Belonuchus Nordmann 1837
publication ID |
https://doi.org/ 10.11646/zootaxa.4449.1.1 |
publication LSID |
lsid:zoobank.org:pub:0734DC4E-2C9B-4CD2-B8FE-8F94589200E6 |
DOI |
https://doi.org/10.5281/zenodo.5950889 |
persistent identifier |
https://treatment.plazi.org/id/215187C8-4C20-B355-FF6B-FC00F4ACD3FC |
treatment provided by |
Plazi |
scientific name |
Belonuchus Nordmann 1837 |
status |
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Belonuchus Nordmann 1837 View in CoL
Belonuchus Nordmann 1837: 129 View in CoL . Type species: Staphylinus haemorrhoidalis Fabricius, 1801 View in CoL , by monotypy.
= Musicoderus Sharp 1885: 455 View in CoL . Type species: Musicoderus cephalotes Sharp 1885 View in CoL , des. by Lucas 1920: 425. Notes: as subgenus of Belonuchus ( Bernhauer 1916c: 266) View in CoL ; as synonym of Belonuchus ( Blackwelder 1943: 421) View in CoL .
Notes: There are 165 valid species of Belonuchus View in CoL distributed in CASA. A recent study ( Chani-Posse et al. 2018) shows Belonuchus View in CoL as a genus of Neotropical origin and generic limits so far undefined. The current concept of the genus is that of Smetana (1995), Chani-Posse (2014a) and Chani-Posse et al. (2018) but most of its Neotropical endemic species were never revised after being described and they do not fit into this concept. Species that may belong to the so called ‘ Belonuchus haemorrhoidalis View in CoL clade’ ( Chani-Posse et al. 2018) are here denoted with an asterisk *.
* Belonuchus abnormalis ( Sharp 1885) View in CoL , comb. nov. ( Figs. 1–2 View FIGURES 1–12 )
Philonthus abnormalis Sharp 1885: 397 View in CoL . Type locality: Mexico: Oaxaca. Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [MCP seen]. Distribution: Mexico, Guatemala. Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).
Remarks. Sharp explicitly mentioned two specimens (male and female) as part of the type series. The two specimens listed below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 1–2 View FIGURES 1–12 ): card-mounted, “ ♂ (in ink) Philonthus abnormalis Type D.S. Oaxaca. Mexico. Höge [in Sharp’s hand] // Type (round white label with red margin) // Oaxaca, Mexico. Hoege. (white label) // B. C. A. Col. I. 2. Philonthus abnormalis, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotype ♀ ( BMNH): with same locality and collection data as lectotype, and additional printed white label “Sp. Figured”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) ; determined as “ Belonuchus ” by J. L. Navarrete-Heredia in 2002.
Belonuchus aequalis Sharp 1876
Belonuchus aequalis Sharp 1876a: 161 View in CoL . Type locality: Pará, Ega, St. Paulo [Amazonas] & Conceicão, Rio Mauhes.
Type depository: BMNH 6 ST: 4 Ega, 1 Pará, 1 St. Paulo [ MCP seen] . FMNH 1 About FMNH ST: 1 Ega. Distribution: Brazil . Notes: distribution in Brazil (Asenjo et al. 2013).
* Belonuchus agilis Erichson 1840 View in CoL
Belonuchus agilis Erichson 1840: 423 View in CoL . Type locality: Cuba. Type depository: ZMHB 2 ST: 1♂, 1♀ [MCP seen].
Distribution: USA, Dominican Rep., Cuba, Jamaica, Haiti, Colombia. Notes: distribution including USA and Dominican Rep. ( Frank 2004); distribution, also within Cuba ( Peck 2005).
= Belonuchus chevrolati Fauvel 1863: 432 View in CoL . Type locality: La Havane, Cuba. Type depository: IRSNB HT ♂ [MCP seen]. Note: as synonym of B. agilis ( Fauvel 1869: 489) View in CoL .
Belonuchus albovariegatus Bernhauer 1916 View in CoL
Belonuchus albovariegatus Bernhauer 1916c: 276 View in CoL . Type locality: West-Columbien: Umgebung Cali am Rio Cauca, St. Antonio, 2000 m. Type depository: FMNH LT ♂, 2 PLT (des. Chani-Posse & Solodovnikov 2017). Distribution: Colombia.
= Linoderus navarretei López-García & Méndez-Rojas 2014: 86 View in CoL . Type locality: Colombia, Quindío Dept., Filandia, Cruces, Parque Regional Natural Barbas-Bremen , 4°42’6.4” N 75°38’16.3” W, 2000 m GoogleMaps . Type depositories: UNCB HT ♂, 13 PT; CIUQ 3 PT. Note : as synonym of B. albovariegatus ( Chani-Posse & Solodovnikov 2017: 90) View in CoL .
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus alternans ( Sharp 1885) View in CoL
Philonthus alternans Sharp 1885: 418 View in CoL . Type locality: Mexico: Jalapa, Oaxaca. Type depository: BMNH 3 ST; FMNH 1ST. Distribution: Mexico. Notes: comb. nov. ex Philonthus ( Navarrete-Heredia et al. 2002: 334) View in CoL ; distribution within Mexico ( Márquez 2006).
* Belonuchus alternatus Sharp 1885 View in CoL
Belonuchus alternatus Sharp 1885: 430 View in CoL . Type locality: Guatemala, near the City, San Geronimo; Nicaragua, Chontales; Panama, Bugaba. Type depository: BMNH #? ST: several, from all type localities [MCP seen]. FMNH 4 ST: Panama, Bugaba. Distribution: Panama, Nicaragua, Guatemala. Note: misspelled as alternans View in CoL in error by Bernhauer & Schubert (1914: 369) (not Philonthus alternans Sharp 1885: 418 View in CoL ).
* Belonuchus altissimus Bernhauer 1916 View in CoL
Belonuchus altissimus Bernhauer 1916c: 275 View in CoL . Type locality: Monte Socorro, West-Cordilleren, West-Columbien [3800m on type label]. Type depository: FMNH HT. Distribution : Colombia .
Belonuchus altivagans Bernhauer 1916 View in CoL
Belonuchus altivagans Bernhauer 1916c: 272 View in CoL . Type locality: West-Columbien: Pacho, 2000m, Ost-Cordilleren, West-Columbien. Type depository: FMNH 2 About FMNH ST. Distribution : Colombia .
Comment. Not Belonuchus , to be placed in a further study.
Belonuchus aluticollis (Bernhauer 1921)
Philonthus aluticollis Bernhauer 1921c: 107 View in CoL . Type locality: Cueva Iturbe, 3700m, Jujuy, Argentina [LT without "Cueva"]. Type depository: FMNH LT ♀, MACN 2 PLT. Distribution: Argentina. Note: comb. nov. ex Philonthus View in CoL and lectotype designation ( Chani-Posse 2010: 8); redescription ( Chani-Posse 2011b: 215).
Belonuchus amplipennis Fauvel 1891 View in CoL
Belonuchus amplipennis Fauvel 1891: 117 View in CoL . Type locality: Colonie Tovar, San Esteban [ Venezuela]. Type depository: IRSNB 2 ST [MCP seen]. Distribution: Venezuela.
Belonuchus amplus Blackwelder 1943 View in CoL
Belonuchus amplus Blackwelder 1943: 425 View in CoL . Type locality: St. Lucia: 0.5 mile northwest of St. Remy on road to Soufrière (2.5 miles southeast of Soufrière) . Type depository: USNM HT ♀, 1 PT ♀. Distribution: Lesser Antilles ( St. Lucia) .
Belonuchus andinus Bernhauer 1916 View in CoL
Belonuchus andinus Bernhauer 1916c: 278 View in CoL (attributed to Fauvel in litt.). Type locality: Yuracarès, Bolivien. Type depository: FMNH 1 About FMNH ST; ZMUH 1 View Materials +ST? [destroyed?] ; IRSNB 2 View Materials +3? ST: only two of them in Bernhauer´s hand [ MCP seen]. Distribution : Bolivia.
Belonuchus angusticeps Scheerpeltz 1960 View in CoL
Belonuchus (Musicoderus) angusticeps Scheerpeltz 1960: 117 View in CoL . Type locality: Cochabamba ( Bolivien), in den Yungas de Palmar , 2000m . Type depository: NMW 2 View Materials ST; FMNH 2 About FMNH ST. Distribution : Bolivia .
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus angusticollis Bernhauer 1906 View in CoL
Belonuchus angusticollis Bernhauer 1906: 330 View in CoL . Type locality: Chanchamayo , Peru . Type depository: FMNH HT. Distribution : Peru . Note: Peru ( Asenjo 2004).
Belonuchus antiguae Blackwelder 1943 View in CoL
Belonuchus antiguae Blackwelder 1943: 434 View in CoL . Type locality: Antigua: near summit of Boggy Peak , parish of St. Mary, elevation 1300 feet . Type depository: USNM HT ♂, 5 PT; BMNH 1 PT; FMNH 1 About FMNH PT. Distribution: Antigua. Notes: distribution within Lesser Antilles (Northern Leeward Is.) ( Peck 2011a); generic affiliation unclear.
Belonuchus apicicornis Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) apicicornis Bernhauer 1916c: 270 View in CoL . Type locality: West-Columbien: Cali am Rio Cauca ( Rio Aquatal , 1800m; S. Antonio, 2000m; Rio Vitaco, 2200m). Type depository: FMNH 2 About FMNH ST;? 1 ST. Distribution: Colombia .
* Belonuchus apiciventris ( Sharp 1885) View in CoL
Philonthus apiciventris Sharp 1885: 419 View in CoL . Type locality: Mexico: Juquila. Type depository: BMNH HT ♂. Distribution: Mexico. Notes: in cave collections ( Navarrete-Heredia & Márquez-Luna 1993); cited as apaciventris, characters. habits, natural history notes, host mushrooms (Navarrete-Heredia 1995); comb. nov. ex Philonthus ( Jiménez-Sánchez et al. 2000) View in CoL ; distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).
Belonuchus argentinus Bernhauer 1916 View in CoL
Belonuchus argentinus Bernhauer 1916c: 278 View in CoL . Type locality: Misiones, Argentina. Type depositary: FMNH HT, 2 PT; MLPA 8 PT (as ST in error, Fernandez et al. 2007). Distribution: Argentina, Brazil, Bolivia. Notes: MLPA "ST", Fernandez et al. 2007, but original reference to "Das typische Stück …"; name cited by Bruch (1915: 503) as nomen nudum, attributed to Bernhauer in litteris; distribution in Brazil (Asenjo et al. 2013); generic affiliation unclear.
* Belonuchus armatus Sharp 1876 View in CoL
Belonuchus armatus Sharp 1876a: 163 View in CoL . Type locality: Abacaxis. Type depository: BMNH HT ♂. Distribution : Brazil (Amazonas) . Note: distribution in Brazil (Asenjo et al. 2013).
Belonuchus arrogans Bernhauer 1916 View in CoL
Belonuchus arrogans Bernhauer 1916c: 274 View in CoL . Type locality: West-Columbien : Ost-Cordilleren, Pacho, 2000 m . Type depository: FMNH HT ♂. Distribution : Colombia. Note: generic affiliation unclear.
* Belonuchus asperatus Bernhauer 1906 View in CoL
Belonuchus asperatus Bernhauer 1906: 336 View in CoL . Type locality: Pachitea, Peru. Type depository: FMNH HT ♂. Distribution: Colombia, Ecuador, Peru, Bolivia. Notes: distribution ( Scheerpeltz 1960: 120; Asenjo 2004).
Belonuchus atticida ( Wasmann 1925) View in CoL
Plociopterus atticida Wasmann 1925: 71 View in CoL . Type locality: Colonia alpina, Rio d. Janeiro [LT]. Type depository: NHME LT ♂, #?PLT. Distribution: Brazil, Paraguay. Notes: comb. nov. ex Plociopterus View in CoL and lectotype designation ( Chatzimanolis & Chani-Posse 2016: 214); with Acromyrmex niger View in CoL , Atta sexdens View in CoL [orig.; Navarrete-Heredia 2001].
= Hesperus eidmanni Scheerpeltz 1936: 484 View in CoL . Type locality: Villa Rica in Paraguay [LT]; Umgebung von Mendes, E. d. Rio [4PLT; Brazil]. Type depository: NMW (Scheerpeltz) LT ♂, 4 PLT (des. Chatzimanolis & Chani- Posse 2016: 214). Distribution: Brazil, Paraguay. Notes: distribution within Brazil (Asenjo et al. 2013); as synonym of B. atticida ( Chatzimanolis & Chani-Posse 2016: 214) View in CoL ; in Atta sexdens View in CoL fungus chamber [orig.]; also Acromyrmex lobicornis View in CoL fungus chamber ( Navarrete-Heredia 2001).
