Jornandes cruralis Distant

Schaffner, JC & Schwartz, MD, 2008, Revision Of The Mexican Genera Ficinus Distant And Jornandes Distant With The Description Of 21 New Species (Heteroptera: Miridae: Orthotylinae: Orthotylini), Bulletin of the American Museum of Natural History 2008 (309), pp. 1-87: 38-45

publication ID

0003-0090

persistent identifier

http://treatment.plazi.org/id/1C1087D2-6779-7A03-423F-99E3FDDBD025

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Felipe

scientific name

Jornandes cruralis Distant
status

 

Jornandes cruralis Distant   Figures 2, 10C, 17–18, 19A

Jornandes cruralis Distant, 1893: 448   , pl. 39, fig. 11 (orig. desc.); Carvalho, 1958: 80 (catalog); Carvalho and Dolling, 1976: 795 (disc. of holotype); Carvalho, 1981: 4 (note); Schuh, 1995: 127 (catalog).

DIAGNOSIS: Recognized by relatively large size; head yellowish brown usually with vertex fuscous, maxillary plate and apex of clypeus dark fuscous to black, antenna and legs pale yellowish brown; pronotum and hemelytron dark fuscous to black (fig. 2); glabrous with only a few scattered setae dorsally; antenna and legs pale; labium nearly reaching posterior margin of mesosternum; cuticular sculpturing extending onto pronotum laterally (fig. 17B); length of antennal segment I less than vertex width, antennal segment II less than pronotal width; metepisternum dorsal to evaporative area of scent gland with microtrichia (fig. 17C). Jornandes cruralis   is similar to J. genetivus   ; nonetheless, these species are easily separated from one another. Jornandes cruralis   has pale antennae and the cuticular sculpturing is extended onto the pronotum, whereas in J. genetivus   the antenna is dark fuscous and the cuticular sculpturing of the hemelytron does not extend onto the pronotum. Additionally the male genitalia are diagnostic for each species. The pale yellowish-brown coloration will distinguish J. sinaloa   from the other two species, all of which share a similar type of male genitalia.

REDESCRIPTION: Male: COLORATION: Head shining yellowish brown, usually with a pale fuscous area on vertex; fuscous to black band extending from maxillary plate across apex of clypeus, and around to other eye, sometimes infuscate on sulcate portion of vertex, stripe on frons between antennal fossa and vertex infuscate; antenna pale yellowish brown with base of segment I fuscous; labium yellow, apex dark fuscous. Pronotum, scutellum and mesoscutum shining dark fuscous to black. Hemelytron with corium shining dark fuscous to black; membrane fuscous with narrow pale area immediately along cuneus. Underside of thorax shining dark fuscous or black except for posterior margin of scent gland evaporatorium, which is yellowish brown; underside of abdomen shining dark fuscous to black. Legs yellowish brown (same color as antenna and posterior area of evaporatorium) with apical tarsomeres pale fuscous. VESTITURE: Body as seen from above shining, almost devoid of setae. Head with sparsely distributed, short setae; antennal segment I with a few scattered setae, remaining segments covered with short decumbent setae. Pronotum glabrous, setae commonly decumbent and located along lateral margins; meso- and metapleurae with scattered erect setae; hemelytron with short decumbent setae located along costal margin and on or near cuneus; underside of abdomen with moderately sparsely distributed semierect setae. GENITALIA (fig. 18): Genital segment with broad, crestlike, serrate tergal process projecting from right of midline on dorsal margin of aperture; ventroposterior margin of capsule with broad notch; distal width of subgenital plate wide, strongly projecting dorsal to aperture of capsule. Left paramere C-shaped in dorsal view; with broad sensory lobe; diameter of paramere gradually expanded to subapical constriction, then expanded to wide, deeply notched mittenlike apical region. Right paramere elongate, slightly more than twice length of left paramere; diameter of paramere widest basally and subapically, distal region gradually tapered to dorsally projecting apex; distal region serrate and with several strong spines. Phallotheca elongate, conical, 1/3 length of spiculum; aperture large, widely open from dorsal surface to apex. Vesica with long, medially

00110992), Santa Lucia, Sinaloa (AMNH_PBI 00111001); scale 5 0.20 mm.

thickened spiculum; apical region of spiculum bifurcate with recurved, narrow, apically pointed, strongly serrate branches; base of spiculum sinuate, broadly affixed to dorsal margin of ductus seminis, without basal spine.

Female: Coloration and vestiture same as for male, except for more ovate body. GENITALIA with ventral margin of VLP not produced ventrally into VUL; SR moderately large, compressed along longitudinal axis; medial region of DLP moderately produced medially; posterior wall with wellsclerotized, convex medial section; IRL deeply incised on ventral margin (fig. 19A), with dorsal portion shorter than narrow, pointed ventral portion; otherwise as in generic description.

MEASUREMENTS: Male: (n 5 13; average given first followed in parentheses by range): Length, 4.28 (3.85–5.00); width, 1.77 (1.62–2.00). Head length, 0.28 (0.24– 0.34); width, 0.94 (0.89–1.02); vertex width, 0.43 (0.41–0.46). Length of antennal segment I, 0.36 (0.32–0.40); II, 1.40 (1.32–1.50); III, 1.10 (1.00–1.21); IV, 0.44 (0.36–0.51). Pronotal length, 0.88 (0.84–0.92) width across base, 1.46 (1.36–1.62). Cuneal length, 0.71 (0.65– 0.84); width across base, 0.63 (0.60–0.68).

Female: (n 5 13; those of holotype given first followed in parentheses by average and range). Length, 4.22 (4.17, 3.80–4.36); width, 1.92 (1.89, 1.76–2.08). Head length, 0.30 (0.31, 0.26–0.36); width, 0.94 (0.92 and 0.96); vertex width, 0.48 (0.48, 0.45–0.50). Length of antennal segment I, 0.30 (0.34, 0.30–0.38); II, 1.30 (1.33, 1.22–1.50); III missing on holotype (0.99, 0.91–1.13); IV, 0.40, 0.36–0.46. Pronotal length, 0.88 (0.88, 0.84–0.96); width across base, 1.46 (1.45, 1.36–1.61). Cuneal length, 0.64 (0.64, 0.58– 0.71); width across base, 0.60 (0.60, 0.58– 0.62).

