Cicindela (Cicindela) sachalinensis niohozana Bates, 1883

Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, Zootaxa 4661 (2), pp. 271-308 : 301

Cicindela (Cicindela) sachalinensis niohozana Bates, 1883

( Figs 8H , 15C )

Cicindela sachalinensis niohozana Bates, 1883:213 .

Cicindela sachalinensis humeralis Beuthin, 1905:186 [synonymy by Horn 1926:211].

Cicindela (Cicindela) sachalinensis Morawitz, 1862 : Rivalier 1950:225 (only nominotypical subspecies).

Specimens examined. (N = 1) [ Japan] 1♂ ( EUMJ), Hokkaido, Mt. Rausu ~ Iwaobetsu , 23–25. VII. 1970, S. Kinoshita leg.

Description. Internal sac everted dors-apically in lateral view. DA, VA, B and BLR well large, followed in order by VLL, DLL and VB.

DLL roundly swollen, vested with fine spines. DA well swollen basally. VA triangular swollen, everted right laterally, vested numerously with short and weakly chitinous spines. BLR oval swollen left ventrally, vested numerously with well sclerotized spines. VB roundly and short swollen, vested with fine spines. VLL roundly and short swollen, everted right laterally, vested numerously with well sclerotized spines. B roundly swollen, everted right laterally, vested numerously with well sclerotized spines. Flagellum well short, curved. Medial tooth short, extending right laterally, widest at middle, thence gently tapered apically.

Remarks. This species shares the characteristics of the position of each bladder with C. (C.) transbaicalica japanensis but is distinguished from the latter by the following characteristics: VA triangular swollen, everted right laterally; medial tooth extending right laterally, widest at middle, thence gently tapered apically.

Figures

FIGURE 8. Habitus of Cosmodela, Cicindela (Cicindela) and Cylindera.A, Cosmodela sp. (cf. separata); B, C. batesi (Fleutiaux) from Iriomote Is., Japan; C, C. aurulenta (Fabricius) from Penang Is., Malaysia; D, C. virgula (Fleutiaux) from Laos; E, C. duponti duponti (Dejean) from South India; F, Cicindela (Cicindela) japana japana Motschulsky; G, C. (C.) gemmata aino Lewis; H, C. (C.) sachaliensis niohozana Bates; I, C. (C.) lewisii lewisii Bates; J, Cylindera (Eugrapha) elisae elisae (Motschulsky); K, Cicindela (Cicindela) transbaicalica japanensis Chaudoir. Scales: 10 mm.

FIGURE 15. Everted internal sacs of Cicindela (Cicindela). A, Cicindela (Cicindela) japana japana Motschulsky; B, C. (C.) transbaicalica japanensis Chaudoir; C, (C.) sachalinensis niohozana Bates; D, C. (C.) lewisii lewisii Bates; E, C. (C.) gemmata aino Lewis. Scales: 1.0 mm.