Belonuchus avidus Erichson 1840 View in CoL
Belonuchus avidus Erichson 1840: 423 View in CoL . Type locality: Novae Valenciae in Columbia [ Venezuela, teste Herman 2001d: 1594]. Type depository: ZMHB 2 ST: 1 ♂, 1 ♀ [MCP seen]. Distribution: Colombia, Venezuela.
Belonuchus barbicornis Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) barbicornis Bernhauer 1916c: 271 View in CoL . Type locality: West-Columbien: Alto de las cruces, 2200m & S. Antonio, 2200m. Type depository: FMNH 5 About FMNH ST. Distribution : Colombia .
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus basiventris ( Sharp 1885) View in CoL
Philonthus apiciventris Sharp 1885: 418 View in CoL . Type locality: Mexico: Cordova, Puebla, Yolos, Oaxaca, Jalapa. Type depository: BMNH 2 ST. Distribution: Mexico. Notes: comb. nov. ex Philonthus ( Jiménez-Sánchez et al. 2000) View in CoL ; in Atta mexicana View in CoL external debris [ Navarrete-Heredia 2001; distribution ( Márquez & Asiain 2012).
* Belonuchus batesi Sharp 1876 View in CoL
Belonuchus batesi Sharp 1876a: 157 View in CoL . Type locality: Ega. Type depository: BMNH HT. Distribution : Brazil (Amazonas) . Note: distribution in Brazil (Asenjo et al. 2013).
* Belonuchus bidens Sharp 1885 View in CoL
Belonuchus bidens Sharp 1885: 430 View in CoL . Type locality: Mexico, Jalapa, Oaxaca, Mirador, Tuxtla; Nicaragua, Chontales. Type depository: BMNH 5 ST: from all type localities [MCP seen]. Distribution: Nicaragua, Mexico. Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2004, 2006, Márquez & Asiain 2012, Asiain et al. 2015).
Belonuchus bipunctatus Bernhauer 1935 View in CoL
Belonuchus bipunctatus Bernhauer 1935: 90 View in CoL . Type locality: Kolumbien; Cali (Villa Elvira, 1800m). Type depository: FMNH 2 ST; BMNH #?. Distribution: Colombia, Costa Rica.
* Belonuchus boops Sharp 1885 View in CoL
Belonuchus boops Sharp 1885: 435 View in CoL . Type locality: Panama, Bugaba . Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [ MCP seen]. Distribution : Panama .
Belonuchus boxi Bernhauer 1934 View in CoL
Belonuchus boxi Bernhauer 1934b: 212 View in CoL . Type locality: Tucumán . Type depository: FMNH 1 About FMNH ST ♂. Distribution : Argentina. Note: generic affiliation unclear.
Belonuchus brasilianus Bernhauer 1917 View in CoL
Belonuchus brasilianus Bernhauer 1917b: 19 View in CoL . Type locality: S. Catharina, Brasilien . Type depository: FMNH HT. Distribution : Brazil (Santa Catarina) . Note: distribution in Brazil (Asenjo et al. 2013); generic affiliation unclear.
Belonuchus breviceps Fauvel 1891 View in CoL
Belonuchus breviceps Fauvel 1891: 117 View in CoL . Type locality: Colonie Tovar [ Venezuela]. Type depository: IRSNB 2 ST [MCP seen]. Distribution: Venezuela. Note: generic affiliation unclear.
Belonuchus bruchi Bernhauer 1912 View in CoL
Belonuchus bruchi Bernhauer 1912b: 175 View in CoL . Type locality: Buenos Aires prov., Argentina. Type depository: FMNH HT. Distribution: Argentina. Note: not bruchi Bernhauer 1934b View in CoL (see duplicatus View in CoL ).
* Belonuchus bruchianus Bernhauer 1911 View in CoL
Belonuchus bruchianus Bernhauer 1911: 413 View in CoL . Type locality: Misiones prov., Argentina . Type depository: FMNH HT ♀. Distribution : Argentina .
Belonuchus bugnioni Fauvel 1901 View in CoL
Belonuchus bugnioni Fauvel 1901: 86 View in CoL . Type locality: Martinique. Type depository: IRSNB HT ♀ [MCP seen]. Distribution: Martinique, Guadeloupe. Note: distribution within (and endemic to) Lesser Antilles ( Peck 2011b: Martinique, Peck et al. 2014: Guadeloupe Archipelago). Note: generic affiliation unclear.
= Belonuchus eximius Bernhauer 1917c: 105 View in CoL . Type locality: Guadeloupe . Type depository: FMNH 2 About FMNH ST. Distribution : Guadeloupe. Note: as synonym of B. bugnioni ( Blackwelder 1943: 428) View in CoL .
Belonuchus cacao Blackwelder 1943 View in CoL
Belonuchus cacao Blackwelder 1943: 434 View in CoL . Type locality: Trinidad: one-half mile south-southwest of Manzanilla junction on the Plum Road. Type depository: USNM HT ♂, 1 PT ♀. Distribution: Venezuela, Trinidad. Note: Venezuela, biology, in Heliconia View in CoL bracts ( Frank & Barrera 2010).
Belonuchus candens ( Erichson 1840) View in CoL
Philonthus candens Erichson 1840: 460 View in CoL . Type locality: Bogotae in Columbia interiore. Type depository: ZMHB 1+3 ST: 1 ♂ in Erichson´s hand, 3 ♀ [MCP seen]. Distribution: Colombia, Brazil, Peru. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 369) View in CoL ; distribution including Peru, Brazil questioned (Asenjo et al. 2013).
Comment. Not Belonuchus , to be placed in a further study.
Belonuchus cariniventris Bernhauer 1918 View in CoL
Belonuchus cariniventris Bernhauer 1918: 225 View in CoL . Type locality: Mexico: Jalappa [Xalapa, Veracruz] . Type depository: FMNH 3 About FMNH ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002); generic affiliation unclear.
Belonuchus castaneus Chani-Posse 2006 View in CoL
Belonuchus castaneus Chani-Posse 2006: 366 View in CoL (new name for Belonuchus richteri Bernhauer 1922: 15 View in CoL ).
= Belonuchus richteri Bernhauer 1922: 15 View in CoL (preoccupied, junior secondary homonym in Belonuchus View in CoL of Philonthus richteri Bernhauer 1912a: 44 View in CoL ). Type locality: Misiones, Argentina. Type depository: FMNH LT ♂, 2 PLT (des. Chani-Posse 2011b: 219). Distribution: Argentina.
Belonuchus cavipennis Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) cavipennis Bernhauer 1916c: 267 View in CoL , attributed to Fauvel in litteris. Type locality: Yuracarès , Bolivien . Type depository: FMNH 1 About FMNH ST; ZMUH 1 View Materials +ST?. Distribution : Bolivia. Note: generic affiliation unclear.
* Belonuchus celatus ( Sharp 1885) View in CoL , comb. nov. ( Figs. 3–4 View FIGURES 1–12 )
Philonthus celatus Sharp 1885: 420 View in CoL . Type locality: Mexico: Mirador , Oaxaca, Tehuacan, Yolos, Cuernavaca. Type depository: BMNH 10 ST: from all type localities [ MCP seen]. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Remarks. Sharp did not specify the number of specimens he examined for naming this species, but it seems likely that he based his description on at least two specimens of different sex. Ten specimens were located in the BMNH under this name. All the ten specimens listed below agree with the original description by Sharp (1885) and are considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated (BMNH, Figs. 3–4 View FIGURES 1–12 ): card-mounted, “ Philonthus celatus Type D.S. Oaxaca. Höge” [in Sharp’s hand] // Type (round white label with red margin) // Oaxaca, Mexico. Hoege (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 2 ♀, one of them “Cuernavaca” (white label) [in Sharp’s hand], the other one “Yolos” (white label) [in Sharp’s hand], both of them with the following additional labels, “Mexico. Salle. Coll (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp. (white label)”; 1 ♂, 4 ♀, card-mounted, ♂ with “ Philonthus celatus Mirador. Höge ” [in Sharp’s hand]), “Mirador, Mexico. Hoege (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp. (white label)”; 2 ♀, card-mounted (one of them also “ Philonthus celatus Type D.S. Tehuacan. Höge” [in Sharp’s hand]), “Tehuacan, Mexico. Hoege (white label) // B. C. A. Col. I. 2. Philonthus celatus, Sharp. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
Belonuchus cephalotes ( Sharp 1885)
Musicoderus cephalotes Sharp 1885: 455 View in CoL . Type locality: Panama, Bugaba 800 to 1500 feet. Type depository: BMNH 12 ST: all of them with labels as "Bugaba" but only two of them with the elevation data [MCP seen]; FMNH 3 ST. Distribution: Panama, Venezuela. Notes: comb. nov. ex Musicoderus ( Scheerpeltz 1933: 1377) View in CoL ; Venezuela, biology, in Heliconia View in CoL bracts ( Frank & Barrera 2010); generic affiliation unclear.
* Belonuchus cifuentesi Rodríguez & Navarrete-Heredia 2016 View in CoL
Belonuchus cifuentesi Rodríguez & Navarrete-Heredia 2016: 445 View in CoL . Type locality: Mexico : Jalisco, Teocuitatlán de Corona, Cerro de García , 20°09´N 103°19´W, 2314 m. Type depository: CZUG HT ♂, 117 PT ( CZUG, MUD, CNIN) GoogleMaps . Distribution: Mexico. Note: in decomposing agave (Agave inaequidens) [orig.].
* Belonuchus clypeatus Sharp 1876 View in CoL
Belonuchus clypeatus Sharp 1876a: 160 View in CoL . Type locality: Ega . Type depository: BMNH HT. Distribution : Brazil (Amazonas) , Peru. Note: Peru (Newton 2015); distribution within Brazil (Asenjo et al. 2013).
Belonuchus coelestinus Bernhauer 1908 View in CoL
Belonuchus coelestinus Bernhauer 1908: 330 View in CoL . Type locality: St. Vincent. Type depository: NMW 1+ST, FMNH 1ST. Distribution: Lesser Antilles ( St. Vincent, Grenada). Note: distribution within Lesser Antilles ( St. Vincent) ( Peck 2010). Note: generic affiliation unclear.
= Belonuchus coeruleus Cameron 1922: 122 View in CoL . Type locality: St. Vincent. Type depository: BMNH 2ST. Distribution: Lesser Antilles ( St. Vincent, Grenada). Notes: as synonym of B. coelestinus ( Blackwelder 1943: 426) View in CoL .
= Belonuchus smithi Cameron 1922: 122 View in CoL (as synonym of B. coeruleus View in CoL ), attributed to Fauvel in litteris. Type locality: St. Vincent. Type depository: BMNH 2 ST. Distribution: Lesser Antilles ( St. Vincent, Grenada). Note: as synonym of B. coelestinus ( Blackwelder, 1943: 426) View in CoL ; unavailable ( ICZN 1999: Art. 11.6).
Belonuchus coelestis Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) coelestis Bernhauer 1916c: 266 View in CoL (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien . Type depository: FMNH 1 About FMNH ST; ZMUH 1 View Materials +ST? Distribution : Bolivia .
Comment. Not Belonuchus , to be placed in a further study.
Belonuchus coeruleispennis Motschulsky 1858 View in CoL
Belonuchus coeruleispennis Motschulsky 1858: 668 View in CoL . Type locality: Brésil intérieur. Type depository: ZMUM? Distribution: Brazil. Note: subsequently spelled coeruleipennis (e.g., Motschulsky 1870: 49; Bernhauer & Schubert 1914: 369), but coeruleispennis View in CoL maintained as correct spelling by Herman (2001d: 2522); distribution within Brazil unknown (Asenjo et al. 2013).
* Belonuchus cognatus Sharp 1885 View in CoL
Belonuchus cognatus Sharp 1885: 429 View in CoL . Type locality: British Honduras, Belize; Guatemala, El Reposo, Las Mercedes, San Isidro, Mirandilla; Nicaragua, Chontales; Panama, Bugaba. Type depository: BMNH #? ST: several, from all type localities [MCP seen]; IRSNB 5 ST: 4 Bugaba, two of them labelled as “Ext-ypis” [MCP seen]; 1 Chontales [MCP seen]; FMNH 3 ST: 3 Bugaba. Distribution: Belize, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Trinidad, Brazil, Argentina, St. Vincent. Notes: distribution within Lesser Antilles ( St. Vincent) ( Peck 2010); within (and including) Brazil (Asenjo et al. 2013).
