Cybaeus adenes, Chamberlin & Ivie, 1932

Bennett, Robb, Copley, Claudia & Copley, Darren, 2019, Cybaeus (Araneae: Cybaeidae): the adenes species group of the Californian clade, Zootaxa 4711 (2), pp. 245-274: 247-248

publication ID

https://doi.org/10.11646/zootaxa.4711.2.2

publication LSID

lsid:zoobank.org:pub:06B09245-2722-4F4D-9076-41E68A17E8A7

persistent identifier

http://treatment.plazi.org/id/2F5B4D33-707D-FFA0-B289-446FFA5EFBB6

treatment provided by

Plazi

scientific name

Cybaeus adenes
status

 

Key to the species of the Cybaeus adenes  group

(males of C. pearcei  and C. torosus  unknown; females of C. adenes  and C. grizzlyi  are apparently morphologically indistinguishable)

1. Males............................................................................................. 2

- Females........................................................................................... 8

2(1). Proximal arm of tegular apophysis with single, pointed tip (Figs 12, 18, 25, 56, 61, 63); with ( Figs 25View FIGURES 25–27, 61–64View FIGURES 61–64) or without (Figs 12, 56) basal, pointed projection or proximally directed dorsal keel............................................ 3

- Proximal arm of tegular apophysis with tip bifid ( Figs 1View FIGURES 1–3, 44View FIGURES 44–46, 50View FIGURES 50–53, 59–60View FIGURES 56–60, 65, 67View FIGURES 65–68) or trifid ( Figs 35View FIGURES 35–37, 57View FIGURES 56–60); proximally directed dorsal keel present (but may be inconspicuous) ( Figs 57, 59–60View FIGURES 56–60, 65, 67View FIGURES 65–68)......................................... 5

3(2). Patellar apophysis with 40–50 peg setae covering dorsal surface (Fig. 13). Proximal arm of tegular apophysis without basal, pointed projection or dorsal keel ( Fig. 56View FIGURES 56–60). West central California: southern San Francisco Bay area to northern Monterey County ( Fig. 69View FIGURE 69)................................................................. amicus Chamberlin & Ivie 

- Patellar apophysis with about 25–30 peg setae dorsally or anterodorsally ( Figs 19View FIGURES 18–20, 26View FIGURES 25–27). Proximal arm of tegular apophysis with basal, pointed projection or dorsal keel ( Figs 61–64View FIGURES 61–64). West or east central California ( Fig. 70View FIGURE 70)................... 4

4(3). Proximal arm of tegular apophysis with simple untwisted tip and prominent, bluntly pointed dorsal keel ( Figs 61–62View FIGURES 61–64). West central California: Contra Costa and Alameda Counties ( Fig. 70View FIGURE 70)................................. grizzlyi Schenkel 

- Proximal arm of tegular apophysis with simple, slightly corkscrewed tip and dorsal keel a broad, bluntly pointed process dorsal to tip ( Figs 63–64View FIGURES 61–64). East central California: Placer and El Dorado Counties ( Fig. 70View FIGURE 70)..... auburn Bennett  spec. nov.

5(2). Patellar apophysis nearly lacking peg setae ( Fig. 36View FIGURES 35–37); proximal arm of tegular apophysis with broad, trifid tip and prominent dorsal keel ( Figs 57–58View FIGURES 56–60). West central California: Santa Clara County to Monterey County ( Fig. 70View FIGURE 70)............................................................................................... reducens Chamberlin & Ivie 

- Ten to 30 peg setae ( Figs 2View FIGURES 1–3, 51View FIGURES 50–53); proximal arm of tegular apophysis with narrower, bifid tip ( Figs 59–60View FIGURES 56–60, 65, 67View FIGURES 65–68), dorsal keel reduced ( Figs 59–60View FIGURES 56–60) or prominent ( Figs 65–68View FIGURES 65–68). West central California: north of Santa Clara County ( Figs 69–70View FIGURE 69View FIGURE 70)..... 6

6(5). Proximal arm of tegular apophysis with divergent tips ( Figs 59–60View FIGURES 56–60). West central California: Sonoma to San Francisco Counties ( Fig. 69View FIGURE 69).................................................................... adenes Chamberlin & Ivie 

- Proximal arm of tegular apophysis with convergent tips ( Figs 65, 67View FIGURES 65–68). West central California: San Mateo and Napa Counties ( Fig. 70View FIGURE 70)........................................................................................... 7

7(6). Tips of proximal arm of tegular apophysis larger, relatively strongly convergent, pincer-like ( Fig. 67View FIGURES 65–68); 13 peg setae anterodorsally on patellar apophysis ( Fig. 45View FIGURES 44–46). West central California: San Mateo County ( Fig. 70View FIGURE 70)............. sanbruno Bennett 

- Tips of proximal arm of tegular apophysis relatively weakly convergent, not pincer-like ( Fig. 65View FIGURES 65–68); about ten peg setae dorsally on patellar apophysis ( Fig. 51View FIGURES 50–53). West central California: Napa County ( Fig. 70View FIGURE 70)............. schusteri Bennett  spec. nov.

