Poecilovitila japonica, Papp, László, 2011

Papp, László, 2011, Oriental Teratomyzidae (Diptera: Schizophora), Zootaxa 2916, pp. 1-34 : 23-25

Poecilovitila japonica sp. nov.

( Figs 85–92 , 118 )

Type material. Holotype 3 ( HNHM): JAPAN: Kyushu, Oike, No. 6, 29. 0 9. 2006, 865 m, Mt Kurodake area, N33º07’29.8” E131º17’34.0”, along stream, leg. M. Földvári.

Description. Measurements in mm: body length 2.62; wing length 2.66; wing width 0.82.

Head. Prefrons (facial plate) with a broad dark sagittal (central) quadratic spot; lateral parts as well as genae with dense whitish yellow microtomentum. Prefrons broader than e.g. in P. defecta . Frons strongly (brightly) shining, its microtrichia rather sparse and short. Gena comparatively narrow, only 0.06 mm below eye, oblique or almost vertical. Genal setae not very long: 3 pairs of long but not peristomal setae, longest one 0.25 mm. Postgena and occiput darker grey. Scape much longer than pedicel or its own width (0.11 mm vs 0.075 mm), without distinct setae. Pedicel with several long setae, longest (subdorsal) 0.11 mm. First flagellomere distinctly longer than high, less “cut” with less dense cilia apically. Arista whitish yellow.

Thorax. Mesonotum rather smooth, moderately shining. Scutellum definitely though finely, granulose (rugose). Mesonotum, scutellum and dorsal parts of pleura dark blackishbrown. Ventral parts of pleura up to 2/5 of anepisternum and 1/3 of anepimeron yellow, strongly contrasting to the dark dorsal parts.

Wing. Ve in R 2+3 much thickened medially, touching costa ( Fig. 118 ). Wing membrane lighter brown, white spot longer than high (0.15 long, 0.11 high). Veins brown but darker on darker areas. Dark brown colour in cell r1 at edge and most apically, cell r2+3 almost entirely dark brown, excluding white spot and a diffuse lighter area below R2+3. Also upper part of r4+5 dark brown. Somewhat darker diffuse colour is in m cells and basally on wing. Length of costal sections (all wing measurements in mm) between R1 and R2+3 1.07, between R2+3 and R4+5 0.80, between R4+5 and M1+2 0.36, no inter-crossvein section, costal seta 0.082, length of basal cell 0.21, length of discal cell 0.37, distance between apices of M veins 0.77, cell r1 only 0.01 mm at narrowest, vein R2+3 0.022 mm thick there. Costal vein continued on a section of 0.035 mm over the apex of R4+5. M3+4 recurved sub-basally, narrowed from 0.10 mm to 0.083 mm at middle. Halter ochre, apical half of knob darker brown.

Legs yellow, apices of femora and the very base of tibiae light brown.

Abdomen. Abdominal tergites dark, blackish brown, sternites yellow, sternal setae very sparse. Sclerites caudal to (inside) pregenital sclerites (ring) very distinct. Male cerci with extremely long setae, also lateral surface of surstylus ( Figs 85, 89 ) with long setae. Surstylus ( Fig. 89 ) subquadrate, with 16–17 small black blunt pegs in its medial apical edge and other 3–4 similar pegs on its caudal subapical part. Subepandrial sclerites ( Fig. 87 ) medium large, quadrate. Distiphallus rather short with short thin curved apical process, mesophallus twice longer, ribbons not thin. Phallapodeme ( Fig. 86 ) comparatively short and robust. Pregonites ( Fig. 91 ) rather large, marginal setae (6) rather short. Postgonites ( Fig. 92 ) thin, curved medially, similarly but less so for pregonites. Epiphallus ( Fig. 92 ) short and rounded, much shorter than broad. Duct of ejaculatory apodeme to basiphallus shorter than length of apodeme. Ejaculatory apodeme ( Figs 88, 90 ) well sclerotised but nowhere dark; its broadened part is almost as long as its rod.

Remark. There are numerous material in collections in Japan, incl. that of T. Saigusa’s collection.

Etymology. The specific epithet of this new species refers to the country where it has been collected.

Figures

FIGURES 85 – 92. Poecilovitila japonica sp. n., male genitalia. 85 = outer genitalia, in caudal view, 86 = phallapodeme, dorsal view, 87 = subepandrial sclerite with contours of surstyli and cerci, ventral view, 88 = ejaculatory apodeme, lateral view, 89 = surstylus, in broadest inner (medial) view, 90 = ejaculatory apodeme, dorsal view, 91 = pregonite with hypandrium, lateral view, 92 = epiphallus and postgonites, caudal view. Scales: 0.2 mm for Figs 85, 87 – 88, 0.1 mm for Figs 86, 89 – 92.

FIGURES 118 – 121. Wings of Poecilovitila new species. 118 = P. japonica sp. nov.; 119 = P. taiwanica sp. nov., holotype female; 120 = P. thaii sp. nov.; 121 = P. variegata sp. nov. Scales: 1.0 mm each.