Orictites omnipunctatus, Balkenohl, 2017

Balkenohl, Michael, 2017, Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini), Contributions to Natural History 35, pp. 1-68 : 40-43

Orictites omnipunctatus sp. nov.

Type material: Holotype: ♂, labels: white, black printed: “ PHILIPPINES, 150 m Palawan, PORT BARTON 14.–18.Dec 1990, Bolm lgt.” ( NHMB).

Paratypes: 4 specimens, same data as holotype ( NHMB / CBB) ; 1 ♂, 1 ♀, 3 specimens. “ SULAWESI: Kotamobagu Metalibaru,  Torosik , Gn. Tongara , 10.XII.1999 700m, and 6.XII.1999, 1100–1450m, leg. A.Riedel ” ( SMNS / CBB) ; 3 ♀, “ SULAWESI, TENGAH : Nr.Morowali , Ranu River Area . 27.i.–20.iv.1980 / M.J.D.Brendell, B.M.1980–280 / Under bark of fallen tree ” ( BMNH / CBB) .

Description ( Figs 7 , 25 , 36 ; pp. 9, 24, 36)

Measurements (n = 10): Body length 3.45–3.95 mm ( = 3.7 mm); width 0.9- 1.03 ( = 0.95 mm); ration length/width of pronotum 1.03–1.11 ( = 1.07); ratio length/width of elytra 2.0–2.09 ( = 2.05).

Colour: Shiny. Red-brown. Elytra red-brown, pronotum darker. Wings of clypeus and supraantennal plates transparent red-brown, antennae, mouthparts, and legs yellowish brown.

Head: A fifth narrower than pronotum. Clypeus limited laterally by small rounded tooth, anterior margin excised, convex between the two clypeal setae, smooth but covered with medium sized punctures, wings obtuse-angular, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches, clypeus and wings reflexed margined, supraantennal plates distinctly vaulted up to mid-eye level, smooth, reflexed margin rounded, elongated as supraorbital carina, frons not separated from clypeus, thought indistinct flat depression visible laterally, supraorbital plates elongated posteriorly as obtuse diverging carina up to neck constriction, separated from supraorbital carinae by broad furrow and from frons by sharp diverging furrow. Frons distinctly convex without median impression, covered irregularly with punctures of large and medium size. Supraorbital setae located in broad supraorbital furrows, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae distinct, regularly rounded, as high as and shorter than eyes, enclosing eyes posteriorly by one sixth, forming obtuse angle at neck. Grooves for reception of antennae short, length about 0.4 of eye diameter. Neck constriction nearly invisible at middle, laterally with densely scattered medium sized punctures. Labrum slightly advanced at middle, 7-setose, ciliate laterally, with fine isodiametric reticulation (120x). Mandibles half as long as head, flattened, stout, broadened at base, gently curved at apex, carinae of scrobe complete, both mandibles rounded towards base, right one ventrally with small tooth at middle. Both mandibles somewhat hollowed out dorsally, both of them with small but distinct tooth near base (visible in completely opened position). Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2 nd segment, 2 nd segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with few punctures and reticulation, Submentum and mentum distinctly separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae short, just reaching over middle of pronotum, scapus knee-like angled, with one seta dorso-apically, scapus and pedicellus nearly smooth, segments 5–10 transverse (L/ W 0.8), antennomeres slightly flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.

Pronotum: Square, distinctly longer than wide, sides straight, gently rounded before posterior angle. Anterior margin straight. Reflexed lateral margin smooth, from posterior angle to base running as straight line. Lateral channel broadened between the two lateral setigerous punctures, smooth, with very few fine punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by twice diameter of pore. Anterior angles not projecting, nearly right-angled in dorsal view, formed by reflexed lateral margin, posterior angle developed as distinct tooth. Anterior transverse line formed by distinct partly connected punctures. Median line deep, broad, surpassing level of anterior transverse line without joining, smaller at base, joining base. Whole surface irregularly covered with medium and big sized punctures. No basal impression, no reticulation at base. Basal channel broad, deep. Flange raised keel-like (lateral view).

Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, completely visible from above in anterior two thirds, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with a few fine, indistinct scars posterior to humerus, channel slightly thickened at humerus, margin bending with channel over rounded humerus up to 5 th stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, broad, punctate, striae 1 and 7 joining at apex, 2 ending free, 3 and 4, 5 and 6 joined apically; striae 6 and 7 shortened at humerus. Striole short but distinct, interval between striole and sutura costiform, raised. All intervals conspicuously convex, thought costate, 2 nd to4 th slightly broader at base, 3 rd and4 th with small tubercle at base, 6 th costate, 7 th and 8 th completely carinae in total. All intervals shiny on disc. Interval 3 with four to seven setigerous punctures (see Tab. 1 ) adjoining stria 3.

Hind wings: Fully developed.

Ventral surface: Proepipleuron small, submarginal furrow distinct, engraved. Epipleuron broadened and with big punctures in basal quarter. Proepisternum somewhat swollen at level of posterior setigerous puncture, proepisternum and episternum covered densely and irregularly with big punctures, prosternite smooth at middle, with confluent double keel at middle, mesosternum smooth at middle. Sternites with big and medium-sized punctures, 3 rd to 5 th with paralateral ambulatory setae at each side, ventral strigae distinct; sternite 6 with big punctures at base, becoming finer at middle, minute towards apex, with two widely separated apical setae at each side, the inner one arising from tooth of margin.

Legs: Anterior tibia with strong digitation, with two big lateral preapical denticles, third one small, apical spine distinctly curved ventral and lateral, no dorsal furrow, basal tarsomere as long as the following three together. Mesotibia with apical spine, furnished with strong seta. All tarsomeres relatively broad.

Male genitalia ( Fig. 25 ): Median lobe slender, more distinctly arcuate at middle, oroficium large, endophallus with some small teeth, apex spatulate. Both parameres slender, dorsal one with seta at apex.

Female genitalia ( Fig. 36 ): Coxostylus broad, slightly curved at apex, with one apical seta. Ensiform seta at middle short, broad. With five longer nematiform setae in middle and basal part.

Variation: For variation of the setigerous punctures on the 3 rd interval of the elytra see Tab. 1 .

Diagnosis: A small pale-brown species with the knob on the clypeus developed as elevation and indistinct flat transverse depression laterally, no humeral tooth, and with tubercles at the base of interval 3 and 4 of the elytra. Distinguished from all other species by the series of setigerous punctures on interval 3 of the elytra, the numerous and dense punctures on many parts of the body, and the coxostylus with one apical seta.

Distribution: Known from Sulawesi and Palawan. The altitude of one collection spot is 150 m, of another one 700 m. One set of specimens was collected under bark of a fallen tree.

Etymology: The name refers to the numerous punctures on many parts of the body.

Figures

Figs 5–8: Orictites spp., habitus. 5: O. plurisetosus sp. nov., holotype, male; 6: O. anteriortenuis sp. nov., holotype, male; 7: O. omnipunctatus sp. nov., holotype, male; 8: O. ampliosetosus sp. nov., holotype, male.

Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm.

Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm.