Pheidole ferruginea, Salata & Fisher, 2020

Salata, Sebastian & Fisher, Brian L., 2020, Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups, ZooKeys 905, pp. 1-235 : 1

Pheidole ferruginea sp. nov. Figs 35A-F , 84P , 86P

Type material.

Holotype. Madagascar. •1 major worker; Antsiranana; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 16 Nov 2003; B.L. Fisher leg.; BLF08998, CASENT0494997, middle specimen on the pin (CASC). Paratypes. Madagascar. •9w., 5s.; same data as for holotype; CASENT0494993-CASENT0494996, CASENT0872087, CASENT0872230-CASENT0872238 (CASC).

Other material.

Madagascar. - Antsiranana: •15w., 6s., 1m., 1q.; Mandraka; -18.91813, 47.91717; alt. 1312 m; 20 Mar 2014; B.L. Fisher et al. leg.; CASENT0375496, CASENT0377119, CASENT0377122, CASENT0377125, CASENT0377128, CASENT0377134, CASENT0377137, CASENT0377144, CASENT0377148, CASENT0377153, CASENT0377156, CASENT0377159, CASENT0378177, CASENT0378178, CASENT0378179, CASENT0378299, CASENT0378301, CASENT0378305, CASENT0378566, CASENT0378569 (CASC). •16w., 5s.; Reg. Analamanga, St. Forestière Mandraka; -18.9183, 47.91687; alt. 1285 m; 25 Mar 2015; B.L. Fisher et al. leg.; CASENT0390402, CASENT0390414, CASENT0390440, CASENT0390443, CASENT0390444, CASENT0390507, CASENT0390537, CASENT0703931, CASENT0720608, CASENT0720646, CASENT0720649, CASENT0720831, CASENT0720836, CASENT0720837, CASENT0728120, CASENT0728121 (CASC). •6w., 3s.; 6.5 km SSW Befingotra, Rés. Anjanaharibe-Sud; -14.75, 49.5; alt. 875 m; 19 Oct 1994; B.L. Fisher leg.; CASENT0198459-CASENT0198461 (CASC). •2s.; 9.2 km WSW Befingotra, Rés. Anjanaharibe-Sud; -14.75, 49.46667; alt. 1200 m; 9 Nov 1994; B.L. Fisher leg.; CASENT0198916 (CASC). •3w., 1s.; Binara Forest; -13.26392, 49.59919; alt. 1065 m; 18 Oct 2013; B.L. Fisher et al. leg.; CASENT0369870, CASENT0369873, CASENT0369876, CASENT0369878 (CASC). •3w., 1s.; Forêt Ambanitaza, 26.1 km 347° Antalaha; -14.67933, 50.18367; alt. 240 m; 26 Nov 2004; B.L. Fisher leg.; CASENT0055590, CASENT0055598 (CASC). •3w., 1s.; Forêt de Binara, 9.1 km 233°SW Daraina; -13.26333, 49.60333; alt. 800 m; 3 Dec 2003; B.L. Fisher leg.; CASENT0043315, CASENT0043364, CASENT0043367, CASENT0043447 (CASC). •12w., 3s., 1m.; Makirovana Forest; -14.17066, 49.95409; alt. 415 m; 28 Apr 2011; B.L. Fisher et al. leg.; CASENT0231230, CASENT0231233, CASENT0231258, CASENT0231270, CASENT0231278, CASENT0236129, CASENT0236143, CASENT0243310, CASENT0243314, CASENT0243315, CASENT0243348 (CASC). •2w.; Masoala National Park; -15.32331, 50.30751; alt. 60 m; 10 Mar 2014; B.L. Fisher et al. leg.; CASENT0376974, CASENT0376995 (CASC). •1s.; Masoala, Cap Est, Forêt d’Andranoanala; -15.26158, 50.4758; alt. 15 m; 15 Mar 2014; B.L. Fisher et al. leg.; CASENT0377900 (CASC). •39w., 7s., 1q.