Gomphostemma longipetalum 3ongch

Bongcheewin, Bhanubong, Ingrouille, Martin J. & Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), Kew Bulletin 77 (1), pp. 27-92 : 45

6. Gomphostemma longipetalum 3ongch . sp. nop.

Type: Burma, Kachin, Mountains East of Fort Hertz, 25°30&N 95°30&E, 600 – 900 m [2000 – 3000 ft], 3 Sept. 1926, Ward 5339 (holotype K [K001081923]).

http://www.ipni.org/urn:lsid:ipni.org:names:55221335-1

Erect herb, up to 0.6 m tall. Stems ascending, woody at nodes, obtusely quadrangular, with longitudinal grooves, tomentose with dense stellate hairs with a 1- celled stalk or sessile stellate hairs, dendroid hairs absent; internodes straight. Leaoes petiolate, spreading, chartaceous; blades elliptic or elliptic-ovate, 10 – 15 × 3 – 5 cm, apex acute, margin shallowly serrate, base attenuate, upper side coarse with 1 – 3-celled simple hairs, bifurcate hairs or stellate hairs, lower side pale green tomentose with dense stellate hairs; petioles 10 – 13 mm long. Inflorescence axillary, with opposite cymes sessile, unbranched then congested with axis not visible between flowers, exceeding 20 mm wide when pressed, excluding corolla, inserted at the upper nodes at which leaves are usually present; verticils manyflowered; bracts elliptic, 6 – 10 × 2 – 3 mm, apex acute, margin serrate, base attenuate, upper side with simple hairs and bifurcate hairs, lower side tomentose with stellate hairs; bracteoles narrowly elliptic, sometimes falcate or ensiform, 6 – 8 × 1 – 1.3 mm. Flowering calyx infundibular, 8 – 10 mm long, 3-ridged, outside with dense stellate hairs, inside tomentose with adpressed short simple hairs; tube 2 – 3 mm long; lobes 2 – 3 mm long, apex filiform. Fruiting calyx not seen. Corolla yellow, 20 – 30 mm long, throat narrow, not enlarged, outside tomentose with sessile stellate hairs and glandular-like simple hairs with a multi-celled stalk, inside glabrous; tube straight, 32 – 22 mm long, exserted from the calyx; posterior lobe hood-like, 6 – 8 × 3 – 3 mm; anterior lobes spreading; median lobe 3 – 5 mm long. Staminal filaments slender. Disk lobed, tumescent behind. Style slender with apex unequally lobed. Nutlets not seen. Fig. 5 .

RECOGNITION. Gomphostemma longipetalum is distinguished from the straight corolla tube species of Gomphostemma in having a much longer corolla, 20 – 30 mm long, calyx lobes not longer than the tube and cymes inserted at the lower nodes where the leaves have fallen. This species could be confused with G. crinitum because of its straight corolla tube and narrow corolla throat but that species

has a shorter corolla, 20 – 30 mm long, and calyx lobes much longer than the tube.

DISTRIBUTION. Myanmar. Map 2 .

SPECIMENS EXAMINED. MYANMAR. Kachin: mountains E of Fort Hertz, 25°30&N 95°30&E, 3 Sept. 1926, Ward 5339 (holotype K).

HABITAT. In mountains; alt. 600 – 900 m.

CONSERVATION STATUS. Gomphostemma longipetalum is known only from the type specimen collected in 1926. Although the label mentioned that this species is ‘fairly abundant at one spot’, the?ata?eficient (??) category is applied due to the lack of further records.

PHENOLOGY. Flowering September.

NOTES. The epithet longipetalum [longi- (in Greek compound), long; -petalum, petal] refers to its corolla length.

Figures

Map 2. Distribution range of Gocphostecca longipetaluc, G. niveuc, G. parvifloruc and G. thocsonii.

Fig. 5. Morphological variation of Gocphostecca javanicuc. A – G leaves, A froc Ridley s.n.; B froc Ridley s.n.; C froc Edaño 17836; D froc Middleton et al. 1307; E froc Burkill & Holttuc 8630; F froc Craiĸ 733; G froc Tsang 27020. H – P calyces, H froc Henry 12011; J froc Craiĸ 733; K froc Middleton et al. 1307; L froc Tsang 27020; M froc Larsen et al. 30833; N froc Kerr 2034; P froc Gardner & Khucchocpoo ST 2117. NB. 4 outer surfaces area and 1 inner surface (on right) shown for each calyx. Only 1 outer surface (on left) showing inducentuc, other 3 outer surfaces without hairs for clarity. Inner surface shows inducentuc. Q – S verticils, Q froc Hooker & Thocson s.n.; R froc Middleton et al. 1307; S froc Burkill & Holttuc 8630. T – U type of hairs, T stellate hairs with a culti-celled stalk, froc Gardner & Khucchocpoo ST 2117; U stellate hairs with a 1-celled stalk, froc Kerr 21078. DRAWN;Y ANDREW;ROWN.