DISCUSSION: The single specimen from near La Laguna, Guerrero, differs somewhat in head color from all others. The center of the vertex and frons is dark brown with areas along the eyes in the more typical yellowish brown.

The male genitalia of the Guerrero specimens differs in the following characteristics from those from Sinaloa: tergal process deflected into genital aperture and slightly more serrate, left paramere longer, and vesical spiculum with longer more serrate apical branches. We are treating the Guerrero and Sinaloa specimens as conspecific pending additional material from intervening localities, which may clarify the variation we have encountered in the male genitalia.

Jornandes cruralis   shares a number of significant characters with J. genetivus   . Both species are almost devoid of setae as seen from above, including the costal margin of the hemelytron. The cuticular sculpturing is more easily seen than in most species of the genus. Both species are essentially the same size, and the color of J. cruralis   is similar to that of males of J. genetivus   that have the dark fuscous to black pronotum.

The male genitalia of both of these species and those of the pale J. sinaloa   indicate a close relationship. The IRL of J. cruralis   and J. sinaloa   are similar in that both have the ventral margin deeply incised (fig. 19A), whereas in J. genetivus   the ventral margin is practically entire (fig. 19B). Both J. cruralis   and J. sinaloa   are distributed predominately on the western coastal plain or ‘‘buried ranges’’ of Mexico, with J. sinaloa   in the

northernmost part in Sinaloa and Sonora and J. cruralis   in south of Sinaloa. The southern populations of J. cruralis   in Guerrero and Morelos seem to overlap some of the distribution of J. genetivus   . These species were found together at one poorly documented collecting event in the environs of Chilpancingo. Whether they were contemporaneously in similar habitat or at equal elevation remains to be determined.

HOST PLANT: Ipomoea murucoides Roem and Schult   ( Convolvulaceae   ).

DISTRIBUTION: Known from 12 localities in Durango, Guerrero, Morelos, and Sinaloa (fig. 10C).

SPECIMENS EXAMINED: MEXICO: Durango: El Salto, 23.79522 ° N 105.36357 ° W, 2578 m, 19 Sep 1976, J. Carayon, 1♀ ( AMNH _ PBI 00243972 View Materials ) ( MNHN). Guerrero: 1 mi NE of La Laguna, 19.91028 ° N 101.75586 ° W, 17 Jul 1984, Carroll , Schaffner , Friedlander , 13 ( AMNH _ PBI 00106416 View Materials ) ( TAMU). 10.3 mi S of Iguala, 17.70023 ° N 100.36666 ° W, 23 Jul 1981, Bogar , Schaffner and Friedlander , Ipomoea murucoides Roem and Schult   ( Convolvulaceae   ), 13 ( AMNH _ PBI 00094261 View Materials ) ( CNC). Ipomoea murucoides Roem and Schult   ( Convolvulaceae   ), 2♀ ( AMNH _ PBI 00106417 View Materials , AMNH _ PBI 00106418 View Materials ) Ipomoea murucoides Roem and Schult.   ( Convolvulaceae   ), 13 ( AMNH _ PBI 00119112 View Materials ) ( TAMU). 22 mi N of Chilpancingo, 17.96733 ° N 99.58776 ° W, 643 m, 24 Aug 1958, H. F. Howden, 13 ( AMNH _ PBI 00110992 View Materials ) ( CNC). 6.2 mi SW of Xochipala, 1728 m, 17.75291 ° N 99.68363 ° W, 13 Jul 1985, Jones and Schaffner , 1♀ ( AMNH _ PBI 00094262 View Materials ) ( CNC). ‘‘ Chilpancingo , Guerrero. [17.55 ° N 99.5 ° W, 1402 m], 4600 ft., Aug., H. H. Smith’’, Holotype, 1♀ ( AMNH _ PBI 00085376 View Materials ) ( BMNH); 24 Jul 1961, R. and K. Dreisbach, 2♀ ( AMNH _ PBI 00133592 View Materials , AMNH _ PBI 00134074 View Materials ) ( USNM). Morelos: 10 km NE of Yautepec, 18.94464 ° N 99.0035 ° W, 1439 m, 13 Sep 1983, H. Brailovsky, 3♀ ( AMNH _ PBI 00245166 View Materials – AMNH _ PBI 00245168 View Materials ) ( UNAM). Yautepec , 18.88333 ° N 99.06667 ° W, 1216 m, 13 Jul 1963, F. D. Parker and L. A. Stange, 13 ( AMNH _ PBI 00245170 View Materials ) ( UCB). Sinaloa: 13 mi E of Concordia, 23.28319 ° N 105.861 ° W, 244 m, 05 Aug 1964, L. A. Kelton, 2♀ ( AMNH _ PBI 00112248 View Materials , AMNH _ PBI 00112249 View Materials ) ( CNC); 09 Aug 1964, L. A. Kelton, 2♀ ( AMNH _ PBI 00112244 View Materials , AMNH _ GoogleMaps  

PBI 00112245) ( CNC). 27 mi E of Villa Union , 23.35201 ° N 106.03997 ° W, 276 m, 26 Jul 1964, H. F. Howden, 2♀ ( AMNH _ PBI 00112246 View Materials , AMNH _ PBI 00112247 View Materials ) ( CNC) GoogleMaps   ; 26 Jul 1964, J. Powell, 13 ( AMNH _ PBI 00119228 View Materials ) ( UCB). Mazatlan , 23.21666 ° N 106.41666 ° W, 20 Sep 1976, J. Carayon, 13 ( AMNH _ PBI 00243968 View Materials ), 2♀ ( AMNH _ PBI 00243970 View Materials , AMNH _ PBI 00243971 View Materials ), 1♀ ( AMNH _ PBI 00243969 View Materials ) ( MNHN). Santa Lucia, 23.49755 ° N 105.92295 ° W, 1219 m, 04 Aug 1964, L. A. Kelton, 103 ( AMNH _ PBI 00111001 View Materials , AMNH _ PBI 00112216 View Materials – AMNH _ PBI 00112224 View Materials ), 19♀ ( AMNH _ PBI 00112225 View Materials – AMNH _ PBI 00112243 View Materials ) ( CNC) GoogleMaps   .

