Turanena Lindholm, 1922

Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), Basteria 80 (1), pp. 5-30 : 10-12

Turanena Lindholm, 1922 ;

Subgenus Turanena Lindholm, 1922

Turanena Lindholm, 1922a: 275 . Type species (by original designation): Buliminus (Pseudonapaeus) herzi O. Boettger, 1889 .

The genus Turanena has a disjunct distribution: several species are found in Central Asia, whereas others are known from the Middle East (Israel, Lebanon, Iraq, northern Iran), Armenia, Turkey, and the southeastern islands of Greece. The species of Central Asia have been assigned to the subgenus Asuranena Schileyko & Moisseeva, 1995. These are T. (A.) albolimbata (Lind- holm, 1927); T. (A.) boamica Kuznetsov, 1999 ; T. (A.) cog- nata (Lindholm, 1927); T. (A.) conicula (E. von Martens, 1882) ; T. (A.) inversa Schileyko & Moisseeva, 1995 ; T. (A.) leptogyra (Lindholm, 1927) ; T. (A.) margaritae Schi- leyko & Moisseeva, 1989; T. (A.) martensiana ( Ancey, 1886) [synonyms: segregatus var. minor E. von Martens, 1874 ; liostracus Westerlund, 1896]; T. (A.) meshkovi Schi- leyko, 1984; T. (A.) stschukini (Lindholm, 1927) and T. (A.) tenuispira Schileyko, 1984 . The synonymy of liostra- cus with martensiana follows Sysoev & Schileyko (2009: 75). They attributed the name liostracus to Ancey (1886), but in reality the name has been introduced by Wester- lund (1896). The name martensiana is a nomen novum for Buliminus segregatus var. minor E. von Martens, 1874 , the latter being a preoccupied name.

The species outside Central Asia have been as- signed to the subgenus Turanena . So far, the following species are known:

T. (T.) albrechti Rähle, 1988 (Turkey: Vilayets Afyonkarahisar, Antalya, Burdur and Isparta)

T. (T.) andronakii ( Lindholm, 1913) (Turkey: Vilayets Artvin and Erzurum) [synonym: yusufelensis E. Gittenberger & Menkhorst, 1993 ]

T. (T.) benjamitica (Benson, 1859) (Israel; Lebanon) [synonym: enabensis Pallary, 1939]

T. (T.) bilgini Schütt & Şeşen, 2002 (Turkey: Vilayets Diyarbakır and Siirt)

T. (T.) carpathia (O. Boettger, 1885) (Greece: Karpathos, Rhodos)

T. (T.) cochlicopoides E. Gittenberger & Menkhorst, 1993 (Turkey: Vilayets Gaziantep and Osmaniye)

T. (T.) conelongata E. Gittenberger & Menkhorst, 1993 (Turkey: Vilayets Batman, Bitlis, Siirt and Van)

T. (T.) demirsoyi Gümüş & Neubert, 2012 (Turkey: Vilayet Erzincan)

T. (T.) forcartiana P. Schnell, 1979 (Turkey: Vilayets Adıyaman, Antalya, Çorum, Mersin, Sivas and Tokat)

T. (T.) hemmeni Bank & Butot, 1990 (Turkey: Vilayets Adana, Denizli, Mersin and Muğla; Greece: Samos)

T. (T.) hermonensis (Forcart, 1981) (Israel: Mount Hermon)

T. (T.) herzi (O. Boettger, 1889) (Armenia; Iran: provinces Azarbayjan-e-Gharbi, Azarbayjan-e-Sharqi, Kordestan, Kermanshah, Alborz and Semnan) [synonym: scalaris Nägele, 1902]

T. (T.) katerinae E. Gittenberger, 1996 (Greece: Crete, Cyprus)

T. (T.) macfadyeni (Pallary, 1939) (Iraq: Sinjar Mountain)

T. (T.) pseudobscura Bank & Neubert, 2016 (Iran: prov. Gilan)

T. (T.) tuccari E. Gittenberger, 1986 (Turkey: Vilayets Antalya and Isparta)

T. (T.) zilchi E. Gittenberger & Menkhorst, 1993 (Turkey: Vilayets Diyarbakır, Hakkâri, Siirt and Van).

The synonymy of enabensis with benjamitica follows Mienis (1977: 83, 84) and was confirmed by us; the synonymy of yusufelensis with andronakii was established by Gittenberger & Menkhorst (2006), and the syn- onymy of scalaris with herzi has been established by Bank & Neubert (2016: 14). The publication date of bilgini is 2002, not 2001 as Schütt (2010: 86) mentioned.

Although a description of bilgini appeared in the 2001 book of Schütt (: 86, 89 fig. b), being a year before the original description by Schütt & Şeşen in 2002, the 2001 description is not valid, as no holotype was selected or no syntypes were fixed, which is required for a zoological taxon described after 1999 (International Code of Zoological Nomenclature, Article 16.4).

Clearly, a major radiation of Turanena is seen in Turkey. Here we add another two new taxa to the fauna of Turkey. We are further aware of two new taxa from Lebanon and Iran, but these will be described elsewhere.