* Belonuchus colon ( Sharp 1885) View in CoL
Philonthus colon Sharp 1885: 419 View in CoL . Type locality: Mexico: Cordova. Type depository: BMNH 2 ST. Distribution: Mexico. Notes: comb. nov. ex Philonthus View in CoL , distribution ( Navarrete-Heredia et al. 2002: 334); distribution within Mexico ( Márquez 2006).
Belonuchus columbicus Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) columbicus Bernhauer 1916c: 271 View in CoL . Type locality: West-Columbien: Umgebung von Cali ( St. Antonio , 2000m). Type depository: FMNH 2 About FMNH ST;? 1 ST. Distribution: Colombia .
Comment. Not Belonuchus , to be placed in a further study.
Belonuchus connexus ( Say 1830) View in CoL
Staphylinus connexus Say 1830: 35 View in CoL , also 1834: 448. Type locality: Mexico. Type depository:? #?ST. Distribution: Mexico. Note: comb. nov. ex Staphylinus ( Erichson 1840: 927) View in CoL ; distribution within Mexico (Navarrete- Heredia et al. 2002). Note: generic affiliation unclear.
Belonuchus convexus ( Bernhauer 1908) View in CoL
Musicoderus convexus Bernhauer 1908: 331 View in CoL . Type locality: Pozuzo, Peru. Type depository: FMNH HT ♀?: Distribution: Peru, Nicaragua. Notes: comb. nov. ex Musicoderus View in CoL , distribution Nicaragua ( Scheerpeltz 1933: 1377); Peru ( Asenjo 2004).
Belonuchus cordilleranus Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) cordilleranus Bernhauer 1916c: 270 View in CoL . Type locality: Columbien: Ost-Cordilleren, 450– 900m. Type depository: FMNH 3 About FMNH ST. Distribution : Colombia. Note: generic affiliation unclear.
* Belonuchus corticalis ( Sharp 1885) View in CoL , comb. nov. ( Figs. 5–6 View FIGURES 1–12 )
Philonthus corticalis Sharp 1885: 413 View in CoL . Type locality: Panama, Volcan de Chiriqui 2000–3000 ft . Type depository: BMNH HT ♂. Distribution : Panama .
Remarks. The “unique” specimen in BMNH is confirmed as the holotype.
Type material examined. Holotype ♂ ( BMNH, Figs. 5–6 View FIGURES 1–12 ): card-mounted, “ ♂ (in ink) Philonthus corticalis Type D.S. V. de Chiriqui 2–3000 ft. [in Sharp’s hand] // Type (round white label with red margin) // V. de Chiriqui 2– 3000 ft., 2–3000 ft. Champion (white label) // B. C. A. Col. I. 2. Philonthus corticalis, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
Belonuchus cribriceps Bernhauer 1921
Belonuchus cribriceps Bernhauer 1921c: 172 View in CoL . Type locality: San Roque, Corrientes [ Argentina]. Type depository: FMNH 1 ST. Distribution: Argentina, Brazil. Note: distribution within (and including) Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.
Belonuchus danforthi Blackwelder 1943 View in CoL
Belonuchus danforthi Blackwelder 1943: 429 View in CoL . Type locality: Puerto Rico: about 13 miles east of Maricao and north of Yauco, elevation about 3000 feet . Type depository: USNM HT ♂. Distribution : Puerto Rico .
* Belonuchus decipiens Sharp 1876 View in CoL
Belonuchus decipiens Sharp 1876a: 159 View in CoL . Type locality: Ega. Type depository: BMNH 6 ST, FMNH 1 ST. Distribution: Brazil, Argentina, Paraguay; Madagascar?; Indonesia (Lombok)? Notes: redescription ( Silvestri 1945); distribution, including Argentina and Paraguay ( Chani-Posse 2011b); records from Brazil only (Asenjo et al. 2013).
* Belonuchus dichrous Erichson 1840 View in CoL
Belonuchus dichrous Erichson 1840: 420 View in CoL . Type locality: Monte Orizaba in Mexico. Type depository: ZMHB 4 ST: 2 in Bernhauer´s hand " dichrous View in CoL ", 2 with "Orizaba" [MCP seen]. Distribution: Mexico, Guatemala, Venezuela. Notes: distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).
* Belonuchus diversicornis Bernhauer 1917 View in CoL
Belonuchus diversicornis Bernhauer 1917c: 104 View in CoL . Type locality: Espirito Santo, Brasilien . Type depository: FMNH HT. Distribution : Brazil (Espírito Santo) . Note: distribution within Brazil (Asenjo et al. 2013).
* Belonuchus docilis Sharp 1885 View in CoL
Belonuchus docilis Sharp 1885: 435 View in CoL . Type locality: Guatemala, Pantaleon; Panama, Bugaba, Tolé. Type depository: BMNH 8 ST: 2 Pantaleón, 3 Bugaba, 3 Tolé [MCP seen]. FMNH 2 ST: 2 Bugaba. Distribution: Guatemala,? Costa Rica, Panama, Brazil. Note: distribution within (and including) Brazil (Asenjo et al. 2013).
Belonuchus dominicus Blackwelder 1943 View in CoL
Belonuchus dominicus Blackwelder 1943: 431 View in CoL . Type locality: Dominica: Hillsboro Estate, at mouth of the Layou River, Leeward side. Type depository: USNM HT ♂, 4 PT; Blackwelder 1 PT; FMNH 1 PT. Distribution: Dominica Note: distribution within Dominica (Peck 2006); generic affiliation unclear.
Belonuchus duplicatus Blackwelder 1944 View in CoL
Belonuchus duplicatus Blackwelder 1944: 137 View in CoL (new name for Belonuchus bruchi Bernhauer 1934: 313 View in CoL ).
= Belonuchus bruchi Bernhauer 1934b: 213 View in CoL (preoccupied, junior primary homonym of Belonuchus bruchi Bernhauer 1912: 175 View in CoL ). Type locality: Tucuman, Argentina. Type depository: FMNH HT. Distribution: Argentina. Note: generic affiliation unclear.
Belonuchus ecuadorensis Bernhauer 1935 View in CoL
Belonuchus (Musicoderus) ecuadorensis Bernhauer 1935: 91 View in CoL . Type locality: Ecuador . Type depository: BMNH #?ST; FMNH 1 About FMNH ST. Distribution : Ecuador .
* Belonuchus ephippiatus ( Say 1830) View in CoL
Staphylinus ephippiatus Say 1830: 35 View in CoL , also 1834: 448. Type locality: Mexico. Type depository: MCZ NT ♂. Distribution: USA, Mexico, Colombia, Venezuela; adventive in Hawaii. Notes: comb. nov. ex Staphylinus ( Erichson 1840: 927) View in CoL ; neotype designation, redescription ( Smetana 1995: 746); distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).
= Belonuchus simplex Sharp 1885: 432 View in CoL . Type locality: Mexico: Oaxaca. Type depository: BMNH LT ♂, 2 PLT; FMNH 1 PLT?. Notes: as synonym of B. ephippiatus ( Fauvel 1901: 86) View in CoL ; lectotype designation ( Smetana 1995: 748).
= Belonuchus punctiventris Casey 1915: 446 View in CoL . Type locality: Benson, Arizona. Type depository: USNM LT ♂, 13 PLT. Notes: synonym of B. ephippiatus ( Smetana 1995: 746) View in CoL ; lectotype designation ( Smetana 1995: 748).
= Belonuchus laticeps Casey 1915: 446 View in CoL . Type locality: near Indio, California. Type depository: USNM LT ♂. Notes: synonym of B. ephippiatus ( Smetana 1995: 746) View in CoL ; lectotype designation ( Smetana 1995: 749).
= Belonuchus jacobianus Casey 1915: 447 View in CoL . Type locality: San Diego, California. Type depository: USNM LT ♂. Notes: synonym of B. ephippiatus ( Smetana 1995: 746) View in CoL ; lectotype designation ( Smetana 1995: 749).
= Belonuchus schaefferi Cooper 1933: 545 View in CoL . Type locality: Brownsville , Texas . Type depository: CUIC HT ♀; AMNH 1 About AMNH PT. Note : synonym of B. ephippiatus ( Smetana 1995: 746) View in CoL .
* Belonuchus erichsoni Bernhauer 1917 View in CoL
Belonuchus erichsoni Bernhauer 1917d: 223 View in CoL . Type locality: [Mexiko]. Type depository: FMNH HT ♂. Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).
* Belonuchus erythropterus Solsky 1868 View in CoL
Belonuchus erythropterus Solsky 1868: 137 View in CoL . Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #? ST, not seen by Smetana (1995). Distribution: USA (TX), Mexico. Note: redescription, distribution ( Smetana 1995: 752); distribution within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006).
Belonuchus excelsus Bernhauer 1917 View in CoL
Belonuchus excelsus Bernhauer 1917b: 19 View in CoL . Type locality: St. Antonio , 2000m, Umgebung von Cali, West- Columbien. Type depository: FMNH HT. Distribution: Colombia .
* Belonuchus extremus ( Sharp 1885) View in CoL , comb. nov. ( Figs. 7–8 View FIGURES 1–12 )
Philonthus extremus Sharp 1885: 415 View in CoL . Type locality: Guatemala, San Gerónimo. Type depository: BMNH HT. Distribution : Guatemala .
Remarks. The “unique” specimen in BMNH is confirmed as the holotype.
Type material examined. Holotype ♂ (BMNH, Figs. 7–8 View FIGURES 1–12 ): card-mounted, “ Philonthus extremus Type D.S. San Geronimo, Guatemala. Champion. [in Sharp’s hand] // Type (round white label with red margin) // San Geronimo, Verapaz. Champion. (white label) // B. C. A. Col. I. 2. Philonthus extremus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus falsus Bernhauer 1917
Belonuchus falsus Bernhauer 1917c: 105 View in CoL . Type locality: Venezuela . Type depository: FMNH 1 About FMNH ST; ZMUH 1 View Materials +ST?. Distribution : Venezuela .
* Belonuchus fassli Bernhauer 1917 View in CoL
Belonuchus fassli Bernhauer 1917a: 37 View in CoL . Type locality: West-Columbien: Umgebung von Cali am Rio Cauca (Cañon del Mte Tolima, 1700m). Type depository: FMNH HT. Distribution: Colombia .
Belonuchus flavicoxis (Bernhauer 1912) View in CoL
Philonthus flavicoxis Bernhauer 1912a: 43 View in CoL . Type locality: Tucuman prov., Argentina. Type depository: FMNH LT ♀. Distribution: Argentina. Notes: comb. nov. ex Philonthus View in CoL and lectotype designation ( Chani-Posse 2010: 8); distribution (Chani-Posse 2004); redescription ( Chani-Posse 2011b: 221).
* Belonuchus flavipennis Solsky 1870 View in CoL
Belonuchus flavipennis Solsky 1870: 263 View in CoL . Type locality: " Mexique " . Type depository: ZIN? HT ♀. Distribution : Mexico ; Costa Rica? , Panama? . Note: distribution, also within Mexico ( Navarrete-Heredia et al. 2002).
* Belonuchus flavocaudatus Bernhauer 1921 View in CoL
Belonuchus flavocaudatus Bernhauer 1921c: 170 View in CoL . Type locality: Hornadita , 3400m, Jujuy, Argentina . Type depository: FMNH 1 About FMNH ST; MACN 1 View Materials ST. Distribution : Argentina .
Belonuchus forsteri Scheerpeltz 1960 View in CoL
Belonuchus (Musicoderus) forsteri Scheerpeltz 1960: 115 View in CoL . Type locality: Bolivien, Yungas de Arepucho , Sihuencas, 2200–2500m . Type depository: NMW ( Scheerpeltz ) HT ♀. Distribution: Bolivia .
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus fossulatus Wasmann 1887 View in CoL
Belonuchus fossulatus Wasmann 1887: 414 View in CoL . Type locality: Blumenau (Prov. St. Catharina, Brasil) . Type depository: NHME (Wasmann) 1ST ♀. Distribution : Brazil (Santa Catarina) . Note: distribution within Brazil (Asenjo et al. 2013).
Belonuchus frater Bernhauer 1906 View in CoL
Belonuchus frater Bernhauer 1906: 333 View in CoL . Type locality: Peru . Type depository: FMNH 1 About FMNH ST. Distribution : Peru . Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.
