Chassalia beamanii T.Y.Yu,

Turner, Ian M. & Cheek, Martin, 2021, Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species, European Journal of Taxonomy 738, pp. 1-60: 12-15

publication ID

https://doi.org/10.5852/ejt.2021.738.1261

DOI

http://doi.org/10.5281/zenodo.4599371

persistent identifier

http://treatment.plazi.org/id/355087D8-FFF2-E46D-181B-FCF4EA7DC735

treatment provided by

Plazi

scientific name

Chassalia beamanii T.Y.Yu
status

sp. nov.

Chassalia beamanii T.Y.Yu  sp. nov.

urn:lsid:ipni.org:names:77215403-1

Figs 6–7View FigView Fig

Diagnosis

Differs from other big-leaved species in having leaves clustered at apex of stem, in infructescences mostly terminal, in fruits significantly obovoid (fruits of most other Asian species of Chassalia  are ovoid or round).

Etymology

Named after American botanist, John Homer Beaman (1929–2015), who collected this species and most other specimens of Chassalia  from around Mount Kinabalu.

Type

MALAYSIA – Borneo , Sabah • Ranau District, near Poring Hot Springs along trail to tourist canopy walkway; 6°03′ N, 110°42′ E; 9 Aug. 1990; John H. Beaman 10924; holotype: K[ K001129721]!; isotype: MSCGoogleMaps  .

Additional material

MALAYSIA – Borneo , Sabah • Ranau District, Kampung Melangkap Tomis, 3 km dari Melangkap Tomis; 10 Sep. 1995; Lorence Lugas 954; K!, SNP  Kawasan Labi ; 14 Jan. 1996; Lorence Lugas 1676; K!, SNP  .

Description

Shrub or small treelet, around 1 m tall, glabrous. Stem terete, hollow, slightly flattened when dry, 4–6(– 8) mm wide; successive nodes flattened at right angles, internode on the flowering branch 1–3 cm long, gradually compressed at apex. Leaves opposite, decussate. Leaf-blades obovate to broadly elliptic, 21– 28 × 7–11 cm, broadest at about middle or above; apex acute to attenuate; base attenuate; midrib slightly raised above, domatia absent; secondary nerves 12–18 on each side of the midrib, prominent above. Petioles flattened, canaliculate, 3–4 × 0.1–0.2 cm, 1 cm long when immature. Stipule not sheathing, broadly flabellate, 5–10 × 8–15 mm, apex broken very easily, base drying deep brown, 5–8 × 8–10 mm, sometimes very small, 3 × 2 mm; margin light brown. Inflorescences compound cymes, 1.5–4 cm long; peduncle 8–20 × 2 mm, rachis and branches 3–15 mm or absent. Bracts 4–9 in each capitulum, 3–6 × 3–4 mm. Flowers not seen. Fruits obovoid, broadest at ¾ from base, 6–7 × 4–6 mm, with 2–3 veins visible on pericarp; pedicel of fruit short, 2–3 mm; calyx tube persistent or broken, 0.5–1 × 1–2 mm, lobes usually broken. Disc persistent, longer than persistent calyx tube, 0.8–1.2 × 0.7–0.8 mm. Pyrenes plano-convex, dorsal surface convex, with two adjacent parallel longitudinal crests, each 0.1 × 0.3 mm, curved, 0.3 mm apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, central part slightly elevated, margin with an endocarpal outgrowth 0.3 × 0.6 mm, with a groove between the edge of endocarp and endocarpal outgrowth 0.2–0.3 mm wide. Raphal opening at base of the endocarpal outgrowth, narrowly ovoid, 0.5 × 0.2 mm, 0.2–0.3 mm from the edge of the pyrene wall, apex with a bony spine, spine triangular, erect, 0.5–0.7 × 0.2–0.3 mm. PGS not detected. Seed light brown, hemi-ellipsoid, slightly crested at central part of dorsal side, 0.3 × 4 mm, endosperm not detected.

Distribution, habitat

Endemic to Borneo. Only known from the Ranau District of Sabah. Lowland forest, alt. 650– 750 m.

MSC

Michigan State University

SNP

Sabah Parks