Palleptoceridae, WICHARD & MÜLLER, 2022

WICHARD, WILFRIED & MÜLLER, PATRICK, 2022, Palleptoceridae fam. nov., an extinct leptoceroid family in mid-Cretaceous Burmese amber (Insecta, Trichoptera), Palaeoentomology 5 (5), pp. 468-474 : 469

Palleptoceridae fam. nov.

Type genus. Palleptocerus gen. nov.

Diagnosis. Ocelli absent. Antennae longer than forewings, scapus conically thickened. Maxillary palps five-segmented, lengths of palp segments in a row: short (1), long (2), long (3) short (4) and longest (5); terminal segment not annulated ( Figs 2A , 3B inset).

Wings: Forewings slightly narrow and apically rounded, hind wings shorter and broader. Sexual dimorphism in terms of wing length and wing venation: Forewing lengths in males a little shorter than in females. The light brown wings of the males are darker in the females. In males forewing venation with forks I and V present ( Fig. 2B ); in females forks I, III, and V present ( Fig. 2C ). In both sexes, discoidal cells open and thyridial cells closed, long. The hind wing venation reduced in both sexes; forks I, II, III, and IV absent, exclusively fork V present; discoidal cells open; crossveins not present ( Fig. 2D ).

Tibial spurs: 2/4/4, metatibia, each with a pair of apical and preapical spurs ( Fig. 1B )

Palleptoceridae fam. nov. is characterized by the apomorphic combination of the following characters: Tibial spur formula 2/4/4. Forewing fork I present without discoidal cell and in hind wing exclusively fork V present.

Figures

FIGURE 1. Palleptocerus grimaldii sp. nov., mating. A, A mating pair, male holotype (right), female paratype 1 (left), ZFMK- TRI000831, in ventral view; scale bar = 2 mm. B, Male paratype, ZFMK-TRI000833, metatibia, each with a pair of apical and preapical spurs (arrows), circle = branching spurs; scale bar = 0.5 mm.

FIGURE 2. Palleptocerus grimaldii sp. nov., characters. A, Maxillary palp of female paratype 1, ZFMK-TRI000831; scale bar = 0.1 mm. B, Wing venations, venations of male forewing, female forewing and hind wing; scale bar = 1 mm.

FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm