Ipomoea clausa Rudolph. ex Ledeb. & Adlerstam

Wood, John R. I., Munoz-Rodriguez, Pablo, Williams, Bethany R. M. & Scotland, Robert W., 2020, A foundation monograph of Ipomoea (Convolvulaceae) in the New World, PhytoKeys 143, pp. 1-823 : 1

215. Ipomoea clausa Rudolph. ex Ledeb. & Adlerstam , Diss. Bot. Pl. Doming.14. 1805. (Ledebour and Adlerstam 1805: 14)

Exogonium pedatum Choisy , Mém. Soc. Phys. Genève 8: 130 [52]. 1838. (Choisy 1838: 130 [52]). Type. DOMINICAN REPUBLIC. Santo Domingo, P.A. Poiteau s.n. (holotype G00418215, isotype P00666136).

Ipomoea viridiflora Urb. , Symb. Antill. 3: 348. 1902. (Urban 1902-3: 348). Type. HAITI. C. Ehrenberg 345 (holotype?B†, isotype US00111489).

Exogonium viridiflorum (Urb.) House , Bull. Torrey Bot. Club 35: 106. 1908. (House 1908a: 106).

Ipomoea buchii Urb. , Symb. Antill 3 (3): 356. 1903. (Urban 1902-03: 356). Type. HAITI. Near Petit Coupe, W. Buch 817 (holotype B†, isotype US00111368).

Ipomoea samanensis Urb. , Repert. Spec. Nov. Regni Veg. 20: 343. 1924. (Urban 1924a: 343). Type. DOMINICAN REPUBLIC. On south side of Samana Bay, W.L. Abbott 1282 (isotype GH00054579).

Ipomoea pitoniana Urb. , Repert. Spec. Nov. Regni Veg. 24: 10. 1927. (Urban 1927: 10). Type. HAITI. Massif du Nord, Port de Paix, Haut Piton, E.L. Ekman H4603 (S07-4774, lectotype designated here; isolectotypes K, S).

Ipomoea selleana Urb. , Repert. Spec. Nov. Regni Veg. 24: 11 1927. (Urban 1927: 11). Type. HAITI. Massif de la Selle. Nouvelle Touraine, Chapelle Faure, 1000 m, E.L. Ekman H1532b (holotype S07-4779).

Ipomoea hospitalis Urb. , Ark Bot. 23 A(5): 102. 1930. (Urban 1930: 102). Type. HAITI. Port-au-Prince, Massif de la Selle, 2 Oct. 1927, E.L. Ekman H9111 (holotype S07-4467).

Ipomoea hotteana Urb. & Ekman , Ark. Bot. 23 A (5): 103. 1930. (Urban 1930: 103). Type. HAITI. Massif de la Hotte, group Morne-Rochelois, Miragoane, limestone cliffs near Etang-Miragoane, E.L. Ekman H7227 (lectotype S07-4472, designated here; isolectotype S).

Type.

DOMINICAN REPUBLIC. P. A. Poiteau (possible fragment US, possible isotypes G, P).

Description.

Slender woody twiner; stems pale brown, usually glabrous. Leaves petiolate, 2-7 × 1.5-6 cm, polymorphic, ovate-deltoid, entire or 3-lobed or palmately divided into 5 pedate, ovate to oblanceolate lobes, apex acute, obtuse or emarginate and mucronate, base truncate or weakly cordate and broadly cuneate onto petiole, abaxially paler, both surfaces glabrous; petioles 1.3-4.2 cm. Inflorescence of pedunculate axillary cymes; peduncles strikingly variable in length from 1-10 cm; bracteoles 1-2 mm, linear-lanceolate, scarious, caducous; secondary peduncles 5-13 mm; pedicels 3-15 mm; sepals subequal, glabrous, coriaceous, margins scarious, outer 8-10 mm, elliptic, rounded or obtuse, inner similar but 9-11 mm; corolla 3-5 cm long, abruptly widened above the short basal tube but not at limb, greenish-white, glabrous, limb weakly lobed, c. 2 cm diam.; stamens included. Capsules globose, glabrous; seeds with long woolly hairs.

Distribution.

Endemic to Hispaniola growing in scrub at low altitudes.

HAITI. Isla Tortue E.L. Ekman H4085 (S); ibid., E.L. Ekman H9744 (K, S); ibid., E.C. Leonard 13901 (K, MO, NY); Massif de Cahos, Gonaïves, E.L. Ekman H9064 (S); Massif des Matheux, E.L. Ekman 9162 (K, S).

DOMINICAN REPUBLIC. Sine loc., R. Schomburgk 1857 (K); Sierra Martín Garcia , Barahona, M. Mejía et al. 1282 (NY); Sierra Prieta, A.H. Liogier 24108 (NY); La Romana, A.H. Liogier 24231 (NY); Peravia, T.A. Zononi et al. 18081 (NY).

Notes. The location of the original material used for the description of Ipomoea clausa is uncertain. There is a fragment at US, which may belong but it was probably based on a duplicate of the same Poiteau collection which is at G and P.

The type of each name represents a form with distinct leaves: Ipomoea clausa has 3-lobed leaves; I. hospitalis has small deltoid leaves c. 2 cm long; I. hotteana is a form with digitate leaves, the terminal lobe oblanceolate c. 4.5 cm long; I. pitoniana has deltoid leaves which are commonly shallowly lobed and c. 4 cm long; I. selleana is similar in leaf form but the leaves are less lobed and the margins strongly undulate. All forms have obscure hairs on the stem and leaf veins but these are more obvious abaxially in I. selleana .

This variable species is in many ways a Hispaniola counterpart of the Cuban Ipomoea alterniflora .

• Species 216-217 are sisters to each other and sisters to the rest of Clades A1-2. They are very different in their calyx structure.