Pardosa armeniaca, Marusik, 2023

Marusik, Yuri M., 2023, A new species related to Pardosa atrata (Araneae, Lycosidae) from Armenia makes the distribution range of the atrata group disjunctive, ZooKeys 1180, pp. 225-235 : 225

Pardosa armeniaca sp. nov.

Figs 1 , 2 , 3A, B, E , 5A-C , 6

Types.

Holotype ♂ and paratype ♀ (ZMMU), Armenia, Gegharkunik Prov., Lake Sevan, env. of Tsovagyugh, Dzknaget River mouth, 40°36'53.4"N, 44°58'10.8"E, 1920 m, bog, 8.05.2021 (Y.M. Marusik).

Etymology.

The species name is derived from the terra typica.

Diagnosis.

The male of the new species differs from those of other species of the Pardosa atrata -group in having longer anterior arm of the tegular apophysis (cf. Fig. 3E, F, G ), relatively longer and more abrupt terminal apophysis directed antero retrolaterally (vs directed almost anteriorly) and different shape of the conductor (cf. Figs 2C, D , 3A, B ). The female of P. armeniaca sp. nov. is very similar to that of P. atrata but differs in having relatively short, straight, and almost parallel receptacles (vs longer, converging, and roundly bent; cf. Figs 4F, I , 5B, C ).

Male. Total length 7.5. Carapace 4.1 long, 2.85 wide. Dark coloured. Carapace dark brown, with thin, light-brown median stripe divided by dark median line, with partly broken submarginal light stripe thinner than dark marginal stripe; cephalic light spot absent. Sternum black. Abdomen almost uniformly dark brown. Legs lacking annulation, coxae-tibiae dark brown, metatarsi and tarsi brown.

Leg lengths:

Palp as in Figs 1C, D , 2 , 3A, B, E ; dark brown with light retrolateral stripe on femur; femur 2.7 times longer than wide; patella 1.3 times longer than wide, tibia almost as wide as long; cymbium as long as femur (in lateral view), tip with 1 claw; bulb 1.3 times longer than wide, tegular apophysis (Tg) large, with long anterior arm, ca. as long as palea; terminal apophysis (Ta) claw-like (in lateral view) and stump-like in anterior view; conductor (Co) massive with rounded notch on tip; embolus (Em) straight in ventral view, roundly bent in anterior; tip not modified.

Female. Total length 8.0. Carapace 4.1 long, 3.15 wide. Colouration as in male, with wider submarginal stripes as wide as marginal stripes. Legs and leg segment length as shown in table below. Femur I with 2 prolateral spines, femur II with 1; tibiae I-IV with 3 pairs of ventral spines including apical; metatarsus I with 3 pairs of ventral spines including apical.

Leg lengths:

Epigyne as in Fig. 5A-C ; epigynal plate ca. 1.3 times wider than long; fovea trapezoidal; ca. 2 times wider than long; septum as long as fovea wide, rounded margin of septal base absent; septal wings (Sw) small, ca. halflength of septal stem (Ss). Receptacles (Re) long (>10 times longer than wide) and straight, almost parallel to axis of epigyne, anterior part slightly widened.

Figures

Figure 1. Pardosa armeniaca sp. nov. A paratype female B-D holotype male A, B habitus, dorsal C whole palp, retrolateral D terminal part of palp, ventral. Scale bars: 0.2 mm, if not otherwise indicated.

Figure 2. Bulb of Pardosa armeniaca sp. nov. A ventral B retro-ventral C anterior D embolic division, ventral. Abbreviations: Co - conductor, Em - embolus, Ta - terminal apophysis, Tg - tegular apophysis. Scale bars: 0.2 mm.

Figure 3. Embolic division (A-D) and tegular apophysis (E-G) of Pardosa armeniaca sp. nov. (A, B, E), P. atrata (C, F from Polar Ural) and P. fuscula (D, G from Washington State). A, C-G ventral B anterior. Abbreviations: Co - conductor, Em - embolus, Te - terminal apophysis. Scale bars: 0.2 mm.

Figure 4. Epigyne of Pardosa armeniaca sp. nov., P. atrata, and P. fuscula A-C P. armeniaca sp. nov. D-F P. atrata G-I P. fuscula A, B, D, E, G, H ventral C, F, I dorsal D from Utsjoki, Finland E from Yamalo Nenets Okrug, Russia G, H from Washington State. Abbreviations: Bm - rounded margin of septal base, Re - receptacle, Sb - septal base, Ss - septal stem, Sw - septal wing. Scale bars: 0.2 mm.

Figure 5. Embolic division and epigyne in Pardosa atrata and P. fuscula A, B P. atrata from Oulu C, D P. fuscula from Washington State A, C embolic division, ventral B, D epigyne, ventral. Abbreviations: Co - conductor, Em - embolus, Te - terminal apophysis. Scale bars: 0.1 mm.

Figure 6. Distribution map with records of Pardosa armeniaca sp. nov. (red square), P. atrata (blue dot), P. fuscula (grey diamond), the type locality of P. narymica (blue triangle), doubtful records of P. atrata (question mark), and Chinese records of P. atrata that may belong to other species (pink dots).