Jornandes genetivus (Distant)   , new combination Figures 2, 10C, 19B, 20–21

Eccritotarsus genetivus Distant, 1884: 284   , pl. 22, fig. 25 (orig. desc.); Atkinson, 1890: 42 (catalog).

Rhinocapsidea genetiva: Reuter, 1908: 54   ; Carvalho, 1952: 79; Carvalho 1958: 79 (catalog); Carvalho and Dolling, 1976: 798 (disc. of lectotype); Carvalho and Wallerstein, 1978: 254 (note); Schuh, 1995: 191 (catalog).

DIAGNOSIS: Recognized by the body entirely dark fuscous to black (some males) or with dorsum of head and pronotum yellowish brown with remainder of body dark fuscous to black (females, some males), antennal segments dark brown to fuscous, femora pale yellowish brown, tibiae brown, tarsi dark fuscous to black (fig. 2); labium not reaching posterior margin of mesosternum, male with vertex width subequal to length of antennal segment I, width clearly longer in female; width of pronotum more or less subequal with length of antennal segment II; metepisternum dorsal to evaporative area of scent gland with microtrichia (fig. 20C). This species shares several significant characters, including the male genitalia, with J. cruralis   and J. sinaloa   . All, as seen from above, are almost devoid of setae including on the costal margin of the hemelytron. The cuticular sculpturing on the hemelytron is prominent and very easy to see. All are about equal in size. The similarity in color between J. genetivus   and J. cruralis   pertains to those males of J. genetivus   that have a uniformly dark fuscous to black pronotum. The antenna of J. genetivus   is dark fuscous whereas it is pale in J. cruralis   and J. sinaloa   . The last species is conspicuously uniformly pale yellowish brown.

REDESCRIPTION: Male: COLORATION: Generally body either entirely dark fuscous to black or frequently with dorsum of head and pronotum yellowish brown with remain- der of body dark fuscous to black. Head ranging from almost completely yellowish brown (rare) to black with fuscous markings occurring as a spot on vertex near each eye, posterior margin of vertex and basal half of clypeus and adjacent region of frons; antennal segments dark brown to fuscous, segment II sometimes darker than others; labium pale yellow, segment I fuscous at base or sometimes entirely so, apex of labium fuscous. Pronotum usually fuscous to black, dorsum also sometimes yellowish brown. Hemelytron dark fuscous to black; coxa usually pale with base of metacoxa pale, femora pale yellowish brown, tibiae brown, tarsi dark fuscous to black. Abdomen dark fuscous to black. VESTITURE: Body as seen from above glabrous, almost devoid of setae. Head with short sparse setae; segment I of antenna with a few scattered semierect setae, remaining segments covered with short decumbent setae with very few, if any, as long as diameter of segment on which located. Pronotum almost without setae, those present most commonly decumbent and located along lateral margins; meso- and metapleurae with scattered erect setae; hemelytron with short decumbent setae located along costal margin and on or near cuneus; underside of abdomen with moderate numbers of semierect setae. GEN- ITALIA (fig. 21): Genital segment with bifurcate tergal process projecting from region slightly to right of midline on dorsal margin of aperture; lateral process smoothly margined, elongate, curved medially with rounded apex, medial base of process with minute dorsal point; ventroposterior margin of capsule with minute notch; distal width of subgenital plate moderately wide, strongly projecting beyond aperture of capsule on right side. Left paramere C-shaped in dorsal view; without sensory lobe; diameter of paramere gradually expanded to wide deeply

0.20 mm.

notched apical region with small ventral point and wide, rounded dorsolateral margin. Right paramere elongate, C-shaped, slightly more than twice length of left paramere; diameter of paramere widest basally, gradually tapered to strongly recurved apex; apex terminating in two small spines; distal region with several stout medial spines. Phallotheca elongate conical, 1/3 length of spiculum; aperture large, open on right side, apex, and distal on left side. Vesica with one, long medially thickened spiculum, situated on dorsal surface of ductus seminis; apically region of spiculum bifurcate with recurved, flattened, strongly serrate branches reaching to base of spiculum, one branch twice as wide as other; base of spiculum sinuate, broadly affixed to dorsal margin of ductus, base of spiculum with narrow, pointed process behind ductus seminis on left side.

Female: Broader in general appearance than male. COLORATION: Generally body with dorsum of head and dorsum of pronotum, sometimes all of prothorax yellowish brown but with some fuscous coloration on head, remainder of body dark fuscous to black; head ranging from almost entirely yellowish brown (rare) to yellowish brown with limited fuscous areas above, apex of clypeus, maxillary plate, and underside of head dark fuscous to black; segment I of antenna brown, remaining segments usually yellowish brown, darker fuscous apically or segments III and IV uniformly pale fuscous; labium pale yellow, fuscous to black at base of segment I and apex of labium. Pronotum ranging from all yellowish brown (rare) to yellowish-brown coloration limited to dorsum, remainder of body same as male.

VESTITURE: Similar to that of males. GENITALIA with strongly sclerotized lateral region of VLP produced anteriorly, ventral margin, not produced ventrally into VUL; SR moderately large, ovoid in longitudinal axis; medial region of DLP relatively small; posterior wall with well-sclerotized, convex medial section with small anterior projection; ventral margin of IRL not deeply incised (fig. 19B) with ventralmost margin blunt; otherwise as in generic description.

