Shearia khakassica Mikhaljova, 2000

Nefediev, P. S., 2019, New data on the millipede genus Shearia Mikhaljova, 2000 from the Russian and Mongolian Altais (Diplopoda: Chordeumatida: Diplomaragnidae), Russian Entomological Journal 28 (2), pp. 219-225 : 221-222

Shearia khakassica Mikhaljova, 2000

Figs 14–20 .

Shearia khakassica Mikhaljova, 2000: 163 , figs.

Shearia khakassica — Mikhaljova,Golovatch,2001:110; Mikhaljova, 2004: 166, 167: figs, 164: map; 2016: 13; 2017: 202, 203: figs, 140: map; Nefediev, Nefedieva, 2006: 98; 2007: 161; 2008b: 62.

Shearia oiskaya — Mikhaljova et al., 2008: 54 .

NEW MATERIAL EXAMINED. 1 ♂, 7 ♀♀, 7 juv., Russia, Republic of Altai, Ulagan District, 9 km S of Lake Dzhulukul, near Lak-Yyash Lakes , Betula rotundifolia thicket, 50.393970°N, 89.674179°E, 2200 m a.s.l., 27.VII.2006, leg. P.S. Nefediev, J.S. Nefedieva.

MATERIAL RE-EXAMINED (specimens previously identified as Shearia oiskaya Mikhaljova, 2000 and published by Mikhaljova et al. [2008]). 7 ♂♂, 2 ♀♀, 22 juv., Russia, Republic of Altai, Ulagan District, 9 km S of Lake Dzhulukul, near Lak-Yyash Lakes, Betula rotundifolia thicket, 50.393970°N, 89.674179°E, 2200 m a.s.l., 27.VII.2006, leg. P.S. Nefediev, J.S. Nefedieva.

DISTRIBUTION. This species is rather widespread in the south of western and central Siberia (W districts of the Republic of Khakassia, S districts of the Kemerovo Area and the Krasnoyarsk Province, as well as SE districts of the Republic of Altai) ( Fig. 14 ) .

REMARKS. A restudy of specimens from near Lak-Yyash Lakes at the border of the Republic of Altai and the Republic of Tyva, previously identified as Shearia oiskaya Mikhaljova, 2000 by Mikhaljova et al. [2008], shows that all of them actually belong to S. khakassica . The above southernmost records of the species are new to the Republic of Altai. Neither the gonopods nor coxae 10, nor coxae 11 of this species have ever been shown using SEM. Hence, they

Figs 8–13. Shearia densecava , male: 8–11 — gonopods; 12 — coxae 10; 13 — coxae 11; 8, 10, 12–13 — oral view; 9, 11 — caudal view (8, 10, 12–13 — Republic of Altai, Lake Taldu-Kiol; 9, 11 — Republic of Altai, Kalbakaya River). Scale bars: 1 mm (8–11), 0.4 mm (12) and 0.3 mm (13).

Рис. 8–13. Shearia densecava , самец: 8–11 — гоноподы; 12 — коксы 10-й пары ног; 13 — коксы 11-й пары ног; 8, 10, 12–13 — спереди; 9, 11 — сЗади (8, 10, 12, 13 — Республика Алтай, оЗ. Талду- Кёль; 9, 11 — Республика Алтай, р. КалбакаЯ). МасШтаб: 1 мм (8–11), 0,4 мм (12) и 0,3 мм (13). are given here ( Figs 15–20 ). An examination of males of the species from the ASU collection has revealed a small posterior process (pp) on each angiocoxite of the posterior gonopods, as opposed to “absent” or “a small fold”, specified in the original description [see Mikhaljova, 2000]. This structure is well visible in a SEM micrograph ( Fig. 18 ), and it must be included into the description of S. khakassica .

Figures

Fig. 14. Distribution of Shearia khakassica (circle) and S. longa (square). Previously known localities marked in black, new records given in white. Рис. 14. Распространение Shearia khakassica (круг) и S. longa (квадрат). Чёрным отмечены ранее иЗвестные места находок, новые находки отмечены белым.

Figs 15–20. Shearia khakassica, male (Republic of Altai, near Lak-Yyash Lakes): 15–18 — gonopods; 19 — coxae 10; 20 — coxae 11; 15, 17, 19–20 — oral view; 16, 18 — caudal view. Scale bars: 1 mm (15–18) and 0.4 mm (19–20). Designations explained in text. Рис. 15–20. Shearia khakassica, самец: 15–18 — гоноподы; 19 — коксы 10-й пары ног; 20 — коксы 11-й пары ног; 15, 17, 19– 20 — спереди; 16, 18 — сЗади. МасШтаб: 1 мм (15–18) и 0,4 мм (19–20). ОбЪЯснение обоЗначений дано в тексте.