Nusasyntrophus echinatus (Heller) Riedel, 2024

Riedel, Alexander, 2024, Taxonomic review of weevils of the tribe Celeuthetini, mainly from Sundaland, Sulawesi and the Lesser Sunda Islands (Coleoptera, Curculionidae, Entiminae), Zootaxa 5464 (1), pp. 1-75 : 43-44

Nusasyntrophus echinatus (Heller) , n. comb.

Syntrophus echinatus (Heller) : Marshall, 1956: 35.

Heteroglymma echinata Heller, 1900: 20 . [incorrect original spelling]

Heteroglymma echinatum Heller, 1900: 20 . Type locality: Lombok, Sapit.

Trigonops bipusulosa Heller, 1940: 93–94 . Type locality: Lombok, Swela. syn. n.

Syntrophus bipusulosus ( Heller, 1940) : Marshall, 1956: 75.

Description. Heteroglymma echinatum Heller , lectotype, male. ( Fig. 24a ). Length. Pronotum plus elytron 5.00 mm. Integument black, largely covered with appressed, greyish-white scales partly abraded. Head with transverse angulate furrow separating rostral base, leading ventrad in front of eye; vertex with radial furrows laterally distinct. Eyes conically protruding. Rostrum 1.14 X longer than wide, sides subparallel; apical declivity with sunk-in scales, with sparse erect setae; dorsum simple. Pronotum 1.14 X longer than wide; with coarse punctures, with greyish-white scales partly abraded; with interspersed suberect claviform scales. Elytra 1.23 X as long as wide; densely covered with greyish-white scales; intervals with interspersed erect light brown scales, longer than interval´s width; interval 6 with small tubercle adorned with few erect dark brown scales. Genitalia. Penis ( Fig. 24b ) with sides of body subparallel to rounded apex. Apodeme 2.00 X as long as body of penis. Transfer apparatus curved. Intraspecific variation. Length. Pronotum plus elytron 4.50–5.80 mm. Female ( Fig. 24d ) with elytra more roundish. Female terminalia. Fig. 24e .

Material examined. Type specimens. Heteroglymma echinatum Heller , lectotype, by present designation, male: INDONESIA, Lombok, Sapit, 2000´, Mai-Juni 1896; labels ( Fig. 24c ); condition fair; genitalia in glycerol in microtube; DNA extraction ARC4030 (GenBank # OR557445) ( SMTD) . Paralectotypes: 2 exx, ARC4662 (DNA not sequenced) , same data as lectotype. 1 ex, Lombok , Fruhstorfer. Trigonops bipusulosa Heller , lectotype ( Fig. 24d ), by present designation, female: INDONESIA, Lombok, Swela, 22–28.IV-1927, leg. B. Rensch; labels ( Fig. 24f ); condition fair; genitalia in glycerol in microtube; DNA extraction ARC4026 (GenBank # OR557442) ( SMTD) . Other material: Lombok Isl. : 1 ex, ARC3556 (GenBank #) , Gn. Rinjani, Senaru, Rinjani-track, S08° 19.429´E116° 24.082´to S08° 20.780´E116° 23.790´, 800–1400 m, 21-23-III-2011; 33 exx, N Tetebatu, S08° 31.9' E116° 25.0', 600-700 m, 06-XII-2004 ( SMNK); 53 exx, N Tetebatu, Rinjani-track, ca . S08° 29.7' E116° 24.2', 800-1450 m, 07- XII-2004 ( SMNK); 1 ex, ARC3557 (GenBank #) , W Sengigi, Gn. Pusuk, S08° 27.899´E116° 04.372´, 200–600 m, 09-III-2011 ( SMNK) .

Notes. The species names N. echinatus (Heller) and N. bipusulosus (Heller) can be considered synonyms with greatest confidence. The type localities, Swela and Sapit, are located in the same area of East Lombok; in fact, the type specimens may come from the same patch of forest as their cox1 sequences differ in only one single position.

Figures

FIGURE 24. Nusasyntrophus echinatus (Heller), a–c, Heteroglymma echinatum Heller, lectotype: (a) Habitus (b) Penis (c) Labels; d–f, Trigonops bipusulosa Heller, lectotype: (a) Habitus (b) Terminalia (c) Labels.