MEASUREMENTS: Male (n 5 20; average followed in parentheses by range): Length, 4.38 (3.94–4.70); width, 1.53 (1.40– 1.66). Head length, 0.23 (0.18–0.26); width, 0.91 (0.84–0.92); vertex width, 0.40 (0.38– 0.44). Length of antennal segment I, 0.38 (0.36–0.42); II, 1.41 (1.28–1.54); III, 1.05 (0.96–1.14); IV, 0.44 (0.38–0.50). Pronotal length, 0.70 (0.64–0.74); width across base, 1.36 (1.22–1.44). Cuneal length, 0.72 (0.64– 0.80); width across base, 0.51 (0.46–0.54).

Female: (n 5 20; average followed in parentheses by range): Length, 4.28 (3.84– 4.60); width, 1.79 (1.64–1.90). Head length, 0.28 (0.24–0.32); width, 0.93 (0.90–0.96); vertex width, 0.45 (0.42–0.46). Length of antennal segment I, 0.38 (0.36–0.40); II, 1.45 (1.32–1.58); III, 1.00 (0.92–1.10); IV, 0.47 (0.38–0.50). Pronotal length, 0.75 (0.70– 0.84); width across base, 1.41 (1.34–1.52).

DISCUSSION: No other species of Jornandes   has the type of color variation exhibited by this species. All specimens as seen from above have dark fuscous to black hemelytra. All females have the dorsum of the pronotum yellowish brown that may extend downward to varying degrees onto the propleuron. The majority of the males available for study have the hemelytra and pronotum dark fuscous to black. The head is usually dark fuscous and with a few brown areas. Other males have the pronotum and head yellowish brown as in the female. Rarely the pronotum is mostly dark with yellowish brown limited to a diffuse medial area. The recurved region of the vesical spiculum in Chiapas males is narrower and situated closer to the main body of the spiculum than in the vesical spiculum of specimens from the reminder of the distribution of J. genetivus   . The vesical spiculum of J. cruralis   , J. genetivus   , and J. sinaloa   are conspicuously bifurcate with broad central trunks and all share apically spinose, elongate right parameres.

The female lectotype is deposited in the Humboldt University Museum in Berlin, Germany. The specimen is teneral, however, the yellowish-brown pronotum, elongate body, and mostly parallel-sided costal margin easily distinguish this species from congeners. The measurements of the lectotype given by Carvalho and Dolling (1976) and as examined by us fit within the range given above.

HOST PLANTS: Montana tomentosa Cerv.   ( Asteraceae   ), Ipomoea walcotiana Rose   ( Convolvulaceae   ), and Euphorbiaceae   .

DISTRIBUTION: Known from 54 localities from northern Puebla and Mexico state south through Morelos, Guerrero, and Oaxaca with one collection in Chiapas (fig. 10C).

