Sánchez-Ruiz, Alexander, Brescovit, Antonio D. & Bonaldo, Alexandre B., 2020, Revision of the spider genus Nyetnops Platnick & Lise (Araneae: Caponiidae) with proposition of the new genus Nopsma, from Central and South America, Zootaxa 4751 (3), pp. 461-486: 474-475

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Nopsma  new genus

Type species: Nopsma juchuy ( Dupérré, 2014) 

Other species included. Nopsma enriquei  n. sp., N. armandoi  n. sp. and N. florencia  n. sp.

Etymology. The generic name is the equivalent in Latin to the expression “there’s no Nops  ”, analogously constructed as the negative substantives usually employed in the language Ocaina, Witoto linguistic family, still spoken in Peruvian and Colombian Amazonia. Gender neutral.

Diagnosis. Members of Nopsma  n. gen. can be distinguished from non-nopine genera by having subsegmented tarsi ( Figs 12KView FIGURE 12; 13View FIGURE 13 F–G) and from other two-eyed nopinae  , except Nyetnops  , by the absence of crista and arolium on anterior metatarsi ( Fig. 13AView FIGURE 13); they differ from Nyetnops  by the presence of a gladius and by the shape of endites, without projected outer sides on anterior margin ( Figs 12EView FIGURE 12; 15GView FIGURE 15; 17EView FIGURE 17; 19FView FIGURE 19); whereas in Nyetnops  the endites anterior margin are round and slightly projected ( Figs 1FView FIGURE 1; 2View FIGURE 2 I–J; 19D). In addition, males of Nopsma  are distinguished from Nyetnops  by the globose tegulum with elongated, prolaterally protruded embolus ( Figs 11View FIGURE 11 B–C; 15B–D; 17B– C, 18B–C). Females are presently known from a single species, but the straight proximal margin of receptaculum ( Fig. 16JView FIGURE 16) may be diagnostic, since in Nyetnops  the proximal margin is slightly concave on both known species ( Fig. 10View FIGURE 10 A–B. Also in the Nopsma  female the sclerotization around spiracles reach the anterior side of anterior spiracles ( Fig. 16JView FIGURE 16), a feature absent in both species of Nyetnops  known by females.

