Anexodus aquilus Pascoe, 1886

Gabris, Radim, Trnka, Filip, Wahab, Rodzay Abdul & Kundrata, Robin, 2017, Taxonomic revision of the endemic Bornean genera Anexodus Pascoe and Pantilema Aurivillius (Coleoptera, Cerambycidae, Lamiinae), ZooKeys 669, pp. 29-51 : 30

Anexodus aquilus Pascoe, 1886 Figs 1-3, 9-11, 17, 21, 22, 28, 32

Anexodus aquilus Pascoe, 1886: 242.

Anexodus kuntzeni Kriesche, 1924: 291; synonymized by Breuning (1950): 258.

Type material examined

( A. aquilus ). Holotype, male, [Malaysia] "Type [circular label with red margin, printed] // N / Borneo [blue oval label, handwritten] // Anexodus / aquilus / type Pasc. [handwritten] // Anexodus / aquilus / N. Borneo Pa [handwritten] // Pascoe / Coll. / 93-60" (BMNH).

Type material examined

( A. kuntzeni ). Holotype, female, [Malaysia] "N:Borneo / Kina-Balu-Geb. / Waterstradt S. [printed] // Anexodus / küntzeni Kriesche / Typ! [handwritten] // Anexodus / aquilus Pasc. / Breuning dét. // HOLOTYPUS / Anexodus / kuntzeni Kriesche 1924 / labelled by MNHUB 2014 [red label, printed]" (MNHUB); male, "N:Borneo / Kina-Balu-Geb. / Waterstradt S. [printed] // Anexodus / küntzeni Kriesche / Paratyp! [handwritten] // PARATYPUS / Anexodus / kuntzeni Kriesche 1924 / labelled by MNHUB 2014 [red label, printed]" (MNHUB); female, "N. Borneo / Kinabalů [handwritten] // Anexodus / küntzeni [handwritten] // Anexodus / aquilus Pasc. / Breuning dét. // PARATYPUS / Anexodus / kuntzeni Kriesche 1924 / labelled by MNHUB 2014" (MNHUB).

Other material examined.

Female, "Malaysia, Sabah, / Crocker Range 20- / IV-2007 Cope / collection" (PCJC); female, "Malaysia, Sabah / Crocker Range / III-22-2009 / local coll" (PCDH); female, "Malaysia, Sabah / Sandakan / II-12-2007 / local coll // Anexodus / aquilus Pasc. / det. D. Heffern ‘07” (PCDH); female, "Malaysia, Sabah / Ranau 700 m / IV-23-2015 / local coll" (PCDH); female, "Malaysia, Sabah / Tenom / III-2-2008 / local coll" (PCDH); male, "Malaysia, Sabah, Mt. / Trus Madi 26-IV-2010 / Cope Collection" (PCJC); male, "Malaysia, Sabah, Mt. / Trus Madi 26-V-2012 / Cope Collection" (PCJC); male, "Malaysia, Sabah / Mt. Trus-Madi / IV-15-2005 / local coll // Anexodus / aquilus / Pascoe / det. J. Sudre" (PCDH).

Differential diagnosis.

This species is similar to A. tufi sp. n. in general habitus, yellowish stripes on dorsal body surface and a labrum with the entire surface punctured but differs in having apex of scape with a distinct lateral hook-shaped projection (simple in A. tufi sp. n.; Figs 21, 22, 26, 27), relatively longer parameres (Fig. 28), and a widened second half of spermatheca (Fig. 32).

Description.

Holotype (male). BL 11.5 mm, BW 3.8 mm. Body brown with yellowish stripes extending from vertex through sides of pronotum to basal part of elytra; mouthparts lighter; antennae reddish brown. Body densely clothed with very short brown pubescence. Head about as wide as anterior margin of pronotum. Labrum transverse, with whole surface moderately sparsely, irregularly punctured (Fig. 17). Eyes moderately emarginate at antennal articulations (Figs 9-11). Antennae 0.9 times as long as body; scape gradually slightly widened towards apex, thickest at apical part, apex with a distinct lateral hook-shaped projection (Fig. 21-22); the relative ratio of antennomere lengths 3.3: 1.6: 1.0: 1.2: 1.1: 0.9: 0.8: 0.6: 0.6: 0.6: 1.1.

Prothorax as long as wide, laterally with one distinct tubercle; pronotal disc with a pair of indistinct tubercles near middle and one median at second half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, two times as wide as long. Elytra elongate, 1.6 times as long as wide at widest part, 1.7 times as long as pronotum, widest near middle; each elytron with three rows of indistinct tubercles, inner row forming a distinct ridge basally; sparsely covered with large deep punctures arranged in slightly irregular rows. Legs long, slender; relative lengths of metatarsomeres 1.0: 0.5: 1.0: 2.0.

Male genitalia with tegmen elongate, widest near middle, basally with very short strut; parameres elongate, 3.5 times longer than wide, apically with long setae (Fig. 28). Penis subparallel-sided, apically truncate; dorsal struts diverged from about 1/3 of penis length. Internal sac long, with paired small medial sclerites and distinct flagellar sclerites.

Variability in males. BL 9.1-12.5 mm, BW 3.1-4.2 mm. Antennae are either reddish brown, brown or black. There is a gradual morphological variation in the pronotal and elytral tubercles, ranging from the less distinct tubercles in the holotype (Figs 1, 9) through the more distinct tubercles in most specimens to the strongly developed tubercles with inner elytral row forming a conspicuous ridge basally in the specimens from Trus Madi (Figs 3, 11).

Female. Most characters same as for males. BL 10.0-13.0 mm, BW 3.2-4.5 mm. Body more convex dorsally, with distinct tubercles on pronotum and elytra. Antennae shorter, 0.6-0.7 times as long as body. Elytra 1.7 times as long as wide, 2.0-2.1 times as long as pronotum. Spermatheca sclerotized, slender, elongate, curved, widened at second half, gradually tapered toward apex; sclerotized part of spermathecal duct simple (Fig. 32).

Distribution.

Malaysia: Borneo (Sabah; Fig. 43).