SPECIMENS EXAMINED: MEXICO: Chiapas: km 20 Motozintla–Comitan, 15.43324 ° N 92.12 ° W, 900 m, 08 Jul 1988, A. Cadena and L. Cervantes, 13 ( AMNH _ PBI 00245155 View Materials ), 1♀ ( AMNH _ PBI 00245156 View Materials ) ( TAMU). Guerrero: 4 mi W of Chilpancingo, 17.56992 ° N 99.56555 ° W, 1506 m, 15 Jul 1984, Carroll, Schaffner, Friedlander, 1♀ ( AMNH _ PBI 00245175 View Materials ) ( TAMU). 6 mi E of Xochipala, 17.7936 ° N 99.54263 ° W, 1067 m, 18 Jul 1984, Carroll, Schaffner, Friedlander, 3♀ ( AMNH _ PBI 00106432 View Materials , AMNH _ PBI 00245177 View Materials – AMNH _ PBI 00245178 View Materials ) ( TAMU). 6 mi NE of Tixtla de Guerrero, 17.64501 ° N 99.36861 ° W, 16 Jul 1984, Carroll, Schaffner, Friedlander, 13 ( AMNH _ PBI 00245022 View Materials ), 2♀ ( AMNH _ PBI 00245023 View Materials , AMNH _ PBI 00245024 View Materials ) ( TAMU). Chilpancingo, 17.55 ° N 99.5 ° W, 1402 m, 24 Jul 1961, R. and K. Dreisbach, 13 ( AMNH _ PBI 00134073 View Materials ) ( USNM). Taxco, 18.55976 ° N 99.61491 ° W, 2072 m, 02 Sep 1966, J. and W. Ivie, 1♀ ( AMNH _ PBI 00108439 View Materials ) ( AMNH); 02 Aug 1938, L. J. Lipovsky, 1♀ ( AMNH _ PBI 00245176 View Materials ) ( UCB). Mexico: Malinalco, 18.95 ° N 99.5 ° W, 1743 m, 26 Jun 1980, Harry Brailovsky, 1♀ ( UNAM); 24 Oct 1977, D. Pluot, 13 ( AMNH _ PBI 00243973 View Materials ), 3♀ ( AMNH _ PBI 00243974 View Materials – AMNH _ PBI 00243976 View Materials ) ( MNHN). Tenayuca, 19.5269 ° N 99.2217 ° W, 2277 m, 28 Oct 1994, H. Brailovsky and C. Mayorga, 1♀ ( AMNH _ PBI 00244691 View Materials ) ( UNAM). Morelos: 10 mi E of Cuernavaca, 18.9166 ° N 99.09631 ° W, 30 Jul 1976, Peigler, Gruetzmacher, R &M Murray, Schaffner, 2♀ ( AMNH _ PBI 00106431 View Materials , AMNH _ PBI 00245043 View Materials ) ( TAMU). 12 mi E of Cuernavaca, 18.89741 ° N 99.06354 ° W, 1280 m, 12 Aug 1954, R. R. Dreisbach, 13 ( AMNH _ PBI 00121643 View Materials ) ( CAS). 4♀ ( AMNH _ PBI 00114327 View Materials – AMNH _ PBI 00114330 View Materials ) ( CNC). 1♀ ( AMNH _ PBI 00127450 View Materials ) ( UMMC); 14 Aug 1954, J. G. Chillcott, 13 ( AMNH _ PBI 00112263 View Materials ), 8♀ ( AMNH _ PBI 00113158 View Materials – AMNH _ PBI 00113163 View Materials , AMNH_PBI 00114334– AMNH _ PBI 00114335 View Materials ) ( CNC); 14 Aug 1954, University of Kansas Mexico Expedition , 1♀ ( AMNH _ PBI 00117947 View Materials ) ( KU). 14 mi S of Cuernavaca, 18.72454 ° N 99.26772 ° W, 975 m, 03 Aug 1954, J. G. Chillcott, 2♀ ( AMNH _ PBI 00112266 View Materials , AMNH _ PBI 00113164 View Materials ) ( CNC). 4.4 mi E of Cuernavaca, 18.91665 ° N 99.18237 ° W, 06 Jul 1974 – 08 Jul 1974, Clark, Murray, Ashe, Schaffner, 1♀ ( AMNH _ PBI 00245042 View Materials ) ( TAMU); 27 Jul 1976 – 29 Jul 1976, Peigler, Gruetzmacher, R. and M. Murray, Schaffner, 1♀ ( AMNH _ PBI 00245041 View Materials ), 13 ( AMNH _ PBI 00245038 View Materials ), 2♀ ( AMNH _ PBI 00245039 View Materials , AMNH _ PBI 00245040 View Materials ) ( TAMU). Acatlipa, 89 km, 18.81667 ° N 99.23333 ° W, 1300 m, 10 Aug 1946, Unknown, 1♀ ( AMNH _ PBI 00108422 View Materials ) ( AMNH). Cuernavaca , 18.91666 ° N 99.25 ° W, 15 Aug 1954, R. R. Dreisbach, 33 ( AMNH _ PBI 00127388 View Materials , AMNH _ PBI 00127446 View Materials – AMNH _ PBI 00127447 View Materials ), 2♀ ( AMNH _ PBI 00127448 View Materials , AMNH _ PBI 00127449 View Materials ) ( UMMC); 12 Jul 1961 – 19 Jul 1961, R. and K. Dreisbach, 2♀ ( AMNH _ PBI 00134070 View Materials , AMNH _ PBI 00134071 View Materials ) ( USNM); 20 Aug 1936, E. D. Ball, 1♀ ( AMNH _ PBI 00134072 View Materials ) ( USNM). El Polvorin, Villa de Ayala , 18.75 ° N 98.98333 ° W, 1147 m, 09 Jul 1972, H. Perez, 1♀ ( AMNH _ PBI 00245173 View Materials ) ( TAMU). 13 ( AMNH _ PBI 00244684 View Materials ) ( UNAM). km 14 Huautla-Jojutla a 123 Art, 18.54324 ° N 99.26731 ° W, 1055 m, 05 Jul 1995, H. Brailovsky and E. Barrera, 13 ( AMNH _ PBI 00244692 View Materials ) ( UNAM). Oaxaca: 1.1 km N of Hwy 190 (rd to Yagul) 29.5 km E of Oaxaca, 16.98293 ° N 96.45529 ° W, 1700 m, 13 Aug 1988, J. K. Liebherr and D. A. Yager, 13 ( AMNH _ PBI 00116224 View Materials ), 1♀ ( AMNH _ PBI 00116225 View Materials ) ( CUIC). 1.1 mi W of El Tule, 16.95676 ° N 96.81482 ° W, 1646 m, 17 Jul 1987, Kovarik and Schaffner, 73 ( AMNH _ PBI 00244685 View Materials – AMNH _ PBI 00244686 View Materials , AMNH_PBI 00245025– AMNH _ PBI 00245029 View Materials ), 5♀ ( AMNH _ PBI 00244687 View Materials , AMNH _ PBI 00245030 View Materials – AMNH _ PBI 00245033 View Materials ) ( TAMU). 10 mi N of Miltepec, 16.71209 ° N 94.61666 ° W, 04 Aug 1976, Peigler, Gruetzmacher, R &M Murray, Schaffner, 13 ( AMNH _ PBI 00094247 View Materials ) ( CNC). Monarda tomentosa Cerv.   ( Asteraceae   ), 13 ( AMNH _ PBI 00244969 View Materials ), 8♀ ( AMNH _ PBI 00244970 View Materials – AMNH _ PBI 00244977 View Materials ), 53 ( AMNH _ PBI 00244957 View Materials – AMNH _ PBI 00244961 View Materials ), 7♀ ( AMNH _ PBI 00244962 View Materials – AMNH _ PBI 00244968 View Materials ) ( TAMU); 15 Jul 1971 – 16 Jul 1971, Clark, Murray, Hart, Schaffner, 13 ( AMNH _ PBI 00112262 View Materials ) ( CNC). Ipomoea walcotiana Rose   ( Convolvulaceae   ), 103 ( AMNH _ PBI 00244842 View Materials – AMNH _ PBI 00244851 View Materials ), 9♀ ( AMNH _ PBI 00244852 View Materials – AMNH _ PBI 00244860 View Materials ), 103 ( AMNH _ PBI 00244820 View Materials – AMNH _ PBI 00244829 View Materials ), 12♀ ( AMNH _ PBI 00244830 View Materials – AMNH _ PBI 00244841 View Materials ) ( TAMU); 11 Jul 1973 – 12 Jul 1973, Mastro and Schaffner, 13 ( AMNH _ PBI 00244869 View Materials ) ( TAMU); 09 Aug 1980, Schaffner, Weaver, Friedlander, Ipomoea walcotiana Rose   ( Convolvulaceae   ), 53 ( AMNH _ PBI 00244861 View Materials – AMNH _ PBI 00244865 View Materials ), 13 ( AMNH _ PBI 00244866 View Materials ), 2♀ ( AMNH _ PBI 00244867 View Materials , AMNH _ PBI 00244868 View Materials ) ( TAMU); 15 Jul 1971, Clark, Murray, Hart, Schaffner, 13 ( AMNH _ PBI 02448670 View Materials ) ( TAMU). 