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 15 Nov 2003; B.L. Fisher et al. leg.; CASENT0045227, CASENT0045231, CASENT0045238, CASENT0045240, CASENT0045243, CASENT0045248, CASENT0045250, CASENT0045252, CASENT0045254, CASENT0045260, CASENT0045264, CASENT0045267, CASENT0045272, CASENT0045334, CASENT0045351, CASENT0045363, CASENT0045366, CASENT0045744, CASENT0045904, CASENT0045987, CASENT0068463, CASENT0068464, CASENT0494859, CASENT0494860, CASENT0494917, CASENT0846566-CASENT0846569, CASENT0846571CASENT0846572, CASENT0846574, CASENT0846577-CASENT0846587 (CASC). •20w., 4s.; Parc National de Marojejy, Manantenina River, 28.0 km 38°NE Andapa, 8.2 km 333°NNW Manantenina; -14.43667, 49.775; alt. 450 m; 12 Nov 2003; B.L. Fisher et al. leg.; CASENT0045803, CASENT0045805, CASENT0045811, CASENT0045832, CASENT0045845, CASENT0045867, CASENT0045876, CASENT0045886, CASENT0045889, CASENT0045899, CASENT0045902, CASENT0045908, CASENT0046015, CASENT0046021, CASENT0047669, CASENT0077149, CASENT0077150, CASENT0077152 (CASC). - Fianarantsoa: •6w., 4s., 1q.; 2 km W Andrambovato, along river Tatamaly; -21.51167, 47.41; alt. 1075 m; 3 Jun 2005; B.L. Fisher et al. leg.; CASENT0060943, CASENT0060944, CASENT0061397, CASENT0061581, CASENT0061621, CASENT0061677, CASENT0109000 (CASC). •1w.; 43 km S Ambalavao, Rés. Andringitra; -22.23333, 47; alt. 825 m; 5 Oct 1993; B.L. Fisher leg.; CASENT0198537 (CASC). •1w.; 45 km S. Ambalavao; -22.21667, 47.01667; alt. 785 m; 25 Sep 1993; B.L. Fisher leg.; CASENT0198538 (CASC). •1w.; 7.6 km 122° Kianjavato, Forêt Classée Vatovavy; -21.4, 47.94; alt. 175 m; 6 Jun 2005; B.L. Fisher et al. leg.; CASENT0061235 (CASC). •3w., 1s.; 9.0 km NE Ivohibe; -22.42667, 46.93833; alt. 900 m; 12 Nov 1997; B.L. Fisher et al. leg.; CASENT0198539, CASENT0198541, CASENT0198918 (CASC). •5w., 2s., 1q.; Ambinanindranomena Non-Protected Area, 39.45 km SE Ambalavao; -21.95386, 47.29427; alt. 1069 m; 1 Feb 2012; Andrianjaka & Ravelomanana leg.; CASENT0293842, CASENT0293876, CASENT0293878, CASENT0293916, CASENT0293917, CASENT0293935 (CASC). •1w., 1s.; Forêt d’Ambalagoavy Nord, Ikongo, Ambatombe; -21.857068, 47.37849; alt. 625 m; 1 Dec 2000; Harin’Hala & Irwin leg.; CASENT0009589, CASENT0009636 (CASC). •2w.; P.N. Ranomafana, Sahavondrona-Ampitamarivo; -21.2575, 47.36015; alt. 1100 m; 18 Mar 2003; Clark leg.; CASENT0052899, CASENT0052904 (CASC). •1w.; P.N. Ranomafana, Tolongoina-Ampasimpotsy 3; -21.47412, 47.55742; alt. 520 m; 11 Apr 2003; Clark leg.; CASENT0052880 (CASC). •1w., 1s., 1q.; Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango; -23.83517, 46.96367; alt. 940 m; 13 Nov 2006; B.L. Fisher et al. leg.; CASENT0128696, CASENT0128700, CASENT0128704 (CASC). •1w., 1s.; Parc National Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango; -23.84083, 46.9575; alt. 1250 m; 17 Nov 2006; B.L. Fisher et al. leg.; CASENT0118391 (CASC). •1w., 1s.; Parc National de Ranomafana, Vatoharanana River, 4.1 km 231°SW Ranomafana; -21.29, 47.43333; alt. 1100 m; 27 Mar 2003; B.L. Fisher et al. leg.; CASENT0039949, CASENT0040207 (CASC). •2w., 1s.; R.S. Ivohibe 8.0 km E Ivohibe; -22.48333, 