Enoclerus mcnallyi Rifkind

Rifkind, Jacques, 2012, New Central American and Mexican Enoclerus Gahan (Coleoptera: Cleridae: Clerinae): Part II, Zootaxa 3397, pp. 1-27 : 5-6

Enoclerus mcnallyi Rifkind , n. sp.

( Figs 4 , 21 )

Type specimen. Holotype male: Mex [ico], Gro. [= Guerrero], 18–20 km SW Filo del Caballo, 9000', Sept. 17, 1989, E. Giesbert, coll. Holotype deposited in CSCA.

Diagnosis. Its distinct pronotal and elytral color pattern will readily separate this attractive species from congeners. It appears closest to E. varius (Klug) , which is known from the Atlantic versant of Mexico (Chipinque Mesa, Nuevo Léon) and Querétaro. They differ in details of coloration and elytral patterning. For example, E. mcnallyi has its vertex, occiput and pronotal posterior darkened, its elytral subbasal markings macular (rather than produced into a fascia), and its posterior elytral fascia transverse rather than sinuate. Lastly, in E. mcnallyi , the pronotum appears proportionally narrower across the disk than in E. varius .

Description (Holotype). Length: 8.00 mm. Color: piceous; frons, ocular notch, labial palpi, pronotal arch (anterior of pronotum before transverse impression), two smallish irregular spots on each side of pronotum, pronotal collar, and as follows on each elytron: a large rounded basal macula dorsally, a large triangulate macula below humerus laterally (narrowly separated from basal macula, complete to sides and to anterior elytral margin ( Fig. 21 )), a rather broad, transverse midelytral fascia, irregularly margined, complete to sides (rounded internally, interrupted before suture), a somewhat narrower, slightly oblique fascia at posterior 1/5 (interrupted laterally at epipleuron, nearly complete internally at suture), a small transverse anteapical spot at apical angle, all yellow ochre; middle disk of pronotum, scutellum, meso– and metasterna (in part), middle anterior of elytra, elytral sutural area (broadly until elytral posterior 1/5, then narrowly to apices), reddish; clypeus (in part), mandibles (in part), maxillary palpi, antennae, meso– and metasterna (in part), abdomen and legs, orange testaceous. Head: antennae elongate, club rather loose, antennomere 11 acute at apex, more than 2X as long as antennomere 10; ocular notch broad; gular process elongate–spatulate; surface shallowly, finely, rather sparsely punctulate, sparsely set with mostly short, mostly suberect, fine, pale setae. Pronotum: as broad as long, narrower than elytra at base; transverse impression deeply incised, sinuate laterally, broadly U–shaped at middle; sides moderately convex; disk notably flattened above; surface shining, very finely, shallowly punctulate, quite sparsely vested with pale, fine, suberect and erect setae of mostly medium length, intermixed with a few darker, longer, erect setae. Elytra: elongate (ratio of length to width 60:33). widest at posterior 1/3, rather shallow in cross section; humeri pronounced, with subcarinate umbones; subbasal tumescences feebly elevated, gently rounded above; disk flattened at middle; surface coarsely, shallowly rugulose–punctate, with additional small defined punctures scattered irregularly; vestiture inconspicuous, consisting of a moderately dense covering of rather short, fine, suberect and erect pale and dark setae, and a few longer, erect setae; sides subparallel, apices separately rounded and apparently rather deeply dehiscent; apical slope moderately shallow. Mesosternum: posterior median process abruptly elevated at base. Abdomen: shining, very sparsely, finely setose; visible sternite 5 broadly, arcuately emarginate, with posterior angles rounded; visible sternite 6 with posterior margin apparently subtruncate; visible tergite 6 convex dorsally (concave ventrally), with sides slightly oblique, hind angles rounded and posterior margin broadly, feebly emarginate at middle, surpassing sternite 6. Legs: robust.

Etymology. I take particular pleasure in naming this rather dapper looking Enoclerus species for Brien McNally: bon vivant, oenophile and friend.

Distribution. Known only from the type locality.

Relationships. The new species appears to belong to a lineage of montane Mexican and Central American species that include the aforementioned E. varius , as well as E. maculicollis (Spinola) , E. whiteheadi Ekis , E. villicus (Gorham) , E. absconditus Rifkind and the three taxa described directly below. They share a similar facies, rather loosely composed antennal club, a broad ocular insertion, and shining, finely punctulate or shallowly roughened and sparsely setose elytra, the latter often marked with three or four sets of pale transverse bands and/or maculae. Except for the Central American species, they are very rarely collected, and no doubt other members of the group await discovery in the mountain forests of México.

Figures

FIGURES 1 – 9. Habitus of: 1. Enoclerus anctus Rifkind n. sp. (holotype female (? )). 2. Enoclerus boquete Rifkind n. sp. (holotype male). 3. Enoclerus skillmani Rifkind n. sp. (holotype male). 4. Enoclerus mcnallyi Rifkind n. sp. (holotype male). 5. Enoclerus fibrillatus Rifkind n. sp. (holotype male). 6. Enoclerus toledoi Rifkind n. sp. (holotype male). 7. Enoclerus boblloydi Rifkind n. sp. (holotype female). 8. Enoclerus gilli Rifkind n. sp. (holotype male). 9. Enoclerus chamelae Rifkind n. sp. (holotype male).

FIGURES 19 – 22. Habitus of: 19. Enoclerus silbermannii aeternitatis Rifkind, n. subsp. (holotype male). 20. Enoclerus albidulus Rifkind n. sp. (holotype male). 21. Enoclerus mcnallyi Rifkind n. sp., detail of anterior elytral markings. 22. Enoclerus toledoi Rifkind n. sp., lateral view of pronotum showing distinctive markings.