21 km E of Mitla, 16.94929 ° N 96.2529 ° W, 1650 m, 17 Aug 1988, Doyen, 2♀ ( AMNH _ PBI 00119463 View Materials , AMNH _ PBI 00119464 View Materials ) ( UCB). 3.4 mi SE of Matatlan, 16.82948 ° N 96.3544 ° W, 1930 m, 06 Aug 1980 – 07 Aug 1980, Schaffner, Weaver, Friedlander, 33 ( AMNH _ PBI 00245034 View Materials – AMNH _ PBI 00245036 View Materials ) ( TAMU). 30 mi NE of Huajuapan de Leon, 18.10746 ° N 97.74907 ° W, 1800 m, 28 Jun 1965, Burke, Meyer, Schaffner, 1♀ ( AMNH _ PBI 00112261 View Materials ) ( CNC). Ipomoea walcotiana Vasey   ( Convolvulaceae   ), 5♀ ( AMNH _ PBI 00244918 View Materials – AMNH _ PBI 00244922 View Materials ), 93 ( AMNH _ PBI 00244908 View Materials – AMNH _ PBI 00244915 View Materials , AMNH _ PBI 00244917 View Materials ), 1♀ ( AMNH _ PBI 00244916 View Materials ) ( TAMU). 31.3 km SE of Huajuapan de Leon, 17.66531 ° N 97.5606 ° W, 2000 m, 11 Aug 1988, J. K. Liebherr and D. A. Yager, 1♀ ( AMNH _ PBI 00116223 View Materials ) ( CUIC). Huajuapan , 17.8 ° N 97.76667 ° W, 1633 m, 25 Aug 1969, L. A. Kelton, ( Euphorbiaceae   ), 173 ( AMNH _ PBI 00112259 View Materials , AMNH _ PBI 00112439 View Materials ,AMNH_PBI 00112962, AMNH_PBI 00113205– AMNH _ PBI 00113218 View Materials ), 26♀ ( AMNH _ PBI 00112260 View Materials , AMNH _ PBI 00113219 View Materials – AMNH _ PBI 00113243 View Materials ), 23 ( AMNH _ PBI 00111003 View Materials , AMNH _ PBI 00112920 View Materials ) ( CNC). Huajuapan de Leon , 17.8 ° N 97.76667 ° W, 1633 m, 17 Sep 1980, Harry Brailovsky, 4♀ ( AMNH _ PBI 00244680 View Materials – AMNH _ PBI 00244683 View Materials ) ( UNAM). 1♀ ( AMNH _ PBI 00133593 View Materials ) ( USNM). Oaxaca , [17.03333 ° N 96.73333 ° W], Deppe, Lectotype, 1♀ ( ZMHB). Puebla: 0.5 km E of Tecamachalco, 18.88333 ° N 97.705 ° W, 2070 m, 29 Jul 1995, T. J. Henry and E. Barrera, Solanum sp.   ( Solanaceae   ), 73 ( AMNH _ PBI 00133548 View Materials , AMNH _ PBI 00133591 View Materials , AMNH_PBI 00133763– AMNH _ PBI 00133767 View Materials ), 20♀ ( AMNH _ PBI 00133768 View Materials – AMNH _ PBI 00133787 View Materials ) ( USNM). 1 mi SW of Zapotitlan, 19.99794 ° N 97.75225 ° W, 1293 m, 11 Jul 1973, Mastro and Schaffner , 103 ( AMNH _ PBI 00244923 View Materials – AMNH _ PBI 00244932 View Materials ), 9♀ ( AMNH _ PBI 00244933 View Materials – AMNH _ PBI 00244941 View Materials ) ( TAMU). 1.1 mi W of Acatlan, 18.19999 ° N 98.06683 ° W, 13 Jul 1974, Clark , Murray , Ashe , Schaffner , 3♀ ( AMNH _ PBI 00119131 View Materials – AMNH _ PBI 00119133 View Materials ) ( CNC). 13 ( AMNH _ PBI 00244956 View Materials ), 2♀ ( AMNH _ PBI 00244954 View Materials , AMNH _ PBI 00244955 View Materials ), 83 ( AMNH _ PBI 00244942 View Materials – AMNH _ PBI 00244949 View Materials ), 4♀ ( AMNH _ PBI 00244950 View Materials – AMNH _ PBI 00244953 View Materials ) ( TAMU). 10 mi SE of Tehuitzingo, 18.24456 ° N 98.20488 ° W, 1189 m, 03 Jul 1953, University of Kansas Mexico Expedition , 13 ( AMNH _ PBI 00108421 View Materials ), 6♀ ( AMNH _ PBI 00169968 View Materials - AMNH _ PBI 00169973 View Materials ) ( AMNH). 4♀ ( AMNH _ PBI 00117943 View Materials – AMNH _ PBI 00117946 View Materials ) ( KU). 11.8 mi NW of Izucar de Matamoros, 18.66106 ° N 98.62248 ° W, 1395 m, 13 Jul 1974, Clark , Murray , Ashe , Schaffner , 4♀ ( AMNH _ PBI 00244715 View Materials – AMNH _ PBI 00244718 View Materials ) ( TAMU). 12 mi NW of Petlalcingo, 18.19489 ° N 98.04667 ° W, 1219 m, 03 Jul 1953, University of Kansas Mexico Expedition , 13 ( AMNH _ PBI 00117802 View Materials ) ( KU). 13 ( AMNH _ PBI 00117942 View Materials ) ( KU). 13.7 mi SW of Izucar de Matamoros, 18.50612 ° N 98.63298 ° W, 1080 m, 21 Jul 1981, Bogar , Schaffner and Friedlander , 1♀ ( AMNH _ PBI 00244721 View Materials ) ( TAMU). 14 km NE of Zinacantepec, 18.43666 ° N 97.215 ° W, 1920 m, 01 Aug 1995, T. J. Henry and H. Brailovsky, 13 ( AMNH _ PBI 00134079 View Materials ) ( USNM). 14 mi N of Acatlan, 19.00658 ° N 97.90422 ° W, 2262 m, 12 Aug 1978, Plitt and Schaffner , 23 ( AMNH _ PBI 00094265 View Materials , AMNH _ PBI 00094268 View Materials ) ( CNC). 103 ( AMNH _ PBI 00244756 View Materials – AMNH _ PBI 00244765 View Materials ), 22♀ ( AMNH _ PBI 00244769 View Materials – AMNH _ PBI 00244790 View Materials ) ( TAMU). 14.4 mi SW of Izucar de Matamoros, 18.5312 ° N 98.57215 ° W, 1268 m, 01 Aug 1976, Peigler , Gruetzmacher, R. and M. Murray, Schaffner , 1♀ ( AMNH _ PBI 00094264 View Materials ) ( CNC). 33 ( AMNH _ PBI 00244693 View Materials – AMNH _ PBI 00244695 View Materials ), 14♀ ( AMNH _ PBI 00244696 View Materials – AMNH _ PBI 00244709 View Materials ) ( TAMU). 13 ( AMNH _ PBI 00107581 View Materials ) ( USNM). 14.5 mi SE of Acatlan, 18.9095 ° N 97.763 ° W, 2035 m, 21 Jul 1981, Bogar , Schaffner and Friedlander , 2♀ ( AMNH _ PBI 00119116 View Materials , AMNH _ PBI 00119117 View Materials ) ( CNC). 143 ( AMNH _ PBI 00244978 View Materials – AMNH _ PBI 00244991 View Materials ), 30♀ ( AMNH _ PBI 00244992 View Materials – AMNH _ PBI 00245021 View Materials ) ( TAMU). 2 km N of Coxcatlan, 18.28307 ° N 97.15849 ° W, 960 m, 20 Aug 1988, J. T. Doyen, 1♀ ( AMNH _ PBI 00119462 View Materials ) ( UCB). 3 km al NW de Tecaltzingo, 18.73191 ° N 97.68368 ° W, 1890 m, 04 Jul 1995, E. Barrera, 2♀ ( AMNH _ PBI 00244688 View Materials , AMNH _ PBI 00244689 View Materials ) ( UNAM); 13 Aug 1992, E. Barrera and C. Mayorga, 1♀ ( AMNH _ PBI 00244690 View Materials ) ( UNAM). 