* Belonuchus gagates Erichson 1840 View in CoL Belonuchus gagates Erichson 1840: 424 View in CoL . Type locality: Puerto Rico & Cuba. Type depository: ZMHB LT ♂, 7
PLT. Distribution: USA (FL), Cuba, Jamaica, Hispaniola, Puerto Rico, Cayman Is., St. Thomas, St. John, Montserrat, Bahamas. Note: lectotype designation, redescription and distribution ( Smetana 1995: 737); distribution, also within Cuba ( Peck 2005); distribution within (and including) Cayman Is. (Frank et al. 2011).
= Belonuchus gowdeyi Bernhauer 1935: 91 View in CoL . Type locality: Hope Gardens, Jamaica. Type depository: FMNH LT, BMNH PLT?. Distribution: Jamaica. Note: lectotype designation ( Smetana 1995: 739); as synonym of B. gagates ( Blackwelder 1943: 424) View in CoL .
= Belonuchus minax Erichson 1840: 424 View in CoL . Type locality: ins. St. Thomae & St. Johannis. Type depository: ZMHB 4 ST [MCP seen]. Distribution: Lesser Antilles (St. Thomas). Note: synonym of B. gagates ( Blackwelder 1943: 424) View in CoL ; not in Smetana (1995: 737).
= Musicoderus nigrocaeruleus Cameron 1913: 175 View in CoL . Type locality: Jamaica, Newcastle district, 3000 ft above sea level. Type depository: BMNH (Cameron) HT ♂. Distribution: Jamaica. Notes: as synonym of B. gagates ( Blackwelder 1943: 424) View in CoL ; although both male and female characters were used in the original description, Cameron (1913) only refers to a “type” deposited in his collection. Smetana (1995: 738) examined eight conspecific specimens under this name in the Cameron collection at the BMNH, one of them being the specimen chosen as “Type” by the author, thus the holotype.
Belonuchus ganglbaueri Bernhauer 1906 View in CoL
Belonuchus ganglbaueri Bernhauer 1906: 331 View in CoL . Type locality: Bolivien . Type depository: FMNH 1 About FMNH ST, NMW #? ST. Distribution : Bolivia. Note: generic affiliation unclear.
Belonuchus godmani ( Sharp 1885) View in CoL
Philonthus godmani Sharp 1885: 413 View in CoL . Type locality: Mexico: Guatemala: Calderas [LT]. Type depository: BMNH LT ♀, 7+ PLT. Distribution: Mexico, Guatemala, Costa Rica, Panama. Notes: comb. nov. ex Philonthus ( Navarrete-Heredia et al. 2002) View in CoL ; distribution, also within Mexico ( Navarrete-Heredia 2002, Márquez 2004, 2006); lectotype designation, redescription, distribution (Márquez 2011: 206).
Comment. Not Belonuchus , to be placed in a further study.
Belonuchus goeldii ( Wasmann 1925) View in CoL
Plociopterus goeldii Wasmann 1925: 72 . Type locality: Colonia alpina, Rio d. Janeiro [LT]. Type depository: NHME LT ♀, #? PLT. Distribution: Brazil, Paraguay, Bolivia. Notes: comb. nov. ex Plociopterus View in CoL and lectotype designation ( Chatzimanolis & Chani-Posse 2016: 215); with Acromyrmex niger View in CoL [orig.; Navarrete-Heredia 2001].
Belonuchus gracilis ( Sharp 1885)
Musicoderus gracilis Sharp 1885: 456 View in CoL . Type locality: Nicaragua, Chontales . Type depository: BMNH HT ♀. Distribution : Nicaragua .
* Belonuchus haemohrroidalis ( Fabricius 1801)
Staphylinus haemorrhoidalis Fabricius 1801: 596 View in CoL . Type locality: America meridionali. Type depository: ZMUK 4ST (until recently at ZMUC, Zimsen 1964): 3ST (former Sehested-Tønder Lund collection), 1ST (former Fabricius´s personal collection, seen AFN 2010). Distribution: Colombia, Venezuela, French Guiana, Peru, Bolivia, Brazil, Paraguay, Argentina. Notes: comb. nov. ex Staphylinus View in CoL , cited as haemoroidalis ( Nordmann 1837: 130); preoccupied, junior primary homonym of Staphylinus haemorrhoidalis Gmelin 1790: 2036 View in CoL and junior primary homonym of Staphylinus haemorrhoidalis Olivier 1795 View in CoL : (42) 11; ICZN application to conserve name ( Herman 2001a: 42), conserved in Opinion 2053 ( ICZN 2003); distribution including Peru, also within Brazil (Asenjo et al. 2013).
Belonuchus hispaniolus Blackwelder 1943 View in CoL
Belonuchus hispaniolus Blackwelder 1943: 422 View in CoL . Type locality: Dominican Republic: San Francisco Mountains . Type depository: USNM HT ♀, 1PT ♀. Distribution : Dominican Rep., Haiti. Note: Haiti ( Frank & Barrera 2010).
Belonuchus holisoides Sharp 1876 View in CoL
Belonuchus holisoides Sharp1876a: 161 View in CoL . Type locality: Ega , Brazil . Type depository: BMNH 2 ST: 1 Ega together with other 4 specimens from probably a different species, all of them under the name " Metoponcus holisoides Sharp View in CoL "[ MCP seen]. Distribution : Brazil (Amazonas) , Argentina. Note: distribution only in Brazil (Asenjo et al. 2013).
Comment. It probably belongs to Holisus Erichson.
Belonuchus iconnicoffi Plavilstschikov 1929 , nomen nudum
Belonuchus iconnicoffi Plavilstschikov 1929: 30 , as "n. sp. (Bernhauer in litteris)"; nomen nudum, no description. Distribution: Peru. Note: listed in error as valid species in Blackwelder (1944) and Asenjo (2004); not in Herman (2001d)
Belonuchus ielskii ( Solsky 1872) View in CoL
Philonthus ielskii Solsky 1872: 296 View in CoL . Type locality: Puna, Lima [Amérique du Sud]. Type depository: ZIN? (Solsky) #?ST. Distribution: Peru. Notes: comb. nov. ex Philonthus ( Bernhauer & Schubert 1914: 370) View in CoL ; as jelskii ( Herman 2001d: 2527, Asenjo 2004: 61). Note: generic affiliation unclear.
* Belonuchus impressifrons Sharp 1876 View in CoL
Belonuchus impressifrons Sharp 1876a: 162 View in CoL . Type locality: Ega. Type depository: BMNH HT. Distribution: Brazil ( Amazonas , São Paulo) . Note: distribution within Brazil (Asenjo et al. 2013).
* Belonuchus infimus ( Sharp 1885) View in CoL , comb. nov.
Philonthus infimus Sharp 1885: 414 View in CoL . Type locality: Guatemala, Las Mercedes 3000 ft, Chiacam; Panama, Volcan de Chiriqui. Type depository: BMNH 8 ST: 5 Las Mercedes, 2 Chiriqui, 1 Chiacam [MCP seen]. Distribution: Guatemala, Panama.
Remarks. Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on more than two specimens, including male/s and female/s. Eight specimens were located in the BMNH under this name. The specimens below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated (BMNH): card-mounted (with additional label “Sp. figured”), “ Philonthus infimus Las Mercedes 3000 ft. Guatemala Champion. [in Sharp’s hand] // Las Mercedes 3000 ft. Champion. (white label) // B. C. A. Col. I. 2. Philonthus infimus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 2 ♂, 2 ♀, card-mounted (one ♀ with additional “Type” round white label with red margin and “Type D.S” on the mounting card [in Sharp’s hand]), “ Philonthus infimus Las Mercedes 3000 ft. Guat. Champion. [in Sharp’s hand] // Las Mercedes 3000 ft. Champion. (white label) // B. C. A. Col. I. 2. Philonthus infimus, Sharp. (white label)”; 2 ♀, card-mounted, “ Philonthus infimus D.S. [in Sharp’s hand] // V. de Chiriqui 4–3000 ft Champion (white label) // B. C. A. Col. I. 2. Philonthus infimus, Sharp. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus iridescens Solsky 1868 View in CoL
Belonuchus iridescens Solsky 1868: 140 View in CoL . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca] . Type depository: ZIN? ( Solsky ) #? ST. Distribution: Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Belonuchus irregularis Bernhauer 1916 View in CoL
Belonuchus irregularis Bernhauer 1916c: 275 View in CoL . Type locality: West-Columbien: Pacho, 2000m, Ost-Cordilleren, West-Columbien. Type depository: FMNH HT. Distribution : Colombia. Note: generic affiliation unclear.
* Belonuchus iteratus ( Sharp 1887) View in CoL , comb. nov.
Philonthus iteratus Sharp 1887: 788 View in CoL . Type locality: Durango (Villa Lerdo). Type depository: BMNH 23 ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Remarks. Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on several specimens, including male/s and female/s. Twenty-three specimens were located in the BMNH under this name.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus jalappensis Bernhauer 1917
Belonuchus jalappensis Bernhauer 1917d: 224 View in CoL . Type locality: Mexico: Jalappa [Xalapa, Veracruz] . Type depository: FMNH 2 About FMNH ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Belonuchus laevipennis Bernhauer 1942 View in CoL
Belonuchus laevipennis Bernhauer 1942: 25 View in CoL . Type locality: Cartago ( Pino ), 1000m [ Costa Rica] . Type depository: FMNH 3 About FMNH ST. Distribution : Costa Rica. Note: generic affiliation unclear.
* Belonuchus latecinctus ( Sharp 1885) View in CoL , comb. nov. ( Figs. 9–10 View FIGURES 1–12 )
Philonthus latecinctus Sharp 1885: 415 View in CoL . Type locality: Guatemala, Cerro Zunil 4000–5000 ft, San Gerónimo. Type depository: BMNH 3 ST: 2 Cerro Zunil , 1 San Gerónimo [ MCP seen]. Distribution: Guatemala .
Remarks. Sharp based his description on three specimens which were located in the BMNH under this name. The specimens below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated (BMNH, Figs. 9–10 View FIGURES 1–12 ): card-mounted, “ Philonthus latecinctus Type D.S. Cerro Zunil 4–6000 ft. Guatem. Champion. [in Sharp’s hand] // Type (round white label with red margin) // Cerro Zunil, Guatemala. Champion. (white label) // B. C. A. Col. I. 2. Philonthus latecinctus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 1 ♂ with same locality and collection data as lectotype and additional printed white label “Sp. Figured”; 1 ♀, cardmounted, “ Philonthus latecinctus D.S. San Gerónimo Champion. [in Sharp’s hand] //San Gerónimo, Guatemala. Champion. (white label) // B. C. A. Col. I. 2. Philonthus latecinctus, Sharp. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus latro Erichson 1840 View in CoL
Belonuchus latro Erichson 1840: 422 View in CoL . Type locality: [no specific locality]. Type depository: ZMHB 1 ST ♀ [ MCP seen]. Distribution : Brazil (São Paulo) ; South America? Note: distribution within (and including) Brazil (Asenjo et al. 2013).
Belonuchus longiceps Bernhauer 1906 View in CoL
Belonuchus longiceps Bernhauer 1906: 332 View in CoL . Type locality: Peru. Type depository: FMNH HT. Distribution: Peru, Bolivia. Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.
* Belonuchus lucilius ( Sharp 1885) View in CoL , comb. nov.
Philonthus lucilius Sharp 1885: 415 View in CoL . Type locality: Panama, Volcan de Chiriqui 2000–3000 ft . Type depository: BMNH HT. Distribution : Panama .
Remarks. The “unique” specimen in BMNH is confirmed as the holotype.
Type material examined. Holotype ♂ ( BMNH): card-mounted, “ Philonthus lucilius Type D.S. V. de Chiriqui 2– 3000 ft . Champion. [in Sharp’s hand] // Type (round white label with red margin) // V. de Chiriqui 2–3000 ft . Champion. (white label) // B. C. A. Col. I. 2. Philonthus lucilius, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
Belonuchus magnificus Bernhauer 1916
Belonuchus magnificus Bernhauer 1916c: 277 View in CoL . Type locality: West-Kolumbien: Umgebung von Cali (S. Antonio, 2000m). Type depository: FMNH 3 About FMNH ST. Distribution : Colombia. Note: generic affiliation unclear.
Belonuchus meridionalis Bernhauer 1916 View in CoL
Belonuchus meridionalis Bernhauer 1916c: 269 View in CoL . Type locality: Argentinien [illeg. locality] & Bolivia. Type depository: FMNH 2 ST, ZMUH 1+ST. Distribution: Argentina, Bolivia. Note: generic affiliation unclear.
Belonuchus mexicanus Solsky 1864 View in CoL
Belonuchus mexicanus Solsky 1864: 440 View in CoL . Type locality: Mexique. Type depository: ZIN? (Solsky) HT. Distribution: Mexico, Brazil, Argentina. Notes: redescription ( Silvestri 1945); distribution within Mexico ( Navarrete-Heredia et al. 2002); distribution within Brazil (Asenjo et al. 2013).
