Trechus brucki Fairmaire, 1862

Faille, Arnaud, Bourdeau, Charles & Fresneda, Javier, 2012, Molecular phylogeny of the Trechus brucki group, with description of two new species from the Pyreneo-Cantabrian area (France, Spain) (Coleoptera, Carabidae, Trechinae), ZooKeys 217, pp. 11-51 : 15-16

Trechus brucki Fairmaire, 1862 Figs 71427, 2833

Type locality.

«Eaux–Bonnes, M. vom Bruck" (Fairmaire, 1862b). France, Pyrénées–Atlantiques.

Type series.

Lectotype (MNHN), present designation: 1 ♂, labelled: "oblongulus Bonnes" [white rectangular label (ms, Fairmaire)], «Bruckii» [white rectangular label (ms, Fairmaire)], "MUSEUM PARIS Collection Léon Fairmaire 1906" [white rectangular label (printed)], «TYPE» [red rectangular label (printed)], "Lectotypus / Trechus bruckii Fairmaire / Faille, Bourdeau & / Fresneda des. 2012" [red rectangular label (printed)], genitalia dissected and mounted in a separate label pinned with the specimen. Paralectotype (MNHN): 1 ♀, same label data and pin as lectotype except "Paralectotypus / Trechus bruckii Fairmaire / Faille, Bourdeau & / Fresneda des. 2012" [red rectangular label (printed)].

Type series of Trechus planiusculus Fairmaire, 1862.

Lectotype (MNHN), present designation: 1 ♀ (red dot), labelled: “oblongus” [white rectangular label (ms, Fairmaire)], “planiusculus” [white rectangular label (ms, Fairmaire)], “Bruckii” [white rectangular label (ms, Fairmaire)], “2203” [white rectangular label (ms, Fairmaire)], "MUSEUM PARIS Collection Léon Fairmaire 1906" [white rectangular label (printed)], “TYPE” [red rectangular label (printed)], "Lectotypus / Trechus planiusculus Fairmaire / Faille, Bourdeau & / Fresneda des. 2012" [red rectangular label (printed)]. Paralectotypes (MNHN): 1 ♀, same label data and pin as lectotype except "Paralectotypus / Trechus planiusculus Frm / Faille, Bourdeau & / Fresneda des. 2012" [red rectangular label (printed)]; 1 ♂, "H Pyrenees 1856 M. Pandellé” [white rectangular label (printed)], “Bruckii” [white rectangular label (ms, Fairmaire)], “COTYPE” [white and red rectangular label (printed)], "R. Jeannel Brucki Fr" [white rectangular label (ms, Jeannel)], "MUSEUM PARIS coll. R. JEANNEL 1931" [white rectangular label (printed)], "Paralectotypus / Trechus planiusculus Frm / Faille, Bourdeau & / Fresneda des. 2012" [red rectangular label (printed)], genitalia dissected and mounted in a separate label pinned with the specimen.

Non Type material.

1 ♀ (MNHN) labelled: “planiusculus” [white rectangular label (ms, Fairmaire?)], “Bruckii” [white rectangular label (ms, Fairmaire)], "MUSEUM PARIS Collection Léon Fairmaire 1906" [white rectangular label (printed)], "R. Jeannel Brucki Fr" [white rectangular label (ms, Jeannel)]. We do not consider this specimen as a syntype of Trechus planiusculus as it is not labeled " oblongus " as the specimen of the type series, suggesting that the specimen arrived in the Fairmaire collection after the description of planiusculus. A second female specimen (MNHN) labelled: "oblongus Arrens" [white rectangular label (ms, Fairmaire)], " TYPE" [white and red rectangular label (printed)], "MUSEUM PARIS Collection Léon Fairmaire 1906" [white rectangular label (printed)]. This specimen could be the reference specimen of Trechus oblongus Schaum, 1862. Reference of the name comes from Schaum (1862: addenda, p. 119): "P. 14 col. 2 Trechus oblongus Schaum;" only the name is mentioned, without any description, number of exemplars studied or locality. It should then be considered as nomen nudum. Jeannel (1927) indicates that Trechus oblongus is a synonym of Trechus brucki with type locality: “Pyrén. occ." We were unable to find the specimen or reference where Jeannel found the type locality.

Taxonomic comments

The study of specimens of Trechus brucki pecoudi from Orhy and of numerous exemplars of Trechus brucki ,including types of the previously described subspecies of Trechus brucki ,demonstrated that none of the characters quoted either by Colas and Gaudin (1935) or by Queinnec and Ollivier (2011) are constant. We consider then the subspecies pecoudi as synonymous of Trechus brucki : Trechus brucki brucki Fairmaire, 1862 = Trechus brucki pecoudi Colas & Gaudin, 1935, syn. n.

Trechus politus and Trechus planiusculus were described by Fairmaire in the volume of the Annales de la Société Entomologique de France of 1861 published in 1862 (Fairmaire, 1862b). As the name Trechus politus was already used for an American species (today Trechisibus politus Brullé, 1842), Fairmaire (1862a) changed the name of this species to Trechus bruckii . Trechus planiusculus was considered synonymous with Trechus brucki by Jeannel (1927), and, moreover, the name planiusculus was preoccupied as it was used by Costa (1858) in a work on Italian fauna. In his works on Trechini , Jeannel (1927, 1941) illustrated the genitalia of a male from the Ossau Valley. Recently, in a revision of the french fauna of Carabidae , Queinnec and Ollivier (2011) suggested that the drawing of Jeannel (1927, 1941) was incorrect, and that the male genitalia of Trechus brucki has a homogeneous shape throughout the distribution area. By examining the types of Fairmaire, we noticed that the drawing of Jeannel (1927, 1941) does not actually match with Trechus brucki . However, by studying specimens from Ossau Valley we found that the drawing of Jeannel actually corresponds to another undescribed species, very narrowly located in the area of Pic de Montagnon ( Bielle–Pyrénées Atlantiques). Here we describe this new species as Trechus bruckoides sp. n.