8(1). Atrium with lateral margins strongly curved antero-medially ( Figs 47–48View FIGURES 47–49); copulatory ducts contiguous (or nearly so) ( Figs 48–49View FIGURES 47–49). West central California: San Mateo County ( Fig. 70View FIGURE 70).................................... sanbruno Bennett 

- Atrium with lateral margins simple, not curved anteriorly ( Figs 4View FIGURES 4–8, 9View FIGURES 9–11, 15View FIGURES 15–17, 21View FIGURES 21–24, 30View FIGURES 28–30, 41View FIGURES 38–43); copulatory ducts widely separated ( Figs 6View FIGURES 4–8, 11View FIGURES 9–11, 17View FIGURES 15–17, 24View FIGURES 21–24, 29View FIGURES 28–30, 43View FIGURES 38–43). East or west central California ( Figs 69–71View FIGURE 69View FIGURE 70View FIGURE 71)............................................. 9

9(8). Atrium weakly to moderately concave and broad, length from epigastric groove to anterior margin <width between lateral margins ( Figs 38, 41View FIGURES 38–43, 21–22View FIGURES 21–24).......................................................................... 10

- Atrium strongly concave and narrower, length 1.2–3.0 times> width ( Figs 6View FIGURES 4–8, 15View FIGURES 15–17, 33View FIGURES 33–34, 54View FIGURES 54–55).......................... 11

10(9). Atrium at anterior margin of vulva ( Figs 38, 41View FIGURES 38–43); vulva compact, apparently strongly reduced ( Figs 39–40, 42–43View FIGURES 38–43). West central California: Santa Clara County to Monterey County ( Fig. 70View FIGURE 70)..................... reducens Chamberlin & Ivie 

- Atrium antero-medially located ( Figs 21–22View FIGURES 21–24); vulva “normal” ( Figs 22–24View FIGURES 21–24). East central California: Placer and El Dorado Counties ( Fig. 70View FIGURE 70)............................................................... auburn Bennett  spec. nov.

11(9). Atrium inverted U-shaped or lyriform, widest anteriorly ( Figs 4, 7View FIGURES 4–8, 9View FIGURES 9–11, 30View FIGURES 28–30); copulatory ducts smoothly curved posteriorly, not turning anteriorly at spermathecal heads ( Figs 6, 8View FIGURES 4–8, 28–29View FIGURES 28–30).................................................. 12

- Atrium inverted U- or vase-shaped (not lyriform), widest posteriorly ( Figs 15View FIGURES 15–17, 53–54View FIGURES 50–53View FIGURES 54–55); copulatory ducts smoothly curved posteriorly ( Figs 16–17View FIGURES 15–17) or turning anteriorly at spermathecal heads ( Figs 34View FIGURES 33–34, 55View FIGURES 54–55)................................ 13

12(11). California: north and west San Francisco Bay area ( Fig. 69View FIGURE 69).............................. adenes Chamberlin & Ivie   

- California: east San Francisco Bay area ( Fig. 70View FIGURE 70).............................................. grizzlyi Schenkel   

13(11). Atrium inverted vase-shaped, lateral margins convex ( Fig. 53View FIGURES 50–53). California: Napa County ( Fig. 70View FIGURE 70)................................................................................................. schusteri Bennett  spec. nov.

- Atrium inverted U-shaped, lateral margins concave ( Figs 15View FIGURES 15–17, 33–34View FIGURES 33–34, 54View FIGURES 54–55). California: San Francisco Bay area counties south of San Francisco and Berkeley to Monterey County ( Figs 69–71View FIGURE 69View FIGURE 70View FIGURE 71)............................................. 14

14(13). Copulatory ducts turning anteriorly at spermathecal heads ( Figs 34View FIGURES 33–34, 55View FIGURES 54–55); length of atrium (from epigastric groove to anterior margin) 2–3 times> width ( Fig. 33View FIGURES 33–34, 54View FIGURES 54–55). West central California: Alameda County ( Fig. 71View FIGURE 71)........................ 15

- Copulatory ducts smoothly curved posteriorly, not turning anteriorly ( Figs 16–17View FIGURES 15–17); length of atrium 1.2–1.6 times> width ( Fig. 15View FIGURES 15–17). West central California: southern San Francisco Bay area to northern Monterey County ( Fig. 69View FIGURE 69)............................................................................................ amicus Chamberlin & Ivie 

15(14). Copulatory ducts broad anteriorly (extending laterally beyond spermathecal bases), heavily sclerotized posteriorly ( Fig. 55View FIGURES 54–55); lateral margins clearly visible ventrally through integument of uncleared epigynum ( Fig. 54View FIGURES 54–55).... torosus Bennett  spec. nov.

- Copulatory ducts narrow anteriorly (not extending laterally beyond spermathecal bases), not heavily sclerotized posteriorly ( Fig. 34View FIGURES 33–34); not clearly visible through epigynal integument ( Fig. 33View FIGURES 33–34)........................ pearcei Bennett  spec. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Cybaeidae

Genus

Cybaeus