Auingeria bellardii (Hoernes, 1852) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard M., 2021, The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group, Zootaxa 5025 (1), pp. 1-75 : 50-51

Auingeria bellardii (Hoernes, 1852) nov. comb.

Figs 4Q, 15A 1 –A 2 , B 1 –B 2 , C, D

* Columbella Bellardii Hörnes —Hörnes 1852: 123, pl. 11, figs 1a–c.

Columbella (Anachis) Bellardii M. Hoernes — Hoernes & Auinger 1880: 99, pl. 11, figs 22a–c.

[ Columbella (Anachis) Bellardii M. Hoernes ] Varietät A— Hoernes & Auinger 1880: 100, pl. 11, figs 23–24.

Columbella Bellardii R. Hoernes et Auinger — Kittl 1887: 248.

A.[nachis]. (A [nachis].) bellardii (Hörn.) — Sieber 1958a: 147.

Columbella (Anachis) bellardii Hörnes, 1856 — Strausz 1966: 297 (pars), pl. 34, figs 18–19.

? Anachis (Anachis) bellardii (M. Hoernes) — Steininger 1973: 429.

Columbella bellardii Hörnes, 1852 — Landau et al. 2019: 170.

Type material. Holotype: NHMW1851 /0013/0007, SL: 15.9 mm, MD: 6.3 mm, Baden ( Austria), illustrated in Hörnes (1852, pl. 11, figs 1a–c), Figs 15A 1 –A 2 .

Additional material. NHMW1866 /0001/0673, SL: 11.8 mm, MD: 5.1 mm, Baden ( Austria) , illustrated in Hoernes & Auinger (1880, pl. 11, figs 22a–c), Fig. 15D . NHMW1949 /0005/0008, SL: 16.7 mm, MD: 6.6 mm, Forchtenau ( Austria) , illustrated in Hoernes &Auinger (1880, pl. 11, figs 23a–c), Fig. 15C . NHMW1853 /0003/0095, SL: 17.8 mm, MD: 7.2 mm, Forchtenau ( Austria) , illustrated in Hoernes & Auinger (1880, pl. 11, figs 24a–c), Figs 4Q, 15B 1 –B 2 . NHMW1870 /0053/0020, 77 spec., Forchtenau ( Austria) ; NHMW1857 /0030/0006, 3 spec., Rohrbach ( Austria) ; NHMW1860 /0010/0002, 2 spec., Sopron ( Hungary) .

Description. Shell medium sized, moderately broad, with gradate spire and subcylindrical whorls. Protoconch conical multispiral of 3.5 convex whorls. Teleoconch of seven whorls. First teleoconch whorl moderately high, smooth. Prominent, orthocline, broad, convex axial ribs on second and subsequent teleoconch whorls. Adsutural spiral cord with indistinct nodes separated by delicate spiral groove on second teleoconch whorl. Later whorls with four spiral cords, overriding axial ribs, forming cancellate sculpture, with weak nodes developed at intersections. Last whorl moderately high, subcylindrical, distinctly constricted at base; axial sculpture weakening over base, strong convex spiral cords on siphonal fasciole. Aperture moderately narrow. Outer lip weakly thickened with about 10 lirae extending deep within aperture. Anal canal incised, V-shaped. Siphonal canal long, narrow, slightly deflected. Columellar callus thickened forming sharply delimited, adherent callus rim, with about five weak columellar denticles. Parietal callus weak.

Shell measurements and ratios. SL: 11.8–17.8 mm, MD: 5.1–6.3 mm, AA: 35–45°, SL/MD: 2.5–2.6, AL/AW: 3.8–4.1, LWH/AH: 1.3.

Discussion. This species was based by Hörnes (1852) on a single juvenile specimen from Baden ( Austria), with prominent sculpture of nodes. In adult shells the nodes are weaker and the last whorl displays a cancellate sculpture. It is separated from its congeners by the presence of axial sculpture.

Palaeoenvironment. Middle neritic to outer neritic. The specimens from the Vienna Basin are found in clay of the Baden Fm., suggesting about 250 m water depth ( Hohenegger et al. 2008).

Distribution in Central Paratethys.? Ottnangian (early Miocene): Kaltenbachgraben (Bavaria) ( Steininger 1973). Badenian (middle Miocene): Carpathian Foredeep: Ostrawa (the Czech Republic) ( Kittl 1887); Vienna Basin: Baden ( Austria); Eisenstadt-Sopron Basin: Sopron ( Hungary) ( Strausz 1966).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width.

FIGURE 15. A1–A2. Auingeria bellardii (Hoernes, 1852), NHMW1851/0013/0007, holotype, Baden (Austria). B1–B2. Auingeria bellardii (Hoernes, 1852), NHMW1853/0003/0095, Forchtenau (Austria). C. Auingeria bellardii (Hoernes, 1852), NHMW1949/0005/0008, Forchtenau (Austria). D. Auingeria bellardii (Hoernes, 1852), NHMW1866/0001/0673, Baden (Austria).E1–E2. Auingeria grussbachensis (Csepreghy-Meznerics, 1969), NHMW1999z0076/0005, Hrušovany nad Jevišovkou (the Czech Republic). F1–F2. Auingeria grussbachensis (Csepreghy-Meznerics, 1969), NHMW1999z0076/0005, Hrušovany nad Jevišovkou (the Czech Republic). G1–G2. Auingeria grussbachensis (Csepreghy-Meznerics, 1969), NHMW1999z0076/0004, Sedlec (Porzteich) (the Czech Republic). H1–H2. Auingeria austriaca (Hoernes & Auinger, 1880), NHMW1863/0015/0628b, paralectotype, Niederleis (Austria). I1–I2. Auingeria austriaca (Hoernes &Auinger, 1880), NHMW1863/0015/0628a, lectotype, Niederleis (Austria). J1–J2. Auingeria austriaca (Hoernes & Auinger, 1880), NHMW2020/0166/0001, paralectotype, Brno- Kníničky (Kinitz).