Epanerchodus flagellifer, Golovatch, Sergei I., Mikhaljova, Elena V. & Chang, Hsueh-Wen, 2011

Golovatch, Sergei I., Mikhaljova, Elena V. & Chang, Hsueh-Wen, 2011, The millipede family Polydesmidae in Taiwan, with descriptions of five new species (Polydesmida, Diplopoda), ZooKeys 93, pp. 9-42 : 25-26

publication ID

https://dx.doi.org/10.3897/zookeys.93.1167

persistent identifier

https://treatment.plazi.org/id/2A4613F9-5AC9-F50F-75AD-7E4D730459D3

treatment provided by

ZooKeys by Pensoft

scientific name

Epanerchodus flagellifer
status

sp. n.

Epanerchodus flagellifer   ZBK sp. n. Figs 6168

Type material:

Holotype ♂ (NMNS-6561-001), Taiwan, Nantou County, Huisun timber land, 24.04.1998, leg. S.H. Wu. Paratypes: 1 ♂ (NMNS-6561-002), same locality, together with holotype; 1 ♂ (NMNS-6561-003), same locality, 04.1998; 1 ♂ (ZMUM), same locality, 25.04.1999; 1 ♂ (TFRI), same locality, 24.04.1999, all leg. S.H. Wu.

Name:

To emphasize the remarkably long, flagelliform distal part of the gonopod telopodite.

Diagnosis:

Differs from other Epanerchodus species in the distal half of the gonopod telopodite being particularly long, flagelliform, coupled with the presence of two rounded teeth at the base of the endomere (see also Key below). From the other congeners known from Taiwan, this new species differs also in the absence of sphaerotrichomes on ♂ legs.

Description:

Length of both sexes ca 8-11 mm; width of pro- and metazona varying between specimens from 0.8-1.0 to 1.4-1.5 mm, respectively. Holotype ca 9 mm long, and 0.9 and 1.4 mm wide on pro- and metazona, respectively. Coloration in alcohol pallid to light grey- to red-brown (Figs 61-65).

All characters as in Epanerchodus orientalis except as follows.

Antennae rather long and evidently clavate (Figs 61, 64), reaching behind segment 3 dorsally.

In width, collum <segment 2 <head = 3 = 4 <5-15(16) (Fig. 62), thereafter body gradually tapering towards telson (Fig. 63). Paraterga well-developed, starting from collum, set high, subhorizontal to very faintly upturned, mostly nearly level to a very faintly convex dorsum, only on collum and segment 2 lying clearly below dorsum; paraterga on collum small, subtriangular, a small lateral incision in front of a rounded caudal corner; front shoulders drawn forward and slightly convex only paraterga 2 and 3, straight on paraterga 4, thereafter increasingly well rounded and directed increasingly caudolaterad; caudal edge straight on segments 2-4, thereafter increasingly bisinuate and acutangular caudolaterally; starting from segment 10, caudal corners extending increasingly beyond rear tergal contour, spiniform on segments 17-19 (Figs 61-65). All poreless segments with three, all pore-bearing ones with four, small but evident incisions, each usually bearing a small seta on top at lateral margin. Metatergal sculpture typical, moderately developed, with three indistinct transverse rows of setiferous, polygonal bosses (Figs 62, 63). Tergal setae very short, mostly retained, a little longer only on collum and in rear row on metatergum 19. Stricture between pro- and metazona wide and smooth. Limbus very thin, microdenticulate. Epiproct rather short, conical (Fig. 65), only slightly bent ventrad, preapical papillae evident. Hypoproct semi-circular (Fig. 65); caudal, paramedian, setiferous papillae evident and well-separated.

Sterna without modifications, very densely setose. Legs rather long, slender, only slightly incrassate (Figs 61, 64-66), ca 1.7-1.8 times (♀) as long as midbody height, prefemora not swollen dorsally, sphaerotrichomes missing (Fig. 66).

Gonopods (Figs 67, 68) with large, subquadrate, medially fused coxae carrying only a few long setae ventrally. Telopodite simple and especially slender, prefemoral portion relatively short, endomere (en) flagelliform, with two small dentiform outgrowths at base; hairy pulvillus evident.

Remarks.

This small species is remarkable in showing a particularly long, flagelliform endomere, coupled with only two short outgrowths at its base.

In Taiwan, Epanerchodus flagellifer sp. n. occurs very locally, having been encountered only at a single locality (Map 2).