Chlaenius (Amblygenius) quadricolororientalis Dejean, 1826

Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid & Garner, Beulah, 2022, Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan, Zootaxa 5115 (4), pp. 451-486 : 458

Chlaenius (Amblygenius) quadricolororientalis Dejean, 1826

( Fig. 3c )

Chlaenius (Amblygenius) quadricolororientalis Dejean, 1826: 339 .

Material examined. PAKISTAN — Punjab • 1 ♀; Choa Saidan Shah, Minhala ; 31 May 2014; Mishkat leg.; PMNH • 1 ♂, 1 ♀, Attock, Akhori ; 29 May 2014; Mishkat leg.; PMNH • Layyah , Chak No. 127 ; 1 ♂, 1 ♀; 05 Aug. 2010; Adnan Bodhla leg.; PMNH • 1 ♂, 1 ♀; Islamabad, Sector E-12, Mazar ; 05 Feb. 2015; Mishkat leg.; PMNH • 1 ♀; Pitfall traps: PMNH ; 14 Apr. 2014; Mishkat leg.; PMNH • 1 ♂; same collection data as preceding; 21 Apr. 2014; Mishkat leg.; PMNH • 1 ♀; Islamabad ; 05 Jan.1983; Manzoor leg.; PMNH .— Khyber Pakhtunkhwa • 1 ♀; Swat, Charbagh (Pitfall Traps); 31 Aug.–02 Sep. 2015; Mishkat leg.; PMNH • 1 specimen; Dera Ismail Khan ; 12 Sep. 1988; Dr. M. Afzal leg.; PMNH • 1 ♀; Kohat ; 22 May 2007; Dr. Muhammad Ather Rafi leg.; NIM • 1 ♂; same collection data as preceding; 12 May 2007 • 1 ♂; same collection data as preceding; 14 Aug. 2007 • 1 ♂; same collection data as preceding; 11 Apr. 2007 • 1 ♂; same collection data as preceding; Apr. 2007 • 1 ♂; Karak ; 03 Aug. 2007; Dr. Muhammad Rafi leg.; NIM • 1 ♂; Peshawar, Pakistan Forest Institute; 18 Mar. 2009; Dr. Muhammad Ather Rafi leg.; NIM.— Gilgit Baltistan • 1 ♂; Chilas ; 07 Jun. 2007; at light; Dr. Muhammad Ather Rafi leg.; NIM .

Measurements. Male (n=10): TL= 15.07 mm, HW= 2.87 mm, PL= 3.36 mm, PW= 4.34 mm, EL= 9.04 mm, EW= 2.74 mm. Ratios PL/PW= 0.79, HW/PW= 0.65 and EL/EW= 3.30. Female (n=10): TL= 15.59 mm, HW= 2.90 mm, PL= 3.50 mm, PW= 4.44 mm, EL= 9.31 mm, EW= 2.90 mm. Ratios PL/PW= 0.78, HW/PW= 0.64 and EL/EW= 3.24

Diagnosis. Head metallic green with setiferous punctures. Antenna article III longer than article I+II; mentum median tooth emarginate at apex; labrum apex slightly emarginate. Labial and maxillary palpi terminal articles subcylindrical and truncate at apex. Pronotum metallic green, quadrate, glabrous with few punctures. Elytra black with a greenish tinge on striae, basal and lateral margins, without marks, intervals smooth and each stria with longitudinal row of indistinctly visible setiferous punctures. Legs reddish brown.

Comments. This species closely resembles Ch. (A.) atripes Chaudoir, 1876 in general body appearance except legs black and antennae dark brown, pronotum lateral margins slightly constricted toward base with latero-basal angles square in Ch. (A.) atripes while antennae and legs reddish brown, pronotum lateral margin not constricted towards base with latero-basal angle rounded and obtuse in Ch. (A.) quadricolor orientalis . This species is widespread in Asia. Although, subspecies orientalis occupies Asia’s western borders including Pakistan and India ( Kirschenhofer, 2004). In this current study, it was recorded from Choa Saidan Shah, Attock, Layyah, Islamabad, Swat, Dera Ismail Khan, Kohat, Karak, Peshawar and Chilas.

Distribution. Afghanistan, Bhutan, China, India (Uttar Pradesh), Nepal and Pakistan ( Chanu & Swaminathan 2017; Lorenz 2018).

Figures

FIGURE 3. Dorsal habitus (a) Chlaenius (Amblygenius) atripes Chaudoir, 1876 imaged from NHMUK collection; (b) Chlaenius (Amblygenius) luculentus Andrewes, 1920 imaged from NHMUK collection; (c) Chlaenius (Amblygenius) quadricolor orientalis Dejean, 1826 imaged from NHMUK collection; (d) Chlaenius (Chlaenius) lafertei Guérin-Méneville, 1843 imaged from NHMUK collection.