Nannaria cryomaia, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

publication ID

http://dx.doi.org/10.3897/zookeys.1030.62544

publication LSID

lsid:zoobank.org:pub:48997D3B-9B4E-4348-8F30-0C8E8672569A

persistent identifier

http://treatment.plazi.org/id/B5C5C054-0EAE-48AB-8422-CC516979A45E

taxon LSID

lsid:zoobank.org:act:B5C5C054-0EAE-48AB-8422-CC516979A45E

treatment provided by

ZooKeys by Pensoft

scientific name

Nannaria cryomaia
status

sp. nov.

Nannaria cryomaia   sp. nov. Figs 96 View Figure 96 , 97 Vernacular name: "The Frozen Head Twisted-Claw Millipede" View Figure 97

Material examined.

Holotype: United States - Tennessee • ♂; Morgan County, campground at Frozen Head State Park ; 36.1321°N, - 84.4978°W; elev. 423 m; 12 May 2017; hand collected; J. Means, D. Hennen, V. Wong leg.; VTEC MPE02642 View Materials . GoogleMaps  

Paratypes: United States - Tennessee • 1 ♂; same collection data as holotype; VMNH MPE03765 View Materials GoogleMaps   2 ♀; same collection data as holotype; VTEC MPE02793 View Materials GoogleMaps   , 94 • 2 ♀; same colletion data as holotype; VMNH MPE02795 View Materials GoogleMaps   , 96.

Other material.

United States - Tennessee • 1 ♂; Morgan County, Frozen Head State Park ; 36.1321°N, - 84.4978°W; 29 May 1980; R. Shelley, MSM leg.; NCSM NAN0452. For detailed collection data see Suppl. material 7 GoogleMaps   .

Diagnosis.

Adult males of Nannaria cryomaia   sp. nov. are distinct from other Nannaria   and the nearby Nannaria   sp. nov. ‘Cratagae’ ( Nannaria wilsoni   species group) based on the following combination of characters: Gonopods. Gonopodal acropodite curving medially before apex, not straight as in Nannaria   sp. nov. ‘Cratagae.’ Apex with distinct constriction (Fig. 96A View Figure 96 , red arrow), tip expanded distally, with small lobed lateral and medial flanges, not with acuminate, triangular lateral flange as in N. kassoni   sp. nov. Apex without ventrally directed bend as in Nannaria   sp. nov. ‘Cratagae.’ Acropodite with expanded medial flange (Fig. 96A View Figure 96 , red triangle). Telopodite basal zone height > 1/3 length of acropodite, not ca. 1/3 as in N. kassoni   sp. nov., and not ca. 1/4 length as in Nannaria   sp. nov. ‘Cratagae.’ Prefemur with medially curving prefemoral process, paralleling curve of acropodite, not straight, acicular as in Nannaria   sp. nov. ‘Cratagae.’ Prefemur with short, triangular prefemoral spine, not pronounced, curving as in N. kassoni   sp. nov. Color. Tergites with faint orange stripes (Fig. 97 View Figure 97 ). Dark brown background. Dorsum of collum smooth with orange margin.

Measurements.

♂ holotype (VTEC, MPE02642): BL = 24.9, CW = 3.9, IW = 2.4, ISW = 0.9, B11W = 4.7, B11H = 3.2; ♀ paratype ( VMNH, MPE02796): BL = 26.6, CW = 3.9, IW = 2.2, ISW = 1.0, B11W = 5.2, B11H = 3.7.

Variation.

No known variation.

Distribution.

Known only from the type locality (Tennessee: Morgan County; Suppl. material 7; Fig. 126 View Figure 126 ). Distribution area: N/A; status: MRE.

Ecology.

Individuals of Nannaria cryomaia   sp. nov. were collected from a mesic hardwood forest dominated by oak, ironwood, beech, and hemlock, under moist leaf litter.

Etymology.

This species is named for its type locality, Frozen Head State Park in the Crab Orchard Mountains of eastern Tennessee. The specific name is a latinized noun in apposition derived from the Greek kryos -, meaning frozen, and maia - for a kind of crab.

Type locality.

United States, Tennessee, Morgan County, campground at Frozen Head State Park; 36.1321°N, - 84.4978°W.