Quaesticulus Schaufuss, 1861

M, José, Costas, Salgado, Moreda, Lucía Labrada & Luque, Carlos G., 2010, A new cave-dwelling species of Quaestus Schaufuss, 1861 from the Montes de Pas, Northern Spain (Coleoptera: Leiodidae: Leptodirini), Zootaxa 2484, pp. 25-34 : 26

Subgenus Quaesticulus Schaufuss, 1861

The following characters define the subgenus Quaesticulus : apices of elytra rounded, not divergent; sutural striae of elytra long and visible; metafemora of males without small tooth on the posterior margin (except Q. adnexus Schaufuss, 1861 ); internal sac of the aedeagus without robust sclerotized pieces and with a stylet lacking the basal plate; spermatheca with a short spermathecal duct and spermathecal membranous gland. This subgenus can be divided into two groups based on the length / width ratios of antennomeres 8 and 11. The species are distributed between the two groups as follows:

adnexus group (9 species): Quaestus adnexus ( Schaufuss, 1861) , Q. angustitarsis ( Español, 1950) , Q. autumnalis ( Escalera, 1898) , Q. canis ( Salgado, 1992) , Q. cisnerosii (Pérez-Arcas, 1872) , Q. noltei ( Coiffait, 1965) , Q. pachecoi ( Bolívar, 1915) , Q. bustilloi Salgado & Fresneda, 2009 and Q. sharpi ( Escalera, 1898) .

filicornis group (3 species): Quaestus filicornis ( Uhagón, 1881) , Q. minos ( Jeannel, 1909) and Q. pasensis sp. nov.