Phanaeus bispinus Bates, 1868

Rosa, Cecilia Lozano De La, Cupello, Mario & Vaz-De-Mello, Fernando Z., 2024, The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution, European Journal of Taxonomy 959 (1), pp. 1-272 : 175-176

Phanaeus bispinus Bates, 1868

Fig. 65B

Phanaeus bispinus Bates, 1868: 89 (original description). Type locality: Ecuador: Pastaza: Pastaza: Canelos. Name-bearing type: holotype by monotypy (MNHN), not examined.

Phanaeus digitalis d’Olsoufieff, 1924: 34 , 84, 147 (original description). Type locality: Guyana. Name-bearing type: holotype by monotypy (MNHN), not examined.

Phanaeus bispinus – Harold 1869d: 1017 (catalog, Phanaeus bispinns [sic]). — Nevinson 1892: 2 (catalog, distribution). — Gillet 1911b: 81 (catalog, distribution). — d’Olsoufieff 1924: 34, 84, 147 (key, redescription, distribution). — Blackwelder 1944: 209 (checklist). — Frey 1963: 559 (comments). — Vulcano & Pereira 1967: 574 (key). — Arnaud 1982: 114 (list of the MNHN types). — Escobar 2000: 209 (list). — Feer 2000: 32 (list for French Guiana); 2013: 767 (list for French Guiana). — Vítolo 2000: 597 (key); 2004: 283 (diagnosis, distribution). — Medina et al. 2001: 140 (checklist). — Pulido-Herrera et al. 2003: 54 (list for Caqueta, Colombia). — Gámez 2004: 57 (list). — Hamel-Leigue et al. 2006: 17 (inventory for Bolivia); 2006: 66 (distribution). — Brûlé et al. 2011a: 193 (list); 2014: 183 (list). — Larsen 2011: 99 (list for Suriname); 2013: 97 (list for Suriname). — Krajcik 2012: 204 (checklist). — Brûlé & Touroult 2013: 41 (list). — Figueroa et al. 2014: 133 (distribution for Peru). — Nunes et al. 2014: 408–410 (cited, list). — Ratcliffe et al. 2015: 197 (list for Peru). — Silva et al. 2016: 555 (ecology, biogeography, list).

Phanaeus (Phanaeus) bispinus – Pereira & Martínez 1956b: 237 (notes).

Phanaeus (Notiophanaeus) bispinus – Edmonds 1994: 33, 35, 100 (key, redescription, distribution). — Vaz-de-Mello 2000: 194 (checklist for Brazil). — Arnaud 2002b: 89 (key, diagnosis). — Vítolo 2004: 283 (diagnosis). — Carvajal et al. 2011: 320–321 (list, cited for Ecuador). — Edmonds & Zídek 2012: 5, 9 (list, key). — Pacheco & Vaz-de-Mello 2015: 2 (checklist). — Boilly et al. 2016: 90, 93, 96 (list for French Guiana, key). — Chamorro et al. 2018: 97 (cited for Ecuador); 2019: 213 (catalogue). — Hielkema & Hielkema 2019: 105 (catalogue for the Guiana).

Phanaeus (Phanaeus) digitalis – Pereira & Martínez 1956b: 237 (cited as synonym of P. bispinus ).

Distribution

Colombia, Venezuela, Guyana, French Guiana, Brazil, Ecuador, Peru, and Bolivia.

Subregion of Venezuela

Peneplain of the Casiquiare River –Upper Orinoco.

Literature records

Edmonds 1994: 35, 100 ( Venezuela: Amazonas). — Arnaud 2002b: 89 ( Venezuela: Amazonas). — Gámez 2004: 57 ( Venezuela). — Vítolo 2004: 283 ( Venezuela: Edmonds 1994). — Hamel-Leigue et al. 2009: 66 ( Venezuela). — Chamorro et al. 2019: 213 ( Venezuela).

Figures

Fig. 65. Species distribution. A. Oxysternon spiniferum Castelnau, 1840. B. Phanaeus bispinus Bates, 1868. Green diamond = CEMT collection data; black circle = literature data.