3 mi N of Petlalcingo, 18.1394 ° N 97.90568 ° W, 1465 m, 03 Aug 1963, F. D. Parker and L. A. Stange, 13 ( AMNH _ PBI 00245169 View Materials ) ( UCB). 33 km SE of Acatlan, nr Salitrillo , 17.97866 ° N 97.87666 ° W, 1500 m, 09 Aug 1988, J. Doyen, 23 ( AMNH _ PBI 00119457 View Materials , AMNH _ PBI 00119531 View Materials ), 8♀ ( AMNH _ PBI 00119520 View Materials – AMNH _ PBI 00119522 View Materials , AMNH_PBI 00119532– AMNH_PBI 00119535, AMNH _ PBI 00119588 View Materials ) ( UCB). 36 km NW of Huajuapan de Leon, 18.07587 ° N 97.9279 ° W, 1600 m, 09 Aug 1988, J. K. Liebherr and D. A. Yager, 2♀ ( AMNH _ PBI 00116226 View Materials , AMNH _ PBI 00116227 View Materials ) ( CUIC). 4 mi SW of Acatepec, 18.9922 ° N 98.32683 ° W, 21 Jul 1984, Carroll , Schaffner , Friedlander , 2♀ ( AMNH _ PBI 00094263 View Materials , AMNH _ PBI 00119130 View Materials ) ( CNC); 11 Jul 1973, Mastro and Schaffner , 13 ( AMNH _ PBI 00094267 View Materials ) ( CNC). 33 ( AMNH _ PBI 00244735 View Materials – AMNH _ PBI 00244737 View Materials ), 4♀ ( AMNH _ PBI 00244738 View Materials – AMNH _ PBI 00244741 View Materials ) ( TAMU). 13 ( AMNH _ PBI 00107582 View Materials ) ( USNM). 4 mi W of Acatepec, 19.03332 ° N 98.34485 ° W, 12 Aug 1978, Plitt and Schaffner , 33 ( AMNH _ PBI 00244766 View Materials – AMNH _ PBI 00244768 View Materials ), 9♀ ( AMNH _ PBI 00244791 View Materials – AMNH _ PBI 00244799 View Materials ) ( TAMU); 09 Aug 1980, Schaffner , Weaver , Friedlander , Ipomoea walcotiana Rose   ( Convolvulaceae   ), 23 ( AMNH _ PBI 00244752 View Materials , AMNH _ PBI 00244753 View Materials ), 2♀ ( AMNH _ PBI 00244754 View Materials , AMNH _ PBI 00244755 View Materials ) ( TAMU); 26 Jul 1973, Mastro and Schaffner , 93 ( AMNH _ PBI 00244724 View Materials – AMNH _ PBI 00244732 View Materials ), 2♀ ( AMNH _ PBI 00244733 View Materials , AMNH _ PBI 00244734 View Materials ) ( TAMU). 4.3 mi SW of Acatepec, 19.03229 ° N 98.28407 ° W, 2141 m, 16 Jul 1971, Clark , Murray , Hart , Schaffner , 3♀ ( AMNH _ PBI 00244742 View Materials – AMNH _ PBI 00244744 View Materials ) ( TAMU). 4.4 mi SW of Acatepec, 18.98809 ° N 98.33118 ° W, 26 Jul 1974, Clark , Murray , Ashe , Schaffner , 133 ( AMNH _ PBI 00244800 View Materials – AMNH _ PBI 00244812 View Materials ), 7♀ ( AMNH _ PBI 00244813 View Materials – AMNH _ PBI 00244819 View Materials ) ( TAMU); 09 Jul 1977, J. C. Schaffner, 63 ( AMNH _ PBI 00094266 View Materials , AMNH _ PBI 00119115 View Materials , AMNH_PBI 00184939– AMNH _ PBI 00184942 View Materials ) ( CNC). 573 ( AMNH _ PBI 00118205 View Materials , AMNH _ PBI 00245048 View Materials – AMNH _ PBI 00245103 View Materials ), 51♀ ( AMNH _ PBI 00245104 View Materials – AMNH _ PBI 00245154 View Materials ) ( TAMU); 09 Jul 1981, Bogar , Schaffner and Friedlander , 13 ( AMNH _ PBI 00244745 View Materials ), 1♀ ( AMNH _ PBI 00244746 View Materials ) ( TAMU); 21 Jul 1984, J. B. Woolley, 13 ( AMNH _ PBI 00244747 View Materials ), 2♀ ( AMNH _ PBI 00244748 View Materials , AMNH _ PBI 00244749 View Materials ) ( TAMU); 24 Jul 1984, Carroll , Schaffner , Woolley , Friedlander , 13( AMNH _ PBI00244750 View Materials )( TAMU); 21 Jul 1984, Carroll , Schaffner , Friedlander , 193 ( AMNH _ PBI 00244871 View Materials – AMNH _ PBI 00244889 View Materials ), 18♀ ( AMNH _ PBI 00244890 View Materials – AMNH _ PBI 00244907 View Materials ) ( TAMU). 5 km NE of Zinacantepec, 18.37 ° N 97.21 ° W, 1290 m, 01 Aug 1995, T. J. Henry and E. Barrera, 23 ( AMNH _ PBI 00133549 View Materials , AMNH _ PBI 00133788 View Materials ), 12♀ ( AMNH _ PBI 00133789 View Materials - AMNH _ PBI 00133800 View Materials ) ( USNM). 5 mi SE of Izucar de Matamoros, 18.54858 ° N 98.41242 ° W, 20 Jul 1984, Carroll , Schaffner , Friedlander , 2♀ ( AMNH _ PBI 00244722 View Materials , AMNH _ PBI 00244723 View Materials ) ( TAMU). 6 km S of Chila, 20.27564 ° N 98.19646 ° W, 1737 m, 10 Aug 1967, H.R. Burke and J. Hafernik, 1♀ ( AMNH _ PBI 00245037 View Materials ) ( TAMU). 6 mi N of Tehuacan, 18.59464 ° N 97.4041 ° W, 1986 m, 22 Aug 1987, J. Doyen, 13 ( AMNH _ PBI 00119460 View Materials ) ( UCB). 6 mi SW of Tehuacan, 18.3883 ° N 97.44836 ° W, 08 Jul 1973 – 10 Jul 1973, Mastro and Schaffner , 13 ( AMNH _ PBI 00106430 View Materials ) ( TAMU); 08 Jul 1981, Bogar , Schaffner and Friedlander , 1♀ ( AMNH _ PBI 00245047 View Materials ) ( TAMU). 6.3 mi N of Tehuacan, 18.53786 ° N 97.41726 ° W, 1798 m, 22 Jul 1987, Kovarik and Schaffner , 13 ( AMNH _ PBI 00245044 View Materials ), 1♀ ( AMNH _ PBI 00245045 View Materials ) ( TAMU); 21 Jun 1987, R. Anderson, 1♀ ( AMNH _ PBI 00245046 View Materials ) ( TAMU). 7 mi N of Atlixco, 18.97124 ° N 98.35886 ° W, 2168 m, 12 Aug 1957, J. A. Chemsak, B.J. Rannells, 13 ( AMNH _ PBI 00119461 View Materials ), 1♀ ( AMNH _ PBI 00119589 View Materials ) ( UCB). 7.3 mi SW of Izucar de Matamoros, 18.52184 ° N 98.57917 ° W, 22 Jul 1981, Bogar , Schaffner and Friedlander , 13 ( AMNH _ PBI 00119114 View Materials ) ( CNC). 23( AMNH _ PBI 00244710 View Materials , AMNH _ PBI 00244711 View Materials ), 3♀ ( AMNH _ PBI 00244712 View Materials – AMNH _ PBI 00244714 View Materials ) ( TAMU); 01 Aug 1976, Peigler , Gruetzmacher, R. and M. Murray, Schaffner , 2♀ ( AMNH _ PBI 00244719 View Materials , AMNH _ PBI 00244720 View Materials ) ( TAMU). Chila , 20.3 ° N 98.2 ° W, 1200 m, 25 Aug 1969, L. A. Kelton, 5♀ ( AMNH _ PBI 00112265 View Materials , AMNH _ PBI 00112963 View Materials , AMNH_PBI 00114331– AMNH _ PBI 00114333 View Materials ) ( CNC). San Martin Texmelucan , 19.28333 ° N 98.43333 ° W, 2237 m, 27 Aug 1969, L. A. Kelton, 1♀ ( AMNH _ PBI 00112264 View Materials ) ( CNC). GoogleMaps  