46.96833; alt. 1200 m; 15 Oct 1997; B.L. Fisher et al. leg.; CASENT0198540, CASENT0198542, CASENT0198919 (CASC). •1w.; Vohiparara Kidonavo 1; -21.22632, 47.37007; alt. 1100 m; 13 Mar 2003; Clark leg.; CASENT0052978 (CASC). - Toamasina: •3w.; Ambanizana, Parc National Masoala; -15.57167, 50.00611; alt. 900-950 m; 26 Feb 2003; Andriamalala et al. leg.; CASENT0008765, CASENT0073478, CASENT0073478 (CASC). •3w., 1s.; Ambatovy, 12.4 km NE Moramanga; -18.83937, 48.30842; alt. 1080 m; 4 Mar 2007; B.L. Fisher et al. leg.; CASENT0123826, CASENT0123867, CASENT0123870, CASENT0123997 (CASC). •1w.; Analamay; -18.80623, 48.33707; alt. 1068 m; 21 Mar 2004; B.L. Fisher et al. leg.; CASENT0046556 (CASC). •2w.; Ankerana; -18.4017, 48.80605; alt. 1035 m; 24 Jan 2012; B.L. Fisher et al. leg.; CASENT0274446, CASENT0274448 (CASC). •6w.; Ankerana; -18.4061, 48.82029; alt. 725 m; 16 Jan 2012; B.L. Fisher et al. leg.; CASENT0274767, CASENT0275345, CASENT0275347, CASENT0275348, CASENT0275462 (CASC). •2w., 2q.; Ankerana; -18.40829, 48.82107; alt. 750 m; 21 Jan 2012; B.L. Fisher et al. leg.; CASENT0274977, CASENT0274978 (CASC). •1w.; Ankerana; -18.4104, 48.8189; alt. 855 m; 22 Jan 2012; B.L. Fisher et al. leg.; CASENT0274417 (CASC). •1w.; Corridor Forestier Analamay-Mantadia, Tsaravoniana; -18.75641, 48.42195; alt. 1036 m; 2 Dec 2012; B.L. Fisher et al. leg.; CASENT0301883 (CASC). •3w., 2s.; F.C. Sandranantitra; -18.04833, 49.09167; alt. 450 m; 18 Jan 1999; Ratsirarson leg.; CASENT0198543, CASENT0198920 (CASC). •2w., 1s.; Ile Sainte Marie, Forêt Kalalao, 9.9 km 34° Ambodifotatra; -16.9225, 49.88733; alt. 100 m; 24 Nov 2005; B.L. Fisher et al. leg.; CASENT0069837, CASENT0069839, CASENT0069844 (CASC). •1w., 1s.; Mahavelona (Foulpointe); -17.66667, 49.5; 1 Nov 1985; A. Pauly leg.; CASENT0095644, CASENT0095650 (CASC). •11w., 11s.; Montagne d’Akirindro 7.6 km 341°NNW Ambinanitelo; -15.28833, 49.54833; alt. 600 m; 17 Mar 2003; B.L. Fisher et al. leg.; CASENT0038892, CASENT0038907, CASENT0039031, CASENT0039045, CASENT0039049, CASENT0039079, CASENT0039080, CASENT0039090, CASENT0039094, CASENT0039109, CASENT0039113, CASENT0039121, CASENT0039194, CASENT0039205, CASENT0039213, CASENT0039217, CASENT0039224, CASENT0039226, CASENT0039233, CASENT0496453 (CASC). •2w.; Montagne d’Anjanaharibe, 18.0 km 21°NNE Ambinanitelo; -15.18833, 49.615; alt. 470 m; 8 Mar 2003; B.L. Fisher et al. leg.; CASENT0037612, CASENT0037639 (CASC). •1w., 3s.; Parc National de Zahamena, Onibe River; -17.75908, 48.85468; alt. 780 m; 21 Feb 2009; B.L. Fisher et al. leg.; CASENT0150453, CASENT0151642, CASENT0153420, CASENT0153422 (CASC). •4w.; Parc National de Zahamena, Tetezambatana forest, near junction of Nosivola and Manakambahiny Rivers; -17.74298, 48.72936; alt. 860 m; 18 Feb 2009; B.L. Fisher et al. leg.; CASENT0151078, CASENT0153357, CASENT0153371, CASENT0153374 (CASC). •4w., 4s.; Parc National Mananara-Nord, 7.1 km 261° Antanambe; -16.455, 49.7875; alt. 225 