= Belonuchus mexicanus var. femoralis Solsky 1864: 440 View in CoL . Type locality: Mexique. Type depository: ZIN? (Solsky) 2 ST. Distribution: Mexico. Notes: not in Bernhauer & Schubert (1914), Blackwelder (1944), Herman (2001d), but as synonym of B. mexicanus in Navarrete-Heredia et al. (2002: 335) View in CoL .
Belonuchus militaris Bernhauer 1921 View in CoL
Belonuchus militaris Bernhauer 1921c: 170 View in CoL . Type locality: Sao Paulo, Brazil . Type depository: FMNH 1 About FMNH ST. Distribution : Brazil (São Paulo) . Note: distribution within Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.
Belonuchus minor Bruch 1915 , nomen nudum
Belonuchus minor Bruch 1915: 504 , attributed to Bernhauer in litteris; nomen nudum. Locality: Misiones. Depository: MLPA (MLP 180). Distribution: Argentina. Note: see Fernández et al. (2007: 38).
Belonuchus miripennis Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) miripennis Bernhauer 1916c: 267 View in CoL . Type locality: Cañon del Mte. Tolima, 1700m, Umgebung von Cali am Cauca, Columbien. Type depository: FMNH 3 About FMNH ST. Distribution : Colombia .
Belonuchus modestus Fauvel 1891 View in CoL
Belonuchus modestus Fauvel 1891: 118 View in CoL . Type locality: Colonie Tovar [ Venezuela]. Type depository: IRSNB (Fauvel) 3 ST [MCP seen]. Distribution: Venezuela, Colombia. Note: generic affiliation unclear.
* Belonuchus montanellus Bernhauer 1916 View in CoL
Belonuchus montanellus Bernhauer 1916c: 272 View in CoL . Type localities: Cañon del Mte. Tolima, 1700m; Alto de las cruces, 2200m (Umgebung von Cali am Cauca). Type depository: FMNH 4 About FMNH ST. Distribution : Colombia .
Belonuchus montanicus Blackwelder 1944 View in CoL
Belonuchus montanicus Blackwelder 1944: 137 View in CoL (new name for Belonuchus montanus Bernhauer 1916: 273 View in CoL ).
= Belonuchus montanus Bernhauer 1916c: 273 View in CoL (preoccupied, junior primary homonym of Belonuchus montanus Fauvel 1895: 267 View in CoL ). Type localities: West-Columbien: Cañon del Mte. Tolima, 1700m; Rio Vitaco, 2200m; Villa Elvira, 1800m. Type depository: FMNH 3 ST. Distribution: Colombia.
Belonuchus monticola Bernhauer 1917 View in CoL
Belonuchus monticola Bernhauer 1917b: 18 View in CoL . Type locality: Rio Vitaco , 2200m, Umgebung von Cali am Rio Cauca, West-Columbien . Type depository: FMNH HT. Distribution : Colombia
* Belonuchus mordens Erichson 1840 View in CoL
Belonuchus mordens Erichson 1840: 422 View in CoL . Type locality: Brasilia. Type depository: ZMHB 1 ST ♂ [MCP seen]. Distribution: Peru, Brazil, Argentina, Bolivia. Notes: frequent guest of Melipona View in CoL nests in Brazil ( Wasmann 1904: 482, Kistner 1982); distribution, including Peru and Bolivia ( Chani-Posse 2011b), within Brazil (Asenjo et al. 2013).
Belonuchus moritzi Bernhauer 1906 View in CoL
Belonuchus moritzi Bernhauer 1906: 331 View in CoL . Type locality: Venezuela . Type depository: FMNH 2 About FMNH ST. Distribution : Venezuel. Note: generic affiliation unclear.
Belonuchus multipunctatus ( Coiffait 1981) View in CoL
Endeius multipunctatus Coiffait 1981: 297 View in CoL . Type locality: Galapagos: I. Santa Cruz versant S.E., basse altitude. Type depository: IRSNB HT ♂; IRSNB, MNHN 258 PT. Distribution: Galapagos Is. (Santa Cruz). Notes: comb. nov. ex Endeius (Chani Posse de Maus 2008: 386) View in CoL ; redescription ( Chani-Posse 2011b: 222); distribution within Galápagos Is. ( Newton & Peck 2006).
Belonuchus mundus Erichson 1840 View in CoL
Belonuchus mundus Erichson 1840: 425 View in CoL . Type locality: ins. St. Vincentii. Type depository: ZMHB? #?ST. Distribution: Lesser Antilles ( St. Lucia, St. Vincent, Grenada), Brazil. Notes: distribution within Lesser Antilles ( St. Vincent) ( Peck 2010); within (and including) Brazil (Asenjo et al. 2013).
* Belonuchus muticus (Sharp 1876) View in CoL , comb. nov. ( Figs. 11–12 View FIGURES 1–12 )
Philonthus muticus Sharp 1876a: 167 View in CoL . Type locality: Ega. Type depository: BMNH 4 ST: 3 Ega , 1 Sao Paulo [ MCP seen]. Distribution: Brazil (Amazonas) . Note: distribution within Brazil (Asenjo et al. 2013).
Remarks. Sharp explicitly mentioned four specimens of both sexes as part of the type series. The specimens listed below agree with the original description by Sharp (1876a) and is considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 11–12 View FIGURES 1–12 ): card-mounted, “ ♂ // Type (round white label with red margin) // Ega (round green label) [in Sharp’s hand]// S America Brazil // Philonthus muticus Type D.S. [in Sharp’s hand] // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 2 ♀ (one without head), card mounted, same locality and collection data as lectotype but with “ SYNTYPE ” (rounded white label with blue margin).
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus nigerrimus Bernhauer 1918 View in CoL
Belonuchus nigerrimus Bernhauer 1918: 225 View in CoL . Type locality: Mexico: Jalappa [Xalapa, Veracruz] . Type depository: FMNH 2 About FMNH ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Belonuchus oakleyi Blackwelder, 1943 View in CoL
Belonuchus oakleyi Blackwelder 1943: 422 View in CoL . Type locality: Puerto Rico: Mayagüez . Type depository: USNM HT. Distribution : Puerto Rico .
* Belonuchus optatus ( Sharp, 1885) View in CoL , comb. nov. ( Figs. 13–14 View FIGURES 13–24 )
Philonthus optatus Sharp 1885: 427 View in CoL . Type locality: Mexico : Guanajuato . Type depository: BMNH HT ♀. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Remarks. The “unique” specimen in BMNH is confirmed as the holotype.
Type material examined. Holotype ♀ (BMNH, Figs. 13–14 View FIGURES 13–24 ): card-mounted, “ Philonthus optatus Type D.S. Guanajuato. Mexico. Sallé. [in Sharp’s hand] // Type (round white label with red margin) //Guanajuato (white label) // Mexico. Salle Coll. (white label) // B. C. A. Col. I. 2. Philonthus optatus, Sharp. (white label) // Belonuchus formosus apud Sallé [in Sallé’s hand]”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus oxyporinus ( Sharp 1885) View in CoL
Philonthus oxyporinus Sharp 1885: 419 View in CoL . Type locality: Mexico: Oaxaca. Type depository: BMNH HT ♀. Distribution: Mexico. Notes: to Belonuchus View in CoL (after Smetana 1995); distribution, biology (Navarrete-Heredia 1995, Jiménez-Sánchez et al. 2000); distribution within Mexico ( Navarrete-Heredia et al. 2002).
Belonuchus palustris Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) palustris Bernhauer 1916c: 268 View in CoL . Type locality: Alto de las Cruces, 2200m. Type depository: FMNH HT. Distribution : Colombia .
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus panamensis Sharp 1885 View in CoL
Belonuchus panamensis Sharp 1885: 434 View in CoL . Type localities: Nicaragua, Chontales; Panama, Volcan de Chiriqui, Bugaba, Tolé. Type depository: BMNH 11 ST: from all type localities [MCP seen]. Distribution: Nicaragua, Costa Rica, Panama.
Belonuchus paradoxipunctatus Bernhauer 1921 View in CoL
Belonuchus paradoxipunctatus Bernhauer 1921c: 171 View in CoL . Type locality: San Roque, Corrientes [ Argentina]. Type depository: FMNH 1 ST. Distribution: Argentina, Brazil. Note: distribution within (and including) Brazil (Asenjo et al. 2013). Note: generic affiliation unclear.
Belonuchus paradoxus Bernhauer 1906 View in CoL
Belonuchus paradoxus Bernhauer 1906: 333 View in CoL . Type locality: Pachitea , Peru . Type depository: FMNH HT. Distribution : Peru . Note: Peru ( Asenjo 2004).
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus pectinipes Bernhauer 1910 View in CoL
Belonuchus pectinipes Bernhauer 1910: 379 View in CoL . Type locality: Cordoba, Veracruz . Type depository: FMNH 4 About FMNH ST. Distribution : Mexico . Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Belonuchus peruvianus Plavilstschikov 1929 , nomen nudum
Belonuchus peruvianus Plavilstschikov 1929: 30 , as "n. sp. (Bernhauer in litteris)"; nomen nudum, no description. Distribution: Peru. Note: as valid sp. in Blackwelder (1944) and Asenjo (2004); not in Herman (2001d).
* Belonuchus piceolus Bernhauer 1935 View in CoL
Belonuchus piceolus Bernhauer 1935: 90 View in CoL . Type locality: Brasilien . Type depository: BMNH #?ST; FMNH 1 About FMNH ST. Distribution : Brazil . Note: distribution within Brazil unknown (Asenjo et al. 2013).
Belonuchus pictipennis Sharp 1885 View in CoL
Belonuchus pictipennis Sharp 1885: 436 View in CoL . Type locality: Bugaba , Panama . Type depository: BMNH 2 ST ♂. Distribution : Panama .
Comment. It may belong to Paederomimus Sharp (Sharp: 436, “very closely allied to B. mimeticus ”) but see Paederomimus mimeticus (Sharp) .
Belonuchus planiceps Erichson 1840 View in CoL
Belonuchus planiceps Erichson 1840: 425 View in CoL . Type locality: Columbia. Type depository: ZMHB 4 ST [MCP seen]. Distribution: Costa Rica, Colombia, Venezuela.
Comment. It probably belongs to Holisus Erichson (Hyptiomina) .
* Belonuchus platypterus ( Sharp 1885) View in CoL , comb. nov. ( Figs. 15–16 View FIGURES 13–24 )
Philonthus platypterus Sharp 1885: 412 View in CoL . Type locality: Guatemala, Cerro Zunil 4 000 – 5000 feet. Type depository: BMNH 3 ST. Distribution : Guatemala .
Remarks. Sharp explicitly mentioned three specimens (two males and one female) as part of the type series. The specimens listed below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 15–16 View FIGURES 13–24 ): card-mounted, “ ♂ (in ink) Philonthus platypterus Type D.S. Cerro Zunil. 4–5000 ft. [in Sharp’s hand] // Type (round white label with red margin) // Cerro Zunil. Guatemala. Champion. // B. C. A. Col. I. 2. Philonthus platypterus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 1 ♂, 1 ♀, same locality and collection data as lectotype but male with additional printed white label “Sp. figured”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus pollens Sharp 1885 View in CoL
Belonuchus pollens Sharp 1885: 431 View in CoL . Type localities: North America; Mexico: Oaxaca; Guatemala: near the city; Honduras. Type depository: BMNH 8 ST: from all type localities except North America [MCP seen]; FMNH 2 ST. Distribution: Mexico, Guatemala, Honduras ["North America" in error]. Notes: not in USA or Canada ( Smetana 1995: 733); distribution within Mexico ( Navarrete-Heredia et al. 2002).
Belonuchus posticalis Laporte de Castelnau 1840 View in CoL
Staphylinus posticalis Laporte de Castelnau 1840: 176 View in CoL . Type locality: Brésil . Type depository: ?? #? ST. Distribution : Brazil . Note: comb. nov. ex Staphylinus ( Bernhauer & Schubert 1914: 371) View in CoL ; distribution within Brazil unknown (Asenjo et al. 2013).
Belonuchus prasinipennis Bernhauer 1917 View in CoL
Belonuchus prasinipennis Bernhauer 1917b: 18 View in CoL (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien . Type depository: FMNH 1 About FMNH ST; ZMUH 1 View Materials ST. Distribution : Bolivia .