AMNH

American Museum of Natural History

MNHN

Museum National d'Histoire Naturelle

CNC

Canadian National Collection of Insects, Arachnids, and Nematodes

USNM

Smithsonian Institution, National Museum of Natural History

UNAM

Universidad Nacional Autonoma de Mexico

UCB

University of California at Berkeley

CAS

California Academy of Sciences

KU

Biodiversity Institute, University of Kansas

CUIC

Cornell University Insect Collection

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Miridae

Genus

Jornandes

Loc

Jornandes cruralis Distant

Schaffner, JC & Schwartz, MD 2008
2008
Loc

Rhinocapsidea genetiva: Reuter, 1908: 54

Schuh, R. T. 1995: 191
Carvalho, J. C. M. & P. Wallerstein 1978: 254
Carvalho, J. C. M. & W. R. Dolling 1976: 798
Carvalho, J. C. M. 1958: 79
Carvalho, J. C. M. 1952: 79
Reuter, O. M. 1908: 54
1908
Loc

Jornandes cruralis

Schuh, R. T. 1995: 127
Carvalho, J. C. M. 1981: 4
Carvalho, J. C. M. & W. R. Dolling 1976: 795
Carvalho, J. C. M. 1958: 80
Distant, W. L. 1893: 448
1893
Loc

Eccritotarsus genetivus

Atkinson, E. T. 1890: 42
Distant, W. L. 1884: 284
1884