m; 16 Nov 2005; B.L. Fisher et al. leg.; CASENT0067223, CASENT0068923, CASENT0069441, CASENT0069445CASENT0069451, CASENT0069605, CASENT0071324 (CASC). •1w., 2s.; Reserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina; -17.924, 49.19967; alt. 390 m; 28 Nov 2005; B.L. Fisher et al. leg.; CASENT0071885, CASENT0071912, CASENT0069185 (CASC). •1w.; Réserve Spéciale Ambatovaky, Sandrangato River; -16.81739, 49.29402; alt. 360 m; 25 Feb 2010; B.L. Fisher et al. leg.; CASENT0163669 (CASC). •8w., 1q.; Réserve Spéciale Ambatovaky, Sandrangato River; -16.77274, 49.26551; alt. 450 m; 20 Feb 2010; B.L. Fisher et al. leg.; CASENT0164042, CASENT0164417, CASENT0164422, CASENT0164440, CASENT0164468, CASENT0164483, CASENT0164485, CASENT0164492, CASENT0164498 (CASC). •5w., 1q.; Réserve Spéciale Ambatovaky, Sandrangato River; -16.7633, 49.26692; alt. 520 m; 22 Feb 2010; B.L. Fisher et al. leg.; CASENT0163718, CASENT0163722, CASENT0163723, CASENT0163833, CASENT0163873, CASENT0163877 (CASC). - Toliara: •1w., 3s.; 10 km NW Enakara, Rés Andohahela; -24.56667, 46.81667; alt. 420 m; 15 Nov 1992; B.L. Fisher leg.; CASENT0198037, CASENT0196935 (CASC). •6w., 1s., 1q.; 11 km NW Enakara, Rés. Andohahela; -24.56667, 46.83333; alt. 800 m; 17 Nov 1992; B.L. Fisher leg.; CASENT0196936, CASENT0198050, CASENT0217984, CASENT0217992, CASENT0217993 (CASC). •5w.; 2.7 km WNW 302° Ste. Luce; -24.77167, 47.17167; alt. 20 m; 9 Dec 1998; B.L. Fisher et al. leg.; CASENT0196937-CASENT0196939, CASENT0198049, CASENT0198051 (CASC). •8w., 2s.; Anosy Region, Anosyenne Mts, 31.2 km NW Manantenina; -24.13894, 47.06804; alt. 1125 m; 26 Feb 2015; B.L. Fisher et al. leg.; CASENT0704213, CASENT0704214, CASENT0704872, CASENT0704873, CASENT0704876, CASENT0704877, CASENT0721011 (CASC). •2w., 1s.; Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro; -24.7585, 46.85367; alt. 275 m; 24 Nov 2006; B.L. Fisher et al. leg.; CASENT0122564, CASENT0129922, CASENT0129927 (CASC). •1w., 1s.; Parc National Andohahela, Manangotry, 33.8 km NW Tolagnaro; -24.75117, 46.85783; alt. 575 m; 24 Nov 2006; B.L. Fisher et al. leg.; CASENT0121878 (CASC). •21w., 6s.; Parc National d’Andohahela, Col du Sedro, 3.8 km 113°ESE Mahamavo, 37.6 km 341°NNW Tolagnaro; -24.76389, 46.75167; alt. 900 m; 21 Jan 2002; B.L. Fisher et al. leg.; CASENT0078392, CASENT0078409, CASENT0078410, CASENT0078412, CASENT0479172, CASENT0479174, CASENT0479187, CASENT0479188, CASENT0479239, CASENT0479260, CASENT0479261, CASENT0479262, CASENT0479299, CASENT0483039, CASENT0483924, CASENT0483951, CASENT0483967, CASENT0483968, CASENT0483980, CASENT0484029, CASENT0484043, CASENT0484060, CASENT0484062, CASENT0484063, CASENT0484111, CASENT0484183 (CASC). •6w., 2s., 1q.; Réserve Spéciale Kalambatritra, Ampanihy; -23.4635, 46.4631; alt. 1270 m; 9 Feb 2009; B.L. Fisher et al. leg.; CASENT0150522, CASENT0150523, CASENT0153214, CASENT0153217, CASENT0217989, CASENT0846595, CASENT0846596, CASENT0846597 (CASC).

