Bellacolumbella embryonalis ( Boettger, 1902 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard M., 2021, The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group, Zootaxa 5025 (1), pp. 1-75 : 57

publication ID	https://doi.org/ 10.11646/zootaxa.5025.1.1
publication LSID	lsid:zoobank.org:pub:35E6C8FA-4078-4C53-9B74-F9618D5E6E60
persistent identifier	https://treatment.plazi.org/id/256F7D50-E253-FFD8-ACD6-301BFBD3B084
treatment provided by	Plazi
scientific name	Bellacolumbella embryonalis ( Boettger, 1902 )
status	comb. nov.

Treatment

Bellacolumbella embryonalis ( Boettger, 1902) View in CoL nov. comb.

Figs 4V, 17A View FIGURE 17 1 –A View FIGURE 1 2 View FIGURE 2 , B 1 –B View FIGURE 1 2 View FIGURE 2

* Columbella (Nitidella) embryonalis View in CoL n. sp. — Boettger 1902: 16.

Columbella (Anachis) embryonalis Bttgr. View in CoL — Boettger 1906: 19.

Pyrene (Anachis) embryonalis (Boettger) — Zilch 1934: 254, pl. 16, fig. 92.

Pyrene (Anachis) sp.— Csepreghy-Meznerics 1956: 403, pl. 7, figs 42–43.

Type materal. Lectotype (designated by Zilch 1934): SMF 360676 (= XII.2226 a), Senckenberg Museum, Frankfurt/ Main, Germany, SL: 5.1 mm, MD: 2.1 mm, Coşteiu de Sus ( Romania), illustrated in Zilch (1934, pl. 16, fig. 92), Figs 4V, 17A View FIGURE 17 1 –A View FIGURE 1 2 View FIGURE 2 . Paralectotypes: SMF 360676 (= XII.2223 a–b), Senckenberg Museum, Frankfurt/Main, Germany, SL: 4.2 mm, MD: 1.7 mm, Coşteiu de Sus ( Romania), Figs 17B View FIGURE 17 1 –B View FIGURE 1 2 View FIGURE 2 .

Additional material. NHMW 2020 View Materials /0173/0002 (ex B. Landau coll.), 2 spec., Coşteiu de Sus ( Romania) .

Description. Shell small, moderately broad fusiform, thin, high spired with deeply incised suture. Large, low conical multispiral protoconch of 3.5 convex whorls, terminating in moderately deep sinusigera. Protoconch microsculpture of delicate axial ribs being most prominent below mid-whorl, adapically fading out. Teleoconch of four whorls. Spire whorls weakly convex to flat-sided with periphery close to abapical suture. Sculpture of closeset, prominent orthocline to weakly opisthocline slightly curved axial ribs, adapically terminating in pointed tips. Spiral sculpture of weak spiral cords on abapical portion of base and siphonal fasciole. Last whorl convex, strongly constricted at base. Aperture moderately wide. Outer lip slightly thickened with five prominent denticles within. Anal canal weakly incised, V-shaped. Siphonal canal moderately long, narrow, deeply notched at tip. Columellar callus sharply delimited, forming narrow callus rim, with three columellar denticles. Small parietal callus pad.

Shell measurements and ratios. SL: 4.2–5.1 mm, MD: 1.7–2.1 mm, AA: 26–29°, SL/MD: 2.5–2.6, AL/AW: 3.1–3.2, LWH/AH: 1.3.

Discussion. This species differs from Bellacolumbella karreri and B. katharinae in the conical spire and the higher number of axial ribs, which develop only delicate nodes at their tips. In addition, the labial denticles are more prominent. Despite having more crowded axial sculpture than its congeners, the peculiar protoconch places it in Bellacolumbella .

Palaeoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés ( Hungary) ( Csepreghy-Meznerics 1956); Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906).