Belonuchus pulchripennis Bernhauer 1908 View in CoL
Belonuchus pulchripennis Bernhauer 1908: 330 View in CoL . Type locality: Montana de Nancho, 7000', central Peru . Type depository: FMNH 2 About FMNH ST; ?? 1 ST. Distribution : Peru . Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.
Belonuchus purpuripennis Bernhauer 1916 View in CoL
Belonuchus purpuripennis Bernhauer 1916c: 278 View in CoL . Type locality: West-Columbien: Umgebung Cali am Rio Cauca, St. Antonio, 2000 m. Type depository: FMNH 2 ST. Distribution: Colombia, Bolivia. Note: generic affiliation unclear.
Belonuchus richteri (Bernhauer, 1912) View in CoL
Philonthus (Chroaptomus) richteri Bernhauer 1912a: 44 View in CoL . Type locality: Tucuman prov., Argentina. Type depository: FMNH HT ♀. Distribution: Argentina. Notes: comb. nov. ex Chroaptomus ( Chani-Posse 2006: 365) View in CoL ; redescription ( Chani-Posse 2011b: 223).
Comment. Not Belonuchus , to be placed in a further study.
* Belonuchus rufipennis ( Fabricius 1801) View in CoL
Staphylinus rufipennis Fabricius 1801: 596 View in CoL . Type locality: America meridionali. Type depository: ZMUK 3 ST (until recently at ZMUC, Zimsen 1964), not seen by Smetana (1995). Distribution: Canada, USA, Mexico, Guatemala, Belize, El Salvador, Nicaragua, Panama, Colombia, Venezuela, French Guiana, Trinidad, Tobago, Jamaica, Peru, Brazil, Argentina, Galapagos Is. (Isabela, S. Crist., S. Cruz), Hawaii; introduced for biocontrol in Italy but not established. Notes: as synonym of B. formosus ( Erichson 1840: 426) View in CoL ; introduction for biocontrol, biology, immature stages ( Silvestri 1945); redescription and distribution ( Smetana 1995: 726); in Atta mexicana View in CoL external debris (occasional) ( Navarrete-Heredia 2001); distribution, also within Mexico ( Navarrete-Heredia et al. 2002, Márquez 2006); distribution within Galapagos Is. ( Newton & Peck 2006); distribution within Brazil (Asenjo et al. 2013); in Heliconia View in CoL bracts ( Frank & Barrera 2010); distribution in carcasses in Argentina ( Aballay et al. 2014).
= Staphylinus formosus Gravenhorst 1806: 72 View in CoL . Type depository: ZMHB LT ♀, 1 PLT. Type locality: Carolina, USA (" Carolina "). Note: as synonym of B. rufipennis View in CoL ( Fauvel 1877: xxvii). Note: lectotype designation ( Smetana 1995: 728).
= Belonuchus pallipes Melsheimer 1844: 35 View in CoL . Type depository: MCZ? #?ST. Type locality: Pennsylvania. Distribution: USA. Note: as synonym of B. rufipennis View in CoL ( Fauvel 1877: xxvii); no specimens under this name in the Melsheimer collection at the MCZ ( Smetana 1995: 728).
= Cafius apicalis Dejean 1833: 63 View in CoL , unavailable, published in synonymy of unavailable name ( ICZN 1999: Art. 11.6). Distribution: Jamaica. Note: as synonym of B. formosus ( Erichson 1840: 426) View in CoL .
* Belonuchus rufiventris ( Sharp 1887) View in CoL , comb. nov. ( Figs. 17–18 View FIGURES 13–24 )
Philonthus rufiventris Sharp 1887: 787 View in CoL . Type locality: Mexico: "Amecameca in Morelos " (=Est. Mexico?). Type depository: BMNH HT ♀ . Distribution: Mexico. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002).
Remarks. The unique female specimen in BMNH is confirmed as the holotype.
Type material examined. Holotype ♀ (BMNH, Figs. 17–18 View FIGURES 13–24 ): card-mounted, “ Philonthus rufiventris Type D.S. Amecameca. Hoge [in Sharp’s hand] // Type (round white label with red margin) // Amecameca, Morelos. Höge. // B. C. A. Col. I. 2. Philonthus rufiventris, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus rufocaudus ( Sharp 1885) View in CoL , comb. nov. ( Figs. 19–20 View FIGURES 13–24 )
Philonthus rufocaudus Sharp 1885: 412 View in CoL . Type locality: Panama, Bugaba , Volcan in Chiriqui . Type depository: BMNH 8 ST: 5 Bugaba , 3 Chiriqui [ MCP seen]. Distribution: Panama .
Remarks. Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on at least one male and one female. Eight specimens were located in the BMNH under this name. The specimens below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 19–20 View FIGURES 13–24 ): card-mounted, “ ♂ (in ink) Philonthus rufocaudus Type D.S. Panama. Bugaba. Champion [in Sharp’s hand] // Type (round white label with red margin) // Bugaba , Panama. Champion. // B. C. A. Col. I. 2. Philonthus rufocaudus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 1 ♂, 3 ♀, card-mounted, with same locality and collection data as lectotype ; 3 ♀, card-mounted, “V. de Chriqui 25–4,000 ft. Champion (white label) // B. C. A. Col. I. 2. Philonthus rufocaudus, Sharp. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
* Belonuchus rufopygus ( Sharp 1885) View in CoL , comb. nov. ( Figs. 21–22 View FIGURES 13–24 )
Philonthus rufopygus Sharp 1885: 411 View in CoL . Type locality: Panama: Bugaba. Type depository: BMNH 8 ST. Distribution: Panama [ Mexico in error]. Note: reported from Mexico ( Zaragoza Caballero 1976, but based on misidentified specimens [J.L. Navarrete-Heredia, unpublished]).
Remarks. Sharp explicitly mentioned several specimens as part of the type series. The specimens listed below agree with the original description by Sharp (1885) and considered to have been syntypes prior to the present lectotype designation.
Type material examined. Lectotype ♂, here designated ( BMNH, Figs. 21–22 View FIGURES 13–24 ): card-mounted, “ ♂ (in ink) Philonthus rufo-pygus Type D.S. Panama. Bugaba. Champion [in Sharp’s hand] // Type (round white label with red margin) // Bugaba , Panama. Champion. // B. C. A. Col. I. 2. Philonthus rufopygus, Sharp. (white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes ( BMNH): 7 ♀ (2 of them mounted together), cardmounted, with same locality and collection data as lectotype .
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
Belonuchus rugicollis Bernhauer 1916
Belonuchus rugicollis Bernhauer 1916c: 276 View in CoL . Type locality: Peru . Type depository: FMNH HT Distribution: Peru . Note: Peru ( Asenjo 2004). Note: generic affiliation unclear.
* Belonuchus satyrus Erichson 1840 View in CoL
Belonuchus satyrus Erichson 1840: 421 View in CoL . Type locality: Columbia. Type depository: ZMHB? #?ST. Distribution: Panama, Colombia, Venezuela, Trinidad; Costa Rica. Notes: biology, in Heliconia View in CoL bracts ( Frank & Barrera 2010).
* Belonuchus serraticornis (Sharp 1876) View in CoL , comb. nov. ( Figs. 23–24 View FIGURES 13–24 )
Philonthus serraticornis Sharp 1876a: 179 View in CoL . Type locality: Ega. Type depository: BMNH HT. Distribution : Brazil (Amazonas) . Note: distribution within Brazil (Asenjo et al. 2013).
Remarks. The “unique” specimen in BMNH is confirmed as the holotype.
Type material examined. Holotype ♂ (incomplete genitalia) ( BMNH, Figs. 23–24 View FIGURES 13–24 ): card-mounted, “ ♂ // Type (round white label with red margin) // Ega (round green label) [in Sharp’s hand]// S America Brazil // Philonthus serraticornis Type D.S. [in Sharp’s hand] // Sharp Coll. 1905.-313. (white label)”.
Comments. In accordance with the diagnosis of Belonuchus ( Chani-Posse et al. 2018) .
Belonuchus setiger Sharp 1876a: 164 View in CoL . Type locality: Ega. Type depository: BMNH 2 ST: 2 ♂ [ MCP seen]. Distribution : Brazil (Amazonas) . Note: distribution within Brazil (Asenjo et al. 2013).
* Belonuchus sharpi Bernhauer 1907 View in CoL
Belonuchus sharpi Bernhauer 1907: 285 View in CoL (replacement name for Belonuchus grandiceps Sharp 1876a: 158 View in CoL ). Distribution: Brazil (Amazonas, Pará), Peru. Note: distribution within Brazil (Asenjo et al. 2013).
= Belonuchus grandiceps Sharp 1876a: 158 View in CoL (preoccupied, junior secondary homonym in Belonuchus View in CoL of Philonthus grandiceps Kraatz 1859a: 95 View in CoL ). Type locality: Ega, Tapajos & St. Paulo [Amazonas]. Type depository: BMNH 5 ST. Note: Peru ( Asenjo 2004).
Belonuchus silvestris Bernhauer 1917 View in CoL
Belonuchus silvestris Bernhauer 1917b: 17 View in CoL . Type locality: Alto de las Cruces, 2200m, Cali am Rio Cauca, West- Columbien. Type depository: FMNH HT. Distribution : Colombia .
Belonuchus similis Sharp 1885 View in CoL
Belonuchus similis Sharp 1885: 436 View in CoL . Type locality: Panama, Volcan de Chiriqui 2500–6000 ft . Type depository: BMNH 3 ST; FMNH 1 About FMNH ST. Distribution : Panama. Note: generic affiliation unclear.
Belonuchus simillimus Bernhauer 1917 View in CoL
Belonuchus simillimus Bernhauer 1917c: 106 View in CoL . Type locality: Villa Elvira 1800m, Westkolumbien. Type depository: FMNH HT. Distribution : Colombia .
Belonuchus skalitskyi Bernhauer 1906
Belonuchus skalitskyi Bernhauer 1906: 334 . Type locality: Pumananea, Peru. Type depository: FMNH HT. Distribution: Peru, Bolivia. Note: Peru ( Asenjo 2004).
Comment. Not Belonuchus , to be placed in a further study.
Belonuchus spinicornis ( Champion 1913) View in CoL
Musicoderus spinicornis Champion 1913: 4 View in CoL . Type locality: Costa Rica, Pitahaya , 1400m . Type depository: BMNH 2 ST: 1 ♂, 1 ♀ [ MCP seen]. Distribution : Costa Rica .
* Belonuchus stenoderus Sharp 1885 View in CoL
Belonuchus stenoderus Sharp 1885: 434 View in CoL . Type locality: Panama, Bugaba. Type depository: BMNH 4 ST; FMNH 1 ST. Distribution: Costa Rica, Panama. Note: the syntype in FMNH does not belong to the ‘ Belonuchus haemorrhoidalis View in CoL clade’ ( Chani-Posse et al. 2018).
Belonuchus subaeneipennis Bernhauer 1916 View in CoL
Belonuchus (Musicoderus) subaeneipennis Bernhauer 1916c: 269 View in CoL . Type locality: West-Columbien: Umgebung von Cali am Rio Cauca (Cañon del Mte Tolima, 1700m). Type depository: FMNH 1 About FMNH ST;? 1 ST. Distribution : Colombia .
Belonuchus subaeneus Bernhauer 1916 View in CoL
Belonuchus subaeneus Bernhauer 1916c: 268 View in CoL (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien . Type depository: FMNH 1 About FMNH ST; ZMUH 1 View Materials +ST?. Distribution : Bolivia .
Belonuchus subsellatus Bernhauer 1916 View in CoL
Belonuchus subsellatus Bernhauer 1916c: 276 View in CoL (attributed to Fauvel in litt.). Type locality: Yuracarès , Bolivien . Type depository: FMNH 1 About FMNH ST, ZMUH 1 View Materials + ST. Distribution : Bolivia. Note: generic affiliation unclear.
Belonuchus superbus Bernhauer 1916 View in CoL
Belonuchus superbus Bernhauer 1916c: 273 View in CoL . Type localities: Alto de las cruces, 2200m ; Rio Vitaco , 2000m ; Rio Aquatal , 1800m ; S. Antonio, 2000m; Villa Elvira, 1800m. Type depository: FMNH 3 About FMNH ST;? 2 ST. Distribution : Colombia. Note: generic affiliation unclear.
* Belonuchus supernus (Herman 2001) View in CoL , comb. nov. ( Figs. 47–48 View FIGURES 45–52 )
Philonthus supernus Herman 2001a: 48 View in CoL (new name for Philonthus excelsus Bernhauer 1941: 287 View in CoL ). Distribution: Peru.
