Thinophilus cataractae, Grichanov, 2023

Grichanov, Igor Ya., 2023, A review of the Afrotropical Thinophilus Wahlberg, 1844 (Diptera: Dolichopodidae), with the descriptions of ten new species, European Journal of Taxonomy 878, pp. 1-52 : 38-41

publication ID 10.5852/ejt.2023.878.2153

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Thinophilus cataractae

sp. nov.

Thinophilus cataractae sp. nov.

Fig. 13 View Fig


Thinophilus cataractae sp. nov. keys to T. fluvialis sp. nov., strongly differing from the latter in morphology of hypopygium, almost entirely black antenna, and black-brown palp on basal half. T. fluvialis male has antenna black dorsally, yellow ventrally; palp entirely yellow (see key above).


The specific epithet ‘ cataractae ’(in Latin) refers to the Niagarakely waterfalls locality in the Niagarakely Forest, where the male type was collected.

Material examined

Holotype MADAGASCAR • ♂; “ Niagarakely Forest [19°09′ S, 48°13′ E]; Moramanga Distr.; Dec. 1955; B. Stuckenberg leg.; Thinophilus tinctus Parent ; P. Vanschuytbroeck det. 1957; I.R.Sc.N.B. I.G. 20938”; RBINS. GoogleMaps

Paratype MADAGASCAR • 1 ♂; same collection data as for holotype; RBINS. GoogleMaps


Male ( Fig. 13A View Fig )

MEASUREMENTS. Body length 3.8 mm; antenna length 1 mm; wing length 3.5 mm; wing width 1.1 mm.

HEAD ( Fig. 13B View Fig ). Postcranium bluish black, grey pollinose; frons and face bluish black; clypeus black, grey pollinose; face under antennae about 2 × as wide as height of postpedicel; clypeus about ⅔ as long as epistoma, about as wide as long; palp black-brown on basal half, yellow distally, bearing black bristly hairs; proboscis dark brown; 2 diverging ocellars; 1 vertical, 1 postvertical, somewhat stronger and longer than, and not in row with upper postoculars; upper postoculars uniseriate, black; lower postoculars multiseriate, white, long; antenna black; scape with scale-like inner projection (spine-like from dorsal view; Fig. 13D View Fig ), as long as scape; pedicel simple, convex on inner side; postpedicel rounded, with short pubescence, as high as long; arista-like stylus dorsal, black and thick basally, thin distally, shortly pubescent; length ratio of scape to pedicel to postpedicel to stylus, 0.07/0.07/0.10/0.84.

THORAX. Metallic, grey dusted; mesonotum black, without matt spots; pleura bluish black; no acrostichals; 5 dorsocentrals of almost equal length; scutellum with 2 strong marginals and 2 minute laterals; 1 dark upper and 1–2 white lower propleural bristles of different length.

LEGS. Fore coxa brown; mid and hind coxae black; legs mostly dirty yellow, darker at knees; tarsi black from tip of basitarsus

FORE LEG. Coxa with long black setae and apical bristles; femur simple, with some short fine ventral setae at base and 3–4 posteroventral bristles at apex; tibia bearing 2 anterodorsal, 2 posterodorsal bristles and 3–4 short apical setae; tibia and tarsus ventrally with elongate simple setulae; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 1.05/1.04/ 0.47/0.22/0.18/0.13/0.16.

MID LEG. Coxa with black setae and 1 bristle; femur ( Fig. 13C View Fig ) with anteroventral row of short dark hairs on basal half; 1 preapical anterior and 4–5 preapical posteroventrals; tibia bearing 2 anterodorsal; 2 posterodorsal, 4 apical bristles; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 1.3/1.39/0.8/0.34/0.2/0.13/0.16.

HIND LEG. Coxa with 1 black exterior bristle; femur ventrally without remarkable setae; 3 anterodorsal bristles; tibia bearing 2 anterodorsal, 2 posterodorsal bristles, few very short ventrals, 4 apicals; segment 5 inconspicuously widened; length of femur, tibia and tarsal segments (in mm): 1.41/1.55/0.46/0.41/0. 28/0.17/0.18.

WING ( Fig. 13E View Fig ). Fumose, without dark spots; veins brown; distal part of M 1+2 straight; tip of R 4+5 parallel with M 1+2; ratio of part of costa between R 2+3 and R 4+5 to that between R 4+5 and M 1+2 (in mm), 0.48/0.25; crossvein dm-m straight; ratio of dm-m to distal part of M 4, 0.32/0.30; anal vein distinct; halter yellow; lower calypter yellow, with white cilia.

ABDOMEN. Black, grey dusted; tergites 2–4 laterally whitish pollinose; setae and hind-marginal bristles black, short; sternites with short setae. Hypopygium ( Fig. 13F View Fig ) black with black appendages; epandrial lobe fingerlike, with strong apical bristle; hypandrium fused with epandrium, short, apically concave; phallosoma broad, acute apically, projected; phallus coiled, long and flat, band-like; surstylus relatively short and narrow (lateral view), not reaching apex of phallosoma ( Fig. 13G View Fig ), with 2 long wormlike ventral processes and 2–3 short spine-like apicoventral setae; cerci dorsally widely separated, elongate-ovate, with narrow finger-like apex, covered with long outer bristles.




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