Diagnosis.

Head in full-face view cordate, widened posteriorly; sides of the head with sparse, long, suberect pilosity; antennal scrobes strongly developed, well delimited and forming distinct dorsal concavity beneath frontal carina; scrobe surface shiny, with sparse, thick, longitudinal and sometimes additional irregular rugae; delimited ventrally and posteriorly by carinulae; propodeal spines long (PSL: 0.23-0.28 (0.26)); first gastral tergite shagreened, at least on its basal part. Minor workers. Head foveolate, sometimes with additional longitudinal rugae on frons; propodeal spines very long (PSL: 0.15-0.18 (0.16)); promesonotum box-like; mesosoma foveolate, sometimes with additional rugae on mesosomal dorsum, and katepisternum with smooth notch.

Description.

Major workers. Measurements (N = 10): HL: 1.18-1.34 (1.25); HW: 1.15-1.31 (1.23); SL: 0.61-0.69 (0.64); EL: 0.13-0.17 (0.15); WL: 0.93-1.05 (0.99); PSL: 0.23-0.28 (0.26); MTL: 0.58-0.66 (0.61); PNW: 0.5-0.6 (0.53); PTW: 0.13-0.17 (0.16); PPW: 0.45-0.58 (0.51); CI: 97.4-101.3 (99.0); SI: 48.7-55.8 (52.1); PSLI: 18.9-21.9 (20.8); PPI: 27.5-33.7 (30.7); PNI: 40.9-45.0 (43.0); MTI: 47.6-52.2 (49.7). Head. In full-face view cordate, widened posteriorly (Fig. 35B ). In lateral view oval; ventral and dorsal faces strongly convex; inner hypostomal teeth visible. Sides of the head with sparse, long, suberect pilosity; whole head with moderately dense, long, suberect to erect pilosity. Antennal scrobes strongly developed, well delimited and forming distinct dorsal concavity beneath frontal carina, scrobe surface shiny, with sparse, thick, longitudinal, and sometimes additional irregular rugae, delimited ventrally and posteriorly by carinulae. Occipital lobes shiny, with sparse and thick rugoreticulation; genae shiny, with moderately dense to dense, fine rugoreticulation; frons and malar area with sparse, thick, and longitudinal rugae, interspaces smooth or with fine rugulae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, narrow, and shallow; median longitudinal carina present but sometimes indistinct; lateral longitudinal carinae absent. Scape, when laid back, reaching slightly beyond the midlength of head; pilosity decumbent to erect (Fig. 35B, D ). Inner hypostomal teeth distinct, closely spaced, triangular, with rounded apex and moderately narrow base; outer hypostomal teeth distinct, slightly lower than inner hypostomal teeth, lobe-like, with base wide and tops directed slightly inward (Fig. 84P ). Mesosoma. In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum with tubercle-like projection, dropping steeply to propodeum; promesonotal groove absent; metanotal groove absent; propodeal spines long, massive basally, with acute apex; humeral area laterally weakly produced (Fig. 35D ). Surface shiny, with thick and dense rugoreticulation, sometimes sculpture weakening on dorsum. Pilosity moderately sparse, long, and erect (Fig. 35D, F ). Petiole. Shagreened; peduncle relatively long, without horizontal lobes on its basal part; node low, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig. 35D, F ). Postpetiole. Shagreened; in dorsal view sides with acute, triangular, moderately long projections; pilosity long, moderately sparse, and erect (Fig. 35D, F ). Petiole. First gastral tergite shagreened, at least on its basal part; pilosity moderately sparse, long, and erect (Fig. 35D, F ). Colour. Head and mesosoma reddish brown to brown; dorsum of mesosoma dark brown; gaster brown; legs yellow to brown (Fig. 35D, F ).