= Philonthus excelsus Bernhauer 1941: 287 View in CoL , also 1951: 277 (preoccupied, junior secondary homonym in Philonthus View in CoL of Hesperus excelsus Cameron 1931: 361 View in CoL , and junior primary homonym of Philonthus excelsus Cameron 1932: 147 View in CoL ). Type locality: Ocaña, 3100m, etc., s Peru, Oberh. Paleo 3240m, S Peru. Type depository: FMNH 21 ST: 1 Ocaña. Note: Peru ( Asenjo 2004).
Remarks. Although references about three different type localities on different dates and a male character are given in the original description, there is no indication about how many specimens were examined. Two specimens were located in the FMNH under this name. The specimen below agrees with the original description and considered to have been a syntype.
Type material examined. Syntype ♂ (FMNH, Figs. 47–48 View FIGURES 45–52 ): card mounted with cutouts at head-thorax and abdomen // Süd – Peru Oberh. Paleo, 3240m 10.4.1936 [in Bernhauer’s hand] (green label) // Hbg. – Süd – Peru Sammelreise 1936. Eing. Nr. 1, 1937 (green label) // 219 (white label) // Philonthus excelsus Brh. Cotypus (white label) // excelsus Benh. Typus Philonthus [in Bernhauer’s hand](red label) // Chicago NHMus M.Bernhauer Collection (white label)”.
* Belonuchus taczanovskii ( Solsky 1872) View in CoL
Philonthus taczanovskii Solsky 1872: 303 . Type locality: Monte Rico, Perou. Type depository: ZIN? (Solsky) #?ST. Distribution: Peru, Ecuador. Note: Peru ( Asenjo 2004).
Belonuchus terminalis (Laporte de Castelnau 1840) View in CoL
Staphylinus terminalis Laporte de Castelnau 1840: 176 View in CoL . Type locality: Brésil. Type depository:? #? ST. Distribution: Brazil. Notes: comb. nov. ex Staphylinus ( Bernhauer & Schubert 1914: 371) View in CoL ; preoccupied, junior primary homonym of Staphylinus terminalis Erichson 1839: 396 View in CoL ; conserved by Opinion 2053 ( ICZN 2003); distribution within Brazil unknown (Asenjo et al. 2013).
* Belonuchus tremolerasi Bernhauer 1912 View in CoL
Belonuchus tremolerasi Bernhauer 1912a: 40 View in CoL . Type locality: Uruguay. Type depository: FMNH 1 ST. Distribution: Uruguay, Argentina, Paraguay.
Belonuchus trinitatis Blackwelder 1943 View in CoL
Belonuchus trinitatis Blackwelder 1943: 423 View in CoL . Type locality: Trinidad: Department of Agriculture Stock Farm at St . Joseph, 6 miles east of Port of Spain . Type depository: USNM HT ♂, 4 PT; FMNH 1 About FMNH PT. Distribution : Trinidad .
* Belonuchus trochanterinus ( Sharp 1885) View in CoL
Philonthus trochanterinus Sharp 1885: 417 View in CoL . Type locality: Guatemala, Capetillo. Type depository: BMNH #? ST. Distribution: Mexico, Guatemala. Note: comb. nov. ex Philonthus View in CoL , distribution within Mexico (Navarrete- Heredia et al. 2002: 335).
Belonuchus tucumanensis Bernhauer 1934 View in CoL
Belonuchus tucumanensis Bernhauer 1934b: 213 View in CoL . Type locality: Tucumán. Type depository: FMNH 1 About FMNH HT. Distribution : Argentina .
* Belonuchus variolosus Solsky 1870 View in CoL
Belonuchus variolosus Solsky 1870: 263 View in CoL . Type locality: " Cayenne " [ French Guiana] . Type depository: ZIN? ( Solsky ) #? ST. Distribution : French Guiana (Cayenne). Note: generic affiliation unclear.
= Belonuchus variolaris View in CoL ; Solsky 1872: 305, misspelling.
Belonuchus venezolanus Bernhauer 1917
Belonuchus venezolanus Bernhauer 1917c: 104 View in CoL . Type localities: Venezuela & Columbien. Type depository: FMNH 3ST; NMW 1+ST?; ZMUH 1+ST?. Distribution: Venezuela, Colombia. Note: generic affiliation unclear.
Belonuchus viridipennis Baudi di Selve 1848 View in CoL
Belonuchus viridipennis Baudi di Selve 1848: 125 View in CoL . Type locality: Novae Valenciae in Columbia [ Venezuela, teste Herman 2001c: 1594]. Type depository: MRSN/MCSN? #?ST. Distribution: Mexico, Guatemala, Panama, Colombia, Venezuela. Note: distribution within Mexico ( Navarrete-Heredia et al. 2002). Note: generic affiliation unclear (based on non-type material in FMNH).
= Belonuchus dejectus Sharp 1885: 433 View in CoL . Type localities: Mexico: Cordova, Jalapa, Las Vigas, Mirador; Guatemala, Capetillo, Cerro Zunil, Balheu, Tamahu, San Joaquin, Purula, El Tumbador, San Gerónimo; Panama: Volcan de Chiriqui 2500–4000 ft. Type depository: BMNH 14+ST. Distribution: Mexico, Guatemala, Panama. Note: as synonym of B. viridipennis ( Sharp 1887: 789) View in CoL .
Belonuchus vulneratus Bernhauer 1906 View in CoL
Belonuchus vulneratus Bernhauer 1906: 335 View in CoL . Type locality: Pachitea , Peru . Type depository: FMNH HT. Distribution : Peru . Note: Peru ( Asenjo 2004). Note: not Belonuchus View in CoL , to be placed in a further study.
Belonuchus weiserianus (Bernhauer 1921) View in CoL
Philonthus weiserianus Bernhauer 1921c: 106 View in CoL . Type locality: Hornadita, 3400m, Jujuy, Argentina. Type depository: FMNH LT ♀. Distribution: Argentina. Note: comb. nov. ex Philonthus View in CoL and lectotype designation ( Chani-Posse 2010: 8); redescription ( Chani-Posse 2011b: 224).
Belonuchus xantholinoides Bernhauer 1917 View in CoL
Belonuchus xantholinoides Bernhauer 1917c: 106 View in CoL . Type localities: Villa Elvira 1800m & S. Antonio 2200m, Westkolumbien. Type depository: FMNH 2 About FMNH ST. Distribution : Colombia .
* Belonuchus xanthomelas Solsky 1868 View in CoL
Belonuchus xanthomelas Solsky 1868: 139 View in CoL . Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #?ST. Distribution: USA (TX), Mexico. Note: generic placement (Navarrete- Heredia 1995); distribution, also within Mexico ( Navarrete-Heredia et al. 2002; Márquez-Luna 2001, Márquez 2006).
* Belonuchus xanthopterus ( Nordmann 1837) View in CoL
Philonthus xanthopterus Nordmann 1837: 99 View in CoL . Type locality: Parae in America meridionali. Type depository: ZMHB 1 ST ♂ [MCP seen]. Distribution: Brazil, Peru, Panama?, Ecuador?. Note: comb. nov. ex Philonthus ( Erichson 1840: 421) View in CoL ; Peru ( Soukup 1942: 177); distribution within Brazil (Asenjo et al. 2013).
* Belonuchus xanthopus Solsky 1868 View in CoL
Belonuchus xanthopus Solsky 1868: 139 View in CoL . Type locality: [ Mexique et probablement de l'Etat d'Oaxaca] . Type depository: ZIN? ( Solsky ) #? ST. Distribution: Mexico . Notes: generic placement (Navarrete-Heredia 1995); distribution within Mexico ( Navarrete-Heredia et al. 2002).
* Belonuchus zunilensis ( Sharp 1885) View in CoL
Philonthus zunilensis Sharp 1885: 418 View in CoL . Type locality: Guatemala: Cerro Zunil 4000–5000 ft. Type depository: BMNH 4 ST. Distribution: Mexico, Guatemala. Notes: comb. nov. ex Philonthus View in CoL , distribution, also within Mexico ( Navarrete-Heredia et al. 2002); distribution ( Asiain et al. 2011).
MCP |
Pontificia Universidade Catolica do Rio Grande do Sul |
FMNH |
Field Museum of Natural History |
USNM |
Smithsonian Institution, National Museum of Natural History |
CZUG |
Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia |
CNIN |
Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico |
CUIC |
Cornell University Insect Collection |
ZIN |
Russian Academy of Sciences, Zoological Institute, Zoological Museum |
NMW |
Naturhistorisches Museum, Wien |
NHME |
Natuurhistorisch Museum |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
|
Phylum |
|
Class |
|
Order |
|
Family |
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SubTribe |
Philonthina |
Belonuchus Nordmann 1837
Chani-Posse, Mariana, Newton, Alfred F., Hansen, Aslak Kappel & Solodovnikov, Alexey 2018 |
Belonuchus
Nordmann 1837 : 129 |
Musicoderus
Sharp 1885 : 455 |
Lucas 1920 : 425 |
Bernhauer 1916c: 266 |
Blackwelder 1943: 421 |
Philonthus abnormalis
Sharp 1885 : 397 |
Belonuchus aequalis
Sharp 1876a : 161 |
Belonuchus agilis
Erichson 1840 : 423 |
Belonuchus chevrolati
Fauvel 1863 : 432 |
Fauvel 1869: 489 |
Belonuchus albovariegatus
Bernhauer 1916c : 276 |
Linoderus navarretei López-García & Méndez-Rojas 2014 : 86
López-García & Méndez-Rojas 2014 : 86 |
Chani-Posse & Solodovnikov 2017: 90 |
Philonthus alternans
Sharp 1885 : 418 |
Navarrete-Heredia et al. 2002 : 334 |
Belonuchus alternatus
Sharp 1885 : 430 |
Bernhauer & Schubert (1914: 369) |
Sharp 1885 : 418 |
Belonuchus altissimus
Bernhauer 1916c : 275 |
Belonuchus altivagans
Bernhauer 1916c : 272 |
Philonthus aluticollis
Bernhauer 1921c : 107 |
Chani-Posse 2010: 8 |
Chani-Posse 2011b: 215 |
Belonuchus amplipennis
Fauvel 1891 : 117 |
Belonuchus amplus
Blackwelder 1943 : 425 |
Belonuchus andinus
Bernhauer 1916c : 278 |
Belonuchus (Musicoderus) angusticeps
Scheerpeltz 1960 : 117 |
Belonuchus angusticollis
Bernhauer 1906 : 330 |
Belonuchus antiguae
Blackwelder 1943 : 434 |
Belonuchus (Musicoderus) apicicornis
Bernhauer 1916c : 270 |
Philonthus apiciventris
Sharp 1885 : 419 |
Belonuchus argentinus
Bernhauer 1916c : 278 |
Bruch (1915: 503) |
Belonuchus armatus
Sharp 1876a : 163 |
Belonuchus arrogans
Bernhauer 1916c : 274 |
Belonuchus asperatus
Bernhauer 1906 : 336 |
Scheerpeltz 1960 : 120 |
Plociopterus atticida
Wasmann 1925 : 71 |
Chatzimanolis & Chani-Posse 2016: 214 |
Hesperus eidmanni
Scheerpeltz 1936 : 484 |
Posse 2016 : 214 |
Chatzimanolis & Chani-Posse 2016: 214 |
Belonuchus avidus
Erichson 1840 : 423 |
