Quedius (Raphirus) umbrinus Erichson, 1839

Salnitska, Maria & Solodovnikov, Alexey, 2018, Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae), Deutsche Entomologische Zeitschrift 65 (2), pp. 117-159 : 134-135

Quedius (Raphirus) umbrinus Erichson, 1839 Fig. 4B

Quedius umbrinus : Herman 2001, 3287 (summary of literature); Kascheev 1989, 36 (records); Assing and Schülke 2012, 475, 477 (diagnosis, distribution and bionomics, aedeagus illustration).

Material examined.

Kazakhstan: 1 ♂, Almaty Area, Dzhungarskiy Alatau Mts, 3 km SSE Lepsinsk, Bulinka River canyon, 1100-1800 m a.s.l., 45°30'N, 80°38'E, Betula sp., Malus, Populus etc. forest, 16-17.VI.2001, S.I. Golovatch leg. (cRyv); 1 ♂, Almaty Area, Talgar District., Ak-Bulak, 43.1613N, 77.2214E, 10-15.V.2014, O. Nakladal leg. (cKoc); 1 ♂, Lle-Alatau NP Talgar env., Ak-Bulak Resort, horse and cow dung, 1690 m a.s.l., 43.27039N, 77.37137E, 12-15.V.2014, M. Kocián leg. (cKoc); 1 ♂, 1 ♀, Lle-Alatau NP Talgar env., SW slope, leaf litter sifting, 1845 m a.s.l., 43.25851N, 77.38501E, 09.V.2014, M. Kocián leg. (cKoc).

Comments on taxonomy, distribution and bionomics.

Among all Middle Asian Raphirus , Q. umbrinus is most similar to Q. novus from which it can be distinguished by the structure of aedeagus: median lobe with distinct ventral tooth near its apex and apical portion slightly curved dorso-ventrally (in lateral view); paramere (underside) with sensory peg setae arranged in wide lateral rows merging at parameral anterior margin.

As a common and widespread species in Europe, Q. umbrinus was noted and illustrated in numerous papers. The latest summary can be found in Assing & Schülke (2012). Based on Kascheev (1989) and material examined here, Q. umbrinus occurs in the mountains of southern Kazakhstan where it can be found in leaf litter and dung at elevations up to 1845 m.