Description. Small caponiids with only two eyes ( Figs 11AView FIGURE 11; 15A, F, HView FIGURE 15; 17AView FIGURE 17; 18AView FIGURE 18). Carapace orange-brown with weak to remarkable pattern, broadly oval, almost sub-circular, widest at front of coxa II, anteriorly narrowed to less than half its maximum width ( Fig. 15FView FIGURE 15). Pars cephalica flattened behind ocular area; pars thoracica strong sloping posteriorly ( Fig. 12AView FIGURE 12); without submarginal depressions opposite intercoxal spaces; thoracic groove almost obsolete or absent. Anterior median eyes dark, situated on slightly elevated black tubercle, separated by about a half of its diameter, set back from anterior margin of clypeus by about twice their diameter ( Fig. 12View FIGURE 12 B–C). Chelicerae pale brown, with median lamina; most of distance between lamina and fang base occupied by white membranous lobe ( Fig. 12View FIGURE 12 H–I); cheliceral paturon with scattered, long, weak bristles; ectal side with stridulatory ridges ( Fig. 12JView FIGURE 12). Endites pale orange except for anterior tips, with a white membranous projection, wide, without projected outer sides on anterior margin ( Fig. 19FView FIGURE 19), convergent along midline, but not touching ( Figs 12EView FIGURE 12; 15GView FIGURE 15; 17EView FIGURE 17; 18EView FIGURE 18), covered with scattered long setae, and with strong distal serrula consisting of single tooth row ( Fig. 12View FIGURE 12 F–G). Labium pale orange, slightly pentagonal, with broad base, fused to sternum along posterior groove ( Figs 11FView FIGURE 11; 15GView FIGURE 15; 17EView FIGURE 17; 18EView FIGURE 18), reborded in apical part ( Fig. 12View FIGURE 12 E–F). Sternum orange, broadly oval, surface with fine reticular lines with numerous long, stiff setae ( Figs 11FView FIGURE 11; 15GView FIGURE 15; 17EView FIGURE 17; 18EView FIGURE 18); pleural membrane with two sclerotized intercoxal extensions between coxae II and III, and III and IV, long precoxal triangles on coxae II, III and IV ( Figs 11FView FIGURE 11; 15GView FIGURE 15; 17EView FIGURE 17; 18EView FIGURE 18). Legs orange, coxae pale orange, formula 4123, without spines; femora and tibiae very enlarged ( Fig. 13FView FIGURE 13); metatarsi entire, with dorsal metatarsal stopper ( Fig. 13HView FIGURE 13), anterior lacking crista but preserving the gladius ( Figs 12View FIGURE 12 K–L; 13A); all tarsi bisegmented ( Figs 12KView FIGURE 12; 13View FIGURE 13 F–G); tarsi with three claws; paired claws usually with 8–11 teeth, most distal of which are largest ( Fig. 13B, I, LView FIGURE 13); unpaired claws short on all legs ( Fig. 13B, C, IView FIGURE 13); ventral frictional setae on tarsi ( Fig. 13B, C, IView FIGURE 13) and several other setae around pretarsal claws. Tibiae, metatarsi, and tarsi with trichobothria in a single row, bases with semicircular rim bearing slight longitudinal ridges ( Fig. 13JView FIGURE 13), tarsal organ exposed, roundish, with marginal ring slightly pronounced ( Fig. 13KView FIGURE 13). Male and female palpal tarsus elongated, without claw, ventralprolateral surfaces densely covered with strong modified setae ( Figs 11BView FIGURE 11; 14AView FIGURE 14; 15CView FIGURE 15; 16AView FIGURE 16; 17BView FIGURE 17; 18BView FIGURE 18); with a pad of fine chemoreceptor setae on dorsal, distal part, with a tibial brush on the prolateral side of palp, near distal end ( Fig. 14BView FIGURE 14); pick on prolateral side of palpal femur, situated near proximal end of podomere ( Fig. 14CView FIGURE 14). Abdomen gray dorsally, lighter ventrally, with only slightly sclerotized epigastric and postepigastric scuta; abdominal pattern slightly outlined ( Figs 15A, HView FIGURE 15; 16AView FIGURE 16) or unnoticeable ( Fig. 18AView FIGURE 18); with two pairs of respiratory spiracles clustered around epigastric groove ( Fig. 15IView FIGURE 15); anterior spiracles leading to wide, short tracheal trunk ending in numerous long tracheoles; posterior spiracles leading to two large tracheal trunks extending anteriorly into prosoma, plus one or two much narrower trunk extending posteriorly for most of abdominal length and several short, small tracheoles extending posteriorly ( Fig. 15JView FIGURE 15). Six spinnerets in typical caponiid arrangement, ALS with one major ampullate gland spigot, PMS with five spigots, and PLS with multiple spigots. Male palpal tibia excavated ventrally ( Figs 14AView FIGURE 14; 16AView FIGURE 16); cymbium elongated, not swollen, tip pointed ( Figs 11View FIGURE 11 B–C; 14A; 15B–D; 16A; 17B–C); tegulum globose originating near base of cymbium ( Figs 11CView FIGURE 11; 15DView FIGURE 15; 17CView FIGURE 17; 18CView FIGURE 18), embolus elongated with a keel bordering the tip ( Figs 14View FIGURE 14 D–F; 16B–E). External female genitalia with anterior plate slightly sclerotized ( Fig. 15IView FIGURE 15); strongly sclerotized around spiracles and laterals extension ( Fig. 15IView FIGURE 15). Internal female genitalia consisting of transverse, anteriorly directed, slightly sclerotized receptaculum (re) with straight distal and proximal margins ( Fig. 16JView FIGURE 16), and a membranous uterus externus which usually extends anteriorly over the receptaculum ( Figs 15JView FIGURE 15; 16View FIGURE 16 G–J).

Distribution. Ecuador, Peru, Colombia and Nicaragua ( Fig. 18GView FIGURE 18).