Herman 2001d : 1594 |
Belonuchus (Musicoderus) barbicornis
Bernhauer 1916c : 271 |
Philonthus apiciventris
Sharp 1885 : 418 |
Belonuchus batesi
Sharp 1876a : 157 |
Belonuchus bidens
Sharp 1885 : 430 |
Belonuchus bipunctatus
Bernhauer 1935 : 90 |
Belonuchus boops
Sharp 1885 : 435 |
Belonuchus boxi
Bernhauer 1934b : 212 |
Belonuchus brasilianus
Bernhauer 1917b : 19 |
Belonuchus breviceps
Fauvel 1891 : 117 |
Belonuchus bruchi
Bernhauer 1912b : 175 |
Belonuchus bruchianus
Bernhauer 1911 : 413 |
Belonuchus bugnioni
Fauvel 1901 : 86 |
Belonuchus eximius
Bernhauer 1917c : 105 |
Blackwelder 1943: 428 |
Belonuchus cacao
Blackwelder 1943 : 434 |
Philonthus candens
Erichson 1840 : 460 |
Bernhauer & Schubert 1914: 369 |
Belonuchus cariniventris
Bernhauer 1918 : 225 |
Belonuchus castaneus
Chani-Posse 2006 : 366 |
Bernhauer 1922 : 15 |
Belonuchus richteri
Bernhauer 1922 : 15 |
Bernhauer 1912a : 44 |
Chani-Posse 2011b : 219 |
Belonuchus (Musicoderus) cavipennis
Bernhauer 1916c : 267 |
Philonthus celatus
Sharp 1885 : 420 |
Musicoderus cephalotes
Sharp 1885 : 455 |
Scheerpeltz 1933 : 1377 |
Belonuchus cifuentesi Rodríguez & Navarrete-Heredia 2016 : 445
Rodríguez & Navarrete-Heredia 2016 : 445 |
Belonuchus clypeatus
Sharp 1876a : 160 |
Belonuchus coelestinus
Bernhauer 1908 : 330 |
Belonuchus coeruleus
Cameron 1922 : 122 |
Blackwelder 1943: 426 |
Belonuchus smithi
Cameron 1922 : 122 |
Blackwelder, 1943: 426 |
Belonuchus (Musicoderus) coelestis
Bernhauer 1916c : 266 |
Belonuchus coeruleispennis
Motschulsky 1858 : 668 |
Motschulsky 1870 : 49 |
Bernhauer & Schubert 1914 : 369 |
Herman (2001d : 2522 |
Belonuchus cognatus
Sharp 1885 : 429 |
Philonthus colon
Sharp 1885 : 419 |
Navarrete-Heredia et al. 2002 : 334 |
Belonuchus (Musicoderus) columbicus
Bernhauer 1916c : 271 |
Staphylinus connexus
Say 1830 : 35 |
Erichson 1840: 927 |
Musicoderus convexus
Bernhauer 1908 : 331 |
Scheerpeltz 1933 : 1377 |
Belonuchus (Musicoderus) cordilleranus
Bernhauer 1916c : 270 |
Philonthus corticalis
Sharp 1885 : 413 |
Belonuchus cribriceps
Bernhauer 1921c : 172 |
Belonuchus danforthi
Blackwelder 1943 : 429 |
Belonuchus decipiens
Sharp 1876a : 159 |
Belonuchus dichrous
Erichson 1840 : 420 |
Belonuchus diversicornis
Bernhauer 1917c : 104 |
Belonuchus docilis
Sharp 1885 : 435 |
Belonuchus dominicus
Blackwelder 1943 : 431 |
Belonuchus duplicatus
Blackwelder 1944 : 137 |
Belonuchus bruchi
Bernhauer 1934b : 213 |
Belonuchus (Musicoderus) ecuadorensis
Bernhauer 1935 : 91 |
Staphylinus ephippiatus
Say 1830 : 35 |
Erichson 1840: 927 |
Smetana 1995: 746 |
Belonuchus simplex
Sharp 1885 : 432 |
Fauvel 1901: 86 |
Smetana 1995: 748 |
Belonuchus punctiventris
Casey 1915 : 446 |
Smetana 1995: 746 |
Smetana 1995: 748 |
Belonuchus laticeps
Casey 1915 : 446 |
Smetana 1995: 746 |
Smetana 1995: 749 |
Belonuchus jacobianus
Casey 1915 : 447 |
Smetana 1995: 746 |
Smetana 1995: 749 |
Belonuchus schaefferi
Cooper 1933 : 545 |
Smetana 1995: 746 |
Belonuchus erichsoni
Bernhauer 1917d : 223 |
Belonuchus erythropterus
Solsky 1868 : 137 |
Smetana 1995: 752 |
Belonuchus excelsus
Bernhauer 1917b : 19 |
Philonthus extremus
Sharp 1885 : 415 |
Belonuchus falsus
Bernhauer 1917c : 105 |
Belonuchus fassli
Bernhauer 1917a : 37 |
Philonthus flavicoxis
Bernhauer 1912a : 43 |
Chani-Posse 2010: 8 |
Chani-Posse 2011b: 221 |
Belonuchus flavipennis
Solsky 1870 : 263 |
Belonuchus flavocaudatus
Bernhauer 1921c : 170 |
Belonuchus (Musicoderus) forsteri
Scheerpeltz 1960 : 115 |
Belonuchus fossulatus
Wasmann 1887 : 414 |
Belonuchus frater
Bernhauer 1906 : 333 |
Belonuchus gagates
Erichson 1840 : 424 |
Belonuchus gowdeyi
Bernhauer 1935 : 91 |
Smetana 1995: 739 |
Blackwelder 1943: 424 |
Belonuchus minax
Erichson 1840 : 424 |
Blackwelder 1943: 424 |
Smetana (1995: 737) |
Musicoderus nigrocaeruleus
Cameron 1913 : 175 |
Blackwelder 1943: 424 |
Smetana (1995: 738) |
Belonuchus ganglbaueri
Bernhauer 1906 : 331 |
Philonthus godmani
Sharp 1885 : 413 |
Plociopterus goeldii
Wasmann 1925 : 72 |
Chatzimanolis & Chani-Posse 2016: 215 |
Musicoderus gracilis
Sharp 1885 : 456 |
Staphylinus haemorrhoidalis
Fabricius 1801 : 596 |
Nordmann 1837: 130 |
Gmelin 1790 : 2036 |
Herman 2001a: 42 |
Belonuchus hispaniolus
Blackwelder 1943 : 422 |
Belonuchus holisoides
Sharp1876a : 161 |
Belonuchus iconnicoffi
Plavilstschikov 1929 : 30 |
Philonthus ielskii
Solsky 1872 : 296 |
Bernhauer & Schubert 1914: 370 |
Herman 2001d : 2527 |
Asenjo 2004 : 61 |
Belonuchus impressifrons
Sharp 1876a : 162 |
Philonthus infimus
Sharp 1885 : 414 |
Belonuchus iridescens
Solsky 1868 : 140 |
Belonuchus irregularis
Bernhauer 1916c : 275 |
Philonthus iteratus
Sharp 1887 : 788 |
Belonuchus jalappensis
Bernhauer 1917d : 224 |
Belonuchus laevipennis
Bernhauer 1942 : 25 |
Philonthus latecinctus
Sharp 1885 : 415 |
Belonuchus latro
Erichson 1840 : 422 |
Belonuchus longiceps
Bernhauer 1906 : 332 |
Philonthus lucilius
Sharp 1885 : 415 |
Belonuchus magnificus
Bernhauer 1916c : 277 |
Belonuchus meridionalis
Bernhauer 1916c : 269 |
Belonuchus mexicanus
Solsky 1864 : 440 |
Belonuchus mexicanus var. femoralis
Solsky 1864 : 440 |
Navarrete-Heredia et al. (2002 : 335 |
Belonuchus militaris
Bernhauer 1921c : 170 |
Belonuchus minor
Bruch 1915 : 504 |
Fernández et al. (2007 : 38 |
Belonuchus (Musicoderus) miripennis
Bernhauer 1916c : 267 |
Belonuchus modestus
Fauvel 1891 : 118 |
Belonuchus montanellus
Bernhauer 1916c : 272 |
Belonuchus montanicus
Blackwelder 1944 : 137 |
Belonuchus montanus
Bernhauer 1916c : 273 |
Fauvel 1895 : 267 |
Belonuchus monticola
Bernhauer 1917b : 18 |
Belonuchus mordens
Erichson 1840 : 422 |
Wasmann 1904 : 482 |
Belonuchus moritzi
Bernhauer 1906 : 331 |
Endeius multipunctatus
Coiffait 1981 : 297 |
Maus 2008 : 386 |
Chani-Posse 2011b: 222 |
Belonuchus mundus
Erichson 1840 : 425 |
Philonthus muticus
Sharp 1876a : 167 |
Belonuchus nigerrimus
Bernhauer 1918 : 225 |
Belonuchus oakleyi
Blackwelder 1943 : 422 |
Philonthus optatus
Sharp 1885 : 427 |
Philonthus oxyporinus
Sharp 1885 : 419 |
Belonuchus (Musicoderus) palustris
Bernhauer 1916c : 268 |
Belonuchus panamensis
Sharp 1885 : 434 |
Belonuchus paradoxipunctatus
Bernhauer 1921c : 171 |
Belonuchus paradoxus
Bernhauer 1906 : 333 |
Belonuchus pectinipes
Bernhauer 1910 : 379 |
Belonuchus peruvianus
Plavilstschikov 1929 : 30 |
Belonuchus piceolus
Bernhauer 1935 : 90 |
Belonuchus pictipennis
Sharp 1885 : 436 |
Belonuchus planiceps
Erichson 1840 : 425 |
Philonthus platypterus
Sharp 1885 : 412 |
Belonuchus pollens
Sharp 1885 : 431 |
Smetana 1995: 733 |
Staphylinus posticalis Laporte de Castelnau 1840 : 176
Castelnau 1840 : 176 |
Bernhauer & Schubert 1914: 371 |
Belonuchus prasinipennis
Bernhauer 1917b : 18 |
Belonuchus pulchripennis
Bernhauer 1908 : 330 |
Belonuchus purpuripennis
Bernhauer 1916c : 278 |
Philonthus (Chroaptomus) richteri
Bernhauer 1912a : 44 |
Chani-Posse 2006: 365 |
Chani-Posse 2011b: 223 |
Staphylinus rufipennis
Fabricius 1801 : 596 |
Erichson 1840: 426 |
Smetana 1995: 726 |
Staphylinus formosus
Gravenhorst 1806 : 72 |
Smetana 1995: 728 |
Belonuchus pallipes
Melsheimer 1844 : 35 |
Smetana 1995: 728 |
Cafius apicalis
Dejean 1833 : 63 |
Erichson 1840: 426 |
Philonthus rufiventris
Sharp 1887 : 787 |
Philonthus rufocaudus
Sharp 1885 : 412 |
Philonthus rufopygus
Sharp 1885 : 411 |
Belonuchus rugicollis
Bernhauer 1916c : 276 |
Belonuchus satyrus
Erichson 1840 : 421 |
Philonthus serraticornis
Sharp 1876a : 179 |
Belonuchus setiger
Sharp 1876a : 164 |
Belonuchus sharpi
Bernhauer 1907 : 285 |
Sharp 1876a : 158 |
Belonuchus grandiceps
Sharp 1876a : 158 |
Kraatz 1859a : 95 |
Belonuchus silvestris
Bernhauer 1917b : 17 |
Belonuchus similis
Sharp 1885 : 436 |
Belonuchus simillimus
Bernhauer 1917c : 106 |
Belonuchus skalitskyi
Bernhauer 1906 : 334 |
Musicoderus spinicornis
Champion 1913 : 4 |
Belonuchus stenoderus
Sharp 1885 : 434 |
Belonuchus (Musicoderus) subaeneipennis
Bernhauer 1916c : 269 |
Belonuchus subaeneus
Bernhauer 1916c : 268 |
Belonuchus subsellatus
Bernhauer 1916c : 276 |
Belonuchus superbus
Bernhauer 1916c : 273 |
Philonthus supernus
Herman 2001a : 48 |
Bernhauer 1941 : 287 |
Philonthus excelsus
Bernhauer 1941 : 287 |
Cameron 1931 : 361 |
Cameron 1932 : 147 |
Philonthus taczanovskii
Solsky 1872 : 303 |
Staphylinus terminalis Laporte de Castelnau 1840 : 176
Castelnau 1840 : 176 |
Bernhauer & Schubert 1914: 371 |
Erichson 1839 : 396 |
Belonuchus tremolerasi
Bernhauer 1912a : 40 |
Belonuchus trinitatis
Blackwelder 1943 : 423 |
Philonthus trochanterinus
Sharp 1885 : 417 |
Belonuchus tucumanensis
Bernhauer 1934b : 213 |
Belonuchus variolosus
Solsky 1870 : 263 |
Belonuchus variolaris
Solsky 1872 : 305 |
Belonuchus venezolanus
Bernhauer 1917c : 104 |
Belonuchus viridipennis Baudi di Selve 1848 : 125
Selve 1848 : 125 |
Herman 2001c : 1594 |
Belonuchus dejectus
Sharp 1885 : 433 |
Sharp 1887: 789 |
Belonuchus vulneratus
Bernhauer 1906 : 335 |
Philonthus weiserianus
Bernhauer 1921c : 106 |
Chani-Posse 2010: 8 |
Chani-Posse 2011b: 224 |
Belonuchus xantholinoides
Bernhauer 1917c : 106 |
Belonuchus xanthomelas
Solsky 1868 : 139 |
Philonthus xanthopterus
Nordmann 1837 : 99 |
Erichson 1840: 421 |
Soukup 1942: 177 |
Belonuchus xanthopus
Solsky 1868 : 139 |
Philonthus zunilensis
Sharp 1885 : 418 |