Minor workers. Measurements (N = 10): HL: 0.52-0.57 (0.55); HW: 0.44-0.48 (0.46); SL: 0.57-0.63 (0.6); EL: 0.11-0.13 (0.12); WL: 0.67-0.72 (0.7); PSL: 0.15-0.18 (0.16); MTL: 0.44-0.49 (0.46); PNW: 0.31-0.35 (0.33); PTW: 0.07-0.09 (0.08); PPW: 0.13-0.14 (0.13); CI: 80.9-86.2 (83.9); SI: 124.6-134.8 (130.0); PSLI: 27.5-31.4 (29.3); PPI: 57.4-67.9 (63.1); PNI: 69.6-74.8 (71.7); MTI: 96.9-102.3 (100.9). Head. Occipital margin straight or indistinctly concave; occipital carina absent (Fig. 35A ). Pilosity sparse, long, suberect to erect. Whole head foveolate, sometimes with additional longitudinal rugae on frons. Clypeus foveolate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing posterior head margin by two-fifths of its length; pilosity suberect to erect (Fig. 35A, C ). Mesosoma. In lateral view, promesonotum box-like; promesonotal groove absent; metanotal groove absent; propodeal spines very long, massive basally, with acute apex (Fig. 35C ). Whole surface foveolate, sometimes with additional rugae on mesosomal dorsum; sometimes katepisternum with smooth notch. Pilosity sparse, long, and erect (Fig. 35C, E ). Petiole. Peduncle relatively long and thin; node low, globular; with few long, erect setae (Fig. 35C, E ). Postpetiole. Short, low, and convex; with few long, erect setae at the anterior edge (Fig. 35C, E ). Petiole. Pilosity sparse and erect (Fig. 35C, E ). Colour. Unicolourous, yellow to brown (Fig. 35C, E ).

Etymology.

Latin for rusty, in reference to body colouration of major workers.

Biology.

The species was collected between 20-1312 m in elevation, in rainforest, montane rainforest, and littoral rainforest. Nests were located in rotten logs and tree stumps, rotten sticks on ground, and the petioles of Melastomataceae .

Comments.

This species is most similar to P. rugocephala sp. nov. Major workers. Pheidole ferruginea sp. nov. can be distinguished from P. rugocephala sp. nov. by the surface of antennal scrobes never foveolate; presence of longitudinal rugae on frons; inner hypostomal teeth newer pointed inward; never smooth surface of promesonotum; and lack of horizontal lobes on the basal part of petiolar peduncle. Minor workers. Pheidole ferruginea sp. nov. can be distinguished from P. rugocephala sp. nov. by long propodeal spines and foveolate clypeus, and lack of promesonotal and metanotal grooves.

Figures

Figure 35. Pheidole ferruginea sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratypeminor worker (CASENT 0494994) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0494997).

Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z).

Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U).