Mitrella elongatissima, Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard M., 2021, The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group, Zootaxa 5025 (1), pp. 1-75 : 24-25

publication ID

https://doi.org/ 10.11646/zootaxa.5025.1.1

publication LSID

lsid:zoobank.org:pub:35E6C8FA-4078-4C53-9B74-F9618D5E6E60

persistent identifier

https://treatment.plazi.org/id/256F7D50-E27C-FFF8-ACD6-37AAFAD2B3F1

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Plazi (2021-08-24 19:56:49, last updated 2023-11-04 17:29:29)

scientific name

Mitrella elongatissima
status

sp. nov.

Mitrella elongatissima nov. sp.

Figs 4C, 9A View FIGURE 9 1 –A View FIGURE 1 2 View FIGURE 2 , B 1 –B View FIGURE 1 2 View FIGURE 2 , C 1 –C View FIGURE 1 2 View FIGURE 2 , D

[ Mitrella ] Petersi nov. form.—Hoernes 1880: 126 (pars) [nomen nudum].

Columbella (Mitrella) Petersi nov. form.— Hoernes & Auinger 1880: 97 (pars), pl. 12, figs 5–6 [non Mitrella petersi Hilber, 1879 ].

Atilia (Macrurella) Petersi Hoernes et Auinger — Cossmann 1901: 245. [non Mitrella petersi Hilber, 1879 ].

Columbella (Mitrella) petersi R. Hö. View in CoL — Boettger 1902: 17 [non Mitrella petersi Hilber, 1879 ].

Columbella (Mitrella) petersi Hö. Au. View in CoL — Boettger 1906: 15 [non Mitrella petersi Hilber, 1879 ].

Mitrella petersi Hörn u. Auing.— Meznerics 1933: 340 [non Mitrella petersi Hilber, 1879 ].

M.[itrella]. (A [tilia].) petersi (R. Hörn. et Au.) — Sieber 1958a: 147 [non Mitrella petersi Hilber, 1879 ].

Mitrella (Atilia) petersi (R. Hörn. Au.) — Sieber 1958b: 148 [non Mitrella petersi Hilber, 1879 ].

Mitra bronni Mich. — Grković 1963: 120, pl. 1, fig. 6 [non Michelotti, 1847].

Type material. Holotype: NHMW 2020 View Materials /0162/0001, Steinebrunn, SL: 11.5 mm, MD: 3.1 mm, illustrated in Hoernes & Auinger (1880, pl. 12, figs 6a–c) as Columbella (Mitrella) Petersi , Figs 4C, 9A View FIGURE 9 1 –A View FIGURE 1 2 View FIGURE 2 . Paratypes: NHMW1846 View Materials /0037/0157, SL: 11.4 mm, MD: 3.4 mm, Steinebrunn , illustrated in Hoernes & Auinger (1880, pl. 12, figs 5a–c) as Columbella (Mitrella) Petersi , Fig. 9D View FIGURE 9 . NHMW 2020 View Materials /0162/0002, SL: 11.2 mm, MD: 3.1 mm, Steinebrunn ( Austria), Figs 9B View FIGURE 9 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 2020 View Materials /0162/0003, SL: 12.8 mm, MD: 3.8 mm, Steinebrunn ( Austria), Figs 9C View FIGURE 9 1 –C View FIGURE 1 2 View FIGURE 2 .

Additional material. NHMW1862 View Materials /0001/0373, 13 spec. Gainfarn ( Austria) . NHMW1871 View Materials /0011/0005, 270 spec., Steinebrunn ( Austria) . NHMW 1884 View Materials /0037/2898 (=A1553), 10 spec., Steinebrunn ( Austria) .

Type locality. Steinebrunn ( Austria), Vienna Basin .

Type stratum. Sandy silt of the Baden Formation.

Age. Middle Miocene, Badenian.

Etymology. Referring to the elongate slender shell.

Diagnosis. Mitrella species of moderately small size, very slender fusiform profile, high conical paucispiral protoconch, high, acute spire, and narrow aperture, with prominent denticles on thickened outer lip, D2 (second from top) stronger. Distinguished from other Paratethyan species by its extraordinarily elongate shape.

Description. Shell moderately small, very slender fusiform, with high spire, and narrowly incised suture. Protoconch paucispiral, high conical, of about two whorls. Teleoconch of nine whorls. Spire whorls high, flat-sided, with periphery at abapical suture. Last whorl high, moderately convex, rounded periphery, weakly contracting, high base. Aperture moderately narrow. Shell surface smooth, glossy; weak spiral cords on abapical portion of base and siphonal fasciole. Outer lip thickened with mid-labial callus pad; about eight prominent denticles, with D2 strongest. Anal canal shallow, U-shaped. Siphonal canal mid-length, narrow with shallow notch. Columella slightly twisted. Columellar callus not thickened, forming thin, narrow rim, weakly delimited from base. No columellar denticles. Parietal callus weakly developed.

Shell measurements and ratios. SL: 11.2–12.8 mm, MD: 3.1–3.8 mm, AA: 17–21°, SL/MD: 3.4–3.8, AL/AW: 3.8–4.3, LWH/AH: 1.2–1.4.

Discussion. Hoernes &Auinger (1880) confused this species with Mitrella petersi ( Hilber, 1879) .All subsequent authors followed the misidentifications by Hoernes & Auinger (1880). A comparison with the lectotype of Mitrella petersi reveals little similarity between the species. Mitrella petersi is much smaller, has a much wider apical angle, and a shorter spire. Moreover, it has a low conical protoconch, whereas the protoconch of M. elongatissima nov. sp. is turreted. For comparison with M. demaintenonae nov. sp., see above.

