Phanaeus haroldi Kirsch, 1871

Rosa, Cecilia Lozano De La, Cupello, Mario & Vaz-De-Mello, Fernando Z., 2024, The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution, European Journal of Taxonomy 959 (1), pp. 1-272 : 176-178

Phanaeus haroldi Kirsch, 1871

Fig. 66B

Phanaeus haroldi Kirsch, 1871: 342 (original description). Type locality: Colombia: Cundinamarca: Bogotá. Name-bearing type: holotype (SMTD), not examined.

Phanaeus schneblei Frey, 1963: 558 (original description). Type locality: Colombia: Cundinamarca: Guayabetal: Monterredondo. Name-bearing type: holotype (NHMB ‒ ex Frey collection), not examined.

Phanaeus haroldi – Kirsch 1871: 342 (358) (cited for Bogotá). — Nevinson 1892: 4 (catalogue). — Gillet 1911b: 83 (catalogue). — d’Olsoufieff 1924: 34, 85, 147 (key, redescription, distribution). — Blackwelder 1944: 209 (checklist). — Vulcano & Pereira 1967: 575 (key). — Amézquita et al. 1999: 119 (biodiversity). — Escobar 2000: 209 (list). — Vítolo 2000: 597 (key); 2004: 283 (diagnosis). — Medina et al. 2001: 140 (checklist for Colombia). — Noriega et al. 2009a: 406 (distribution). — Krajcik 2012: 204 (checklist). — Ratcliffe et al. 2015: 197 (checklist for Peru).

Phanaeus schneblei – Frey 1967: 405 (synonymy with P. haroldi ). — Arnaud 2002b: 82 (cited as synonym of P. haroldi ).

Phanaeus (Notiophanaeus) haroldi – Edmonds 1994: 21, 24, 101 (key, redescription, distribution). — Vaz-de-Mello 2000: 194 (cited for Brazil). — Arnaud 2002b: 82 (key, diagnosis). — Carvajal et al. 2011: 320–321 (cited for Ecuador). — Edmonds & Zídek 2012: 12 (key). — Figueroa et al. 2014: 133 (distribution for Peru). — Chamorro et al. 2018: 98 (cited for Ecuador); 2019: 217 (catalogue). — Hielkema & Hielkema 2019: 106 (catalogue for the Guiana).

Distribution

Colombia, Venezuela, Brazil, Ecuador, and Peru.

Subregions of Venezuela

Peneplain of the Caura and Paragua rivers, Peneplain of the Casiquiare River –Upper Orinoco, System of hills and low piedmont mountains of the Guiana Shield, Andes mountains, Central Coast Mountain Range, and Guiana Shield.

Literature records

Blanco 1988: 45 ( Venezuela: Táchira). — Martínez & Clavijo 1990: 7 ( Venezuela: Territorio Federal de Amazonas). — Edmonds 1994: 21, 24, 101 ( Venezuela: Amazonas, Bolívar and Táchira). — Arnaud 2002b: 82 ( Venezuela: Amazonas, Aragua, Táchira and Bolívar). — Gámez 2004: 48, 53, 57, 59 ( Venezuela: Cordillera de Los Andes, Táchira). — Vítolo 2004: 283 ( Venezuela: Edmonds 1994). — Noriega et al. 2009: 406 ( Venezuela: Amazonas, Aragua, Bolívar and Táchira states). — Edmonds & Zídek 2012: 12 ( Venezuela). — Gámez & Acconcia 2018: 65, 73 ( Venezuela: Barinas state: Bolívar). — Chamorro et al. 2019: 217 ( Venezuela).

Figures

Fig. 66. Species distribution. A. Phanaeus bordoni Arnaud, 1997. B. Phanaeus haroldi Kirsch, 1871. Green diamond = CEMT collection data; black circle = literature data.