in males, teeth broad; protibia in males with two teeth and in females with three teeth; males with protarsomere II noticeably wider than mesotarsomere II; spurs of metatibia even in shape ( Fig. 6C View FIGURE 6 ).................................. Liogenys lucialmeidae Cherman   , new species

9. Elytra shiny or semiopaque, never pruinose; ventrites and pygidium with few bristles; males with two protibial teeth and females with three teeth.......................................................... Liogenys kuntzeni Moser, 1921  

- Elytra opaque, noticeably pruinose ( Fig. 7E View FIGURE 7 ); ventrites and pygidium with thick, scale-like bristles; both sexes with three protibial teeth........................................................ .. Liogenys martinezi Cherman   , new species