Mitrella liguloides Sacco, 1890 , from the Tortonian of Stazzano ( Italy), might represent a closely related species, but is twice the size. It has a very similar spire and an identical narrow aperture, but is slightly less slender, bears columellar denticles and has a deeply-incised siphonal notch (see Sacco 1890; Ferrero Mortara et al. 1981, pl. 58, figs 13a–b). Mitrella teres ( Bellardi, 1848) , from the Pliocene of Italy, is reminiscent of M. elongatissima in its high and slender spire, but differs in its convex whorls, higher last whorl, and longer siphonal canal (see Ferrero Mortara et al. 1981, pl. 57, figs 7a–b; Chirli 2002, pl. 6, figs 7–12).

Palaeoenvironment. The frequent occurrence at Steinebrunn suggests shallow, inner neritic environments with sea grass (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn ( Austria) ( Sieber 1958b; own data); southern Pannonian Basin: Tešanj ( Bosnia-Herzegovina) ( Grković 1963).

Bellardi, l. (1848) Monografia delle Columbelle fossili del Piemonte. Accademia delle scienze di Torino, Dalla Stamperia reale, 22 pp. [https: // www. biodiversitylibrary. org / item / 121649 # page / 3 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 59632

Boettger, O. (1902) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. (Mit einem Situationsplan der Fundpunkte). II. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 51, 1 - 200. [https: // www. biodiversitylibrary. org / item / 109496 # page / 283 / mode / 1 up]

Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]

Chirli, C. (2002) Malacofauna Pliocenica Toscana, 3. Superfamiglia Muricoidea-Cancellarioidea. C. Chirli, Firenze, 92 pp.

Cossmann, M. (1901) Essais de Paleoconchologie Comparee. Livraison 4. Cossmann, Lamarre &, C., Paris, 293 pp., 10 pls. [https: // www. biodiversitylibrary. org / item / 111984 # page / 11 / mode / 1 up]

Ferrero Mortara, E., Montefameglio, L., Pavia, G. & Tampieri, R. (1981) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte I. Museo Regionale di Scienze Naturali, Catalogo, 6, 1 - 327. [https: // www. biodiversitylibrary. org / item / 269998 # page / 5 / mode / 1 up]

Grkovic, Z. B. (1963) Nalazak faune marinskog torton u okolini Tesnja (Severna Bosna). Geoloski glasnik, 8, 117 - 122.

Hilber, V. (1879) Neue Conchylien aus den mittelsteierischen Mediterranschichten. Sitzungsberichte der kaiserlichen Akademie der Wissenschaften, der Mathematisch-Naturwissenschaftlichen Classe, 79, 416 - 464. [https: // www. zobodat. at / pdf / SBAWW _ 79 _ 0416 - 0464. pdf]

Hoernes, R. & Auinger, M. (1880, 1890) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran-Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls 7 - 12 (1880), 233 - 282, pls. 29 - 36 (1890). [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]

Meznerics, I. (1933) Die Minutien der tortonischen Ablagerungen von Steinabrunn in Niederosterreich. Annalen des Naturhistorischen Museums Wien, 46, 319 - 359. [https: // www. zobodat. at / pdf / ANNA _ 46 _ 0319 - 0359. pdf]

Michelotti, G. (1847) Description des fossiles des terrains miocenes de l'Italie septentrionale. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem, Series 2, 3 (2), 1 - 408. [https: // books. google. be / books? id = aQVqpL 58 HrsC & ots = rhEPJ _ C 6 hy & dq = Michelotti % 2 C % 20 G. % 091847 % 09 Description % 20 des % 20 fossiles % 20 de s % 20 terrains % 20 mioc % C 3 % A 8 nes % 20 de % 20 l'Italie % 20 septentrionale. & lr & pg = PP 5 # v = onepage & q = Michelotti, % 20 G. % 091847 % 09 Description % 20 des % 2]

Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Sieber, R. (1958 b) Die Tortonfauna von Steinabrunn bei Drasenhofen (Bez. Mistelbach, N. - O.). Verhandlungen der Geologischen Bundesanstalt, 1958, 142 - 155. [https: // opac. geologie. ac. at / ais 312 / dokumente / VH 1958 _ 142 _ A. pdf]

Gallery Image

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

Gallery Image

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width.

Gallery Image

FIGURE 9 A1–A2. Mitrella elongatissima nov. sp., NHMW 2020/0162/0001, holotype, Steinebrunn (Austria). B1–B2. Mitrella elongatissima nov. sp., NHMW 2020/0162/0002, paratype, Steinebrunn (Austria). C1–C2. Mitrella elongatissima nov. sp., NHMW 2020/0162/0003, paratype, Steinebrunn (Austria). D. Mitrella elongatissima nov. sp., NHMW1846/0037/0157, paratype, Steinebrunn (Austria). E1–E2. Mitrella demaintenonae nov. sp., NHMW1878/0001/0081, paratype, Möllersdorf; F1–F2. Mitrella demaintenonae nov. sp., NHMW2020/0162/0002, Coşteiu de Sus (Romania). G1–G2. Mitrella demaintenonae nov. sp., NHMW 2020/0161/0001, holotype, Coşteiu de Sus (Romania). H. Mitrella demaintenonae nov. sp., NHMW1867/ 0019/0049b, Coşteiu de Sus (Romania).

Kingdom

Plantae

Phylum

Tracheophyta

Class

Magnoliopsida

Order

Magnoliales

Family

Annonaceae

Genus

Mitrella