Agathidium virile Fall

WHEELER, QUENTIN D. & MILLER, KELLY B., 2005, Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part I. Coleoptera: Leiodidae, Bulletin of the American Museum of Natural History 2005 (290), pp. 1-95 : 74-79

Agathidium virile Fall Figures 101 , 149–156

Agathidium virile Fall, 1901: 219, 1934 . Agathidium hebetatum Hatch, 1957: 35 . NEW SYN­

ONYM.

slightly expanded subapically, apex blunt (fig. 148), in lateral aspect apex recurved dorsally, apical quarter approximately straight, irregularly shaped along ventral orifice, with almost toothlike projection (fig. 147); operculum deeply emarginate, each part elongate, bluntly rounded (fig. 148); lateral lobes blunt (figs. 147, 148).

Female tarsi 5–4–4.

DISTRIBUTION: This species is known from British Columbia, Canada (fig. 100).

SPECIMENS EXAMINED: CANADA: British Columbia: Toco , 28 Apr 1984, R and J Camenisch (1, PECK) ; Queen Charlotte Islands, Moresby Island, 2.5 mi N Mosquito Lake Campground, 5 Jun 1984, Spruce forest litter, RS Anderson (1, PECK) ; Charlotte Islands, Graham Island 1 mi NW Tlell, 27 Jun 1984, Sitka spruce, hemlock forest, RS Anderson (1, PECK) ; Stickeen R. Canyon, Liebeck (1, MCZC) .

DISCUSSION: There have been no reported hosts. The type was collected from ‘‘sifting

TYPE MATERIAL: Agathidium virile : holotype, Ƌ in MCZC labeled ‘‘Ƌ/ Pom Cal Mar. 25.93 [date handwritten]/ TYPE virile [‘TYPE’ underlined in red, ‘virile’ handwritten]/ M.C.Z. Type 24045 [red label]/ H.C. FALL COLLECTION / HOLOTYPE, Agathidium virile Fall, 1901 [red label with black line border]’’.

Agathidium hebetatum : holotype, Ƌ, USNM, labeled ‘‘McMinnville, ORE X­20– 41 [handwritten]/ TYPE Ƌ Agathidium (s.str.) hebetatum 1955 —M. Hatch [handwritten, red label]/ HOLOTYPE Agathidium hebatatum Hatch, 1957 [red label with black line border]’’.

TYPE LOCALITY: Agathidium virile : United States, California, Los Angeles Co., Pomona.

Agathidium hebatatum : United States, Oregon, Yamhill Co., McMinnville.

DIAGNOSIS: This variable species is distinguished from others in the A. revolvens species group by the following combination of characters: body broadly oval, convex, partially contractile (figs. 153, 154); dorsal punctation dense, confused, coarsely to weakly impressed (figs. 149–151); metafemoral tooth apical, broad; aedeagus narrowed at apex, sometimes slightly widened and spatulate at apex (fig. 156); and ventral operculum deeply divided, each lobe apically truncate (fig. 156).

DESCRIPTION: Body very broadly elongate oval, partially contractile, moderately convex (figs. 153, 154); TBL = 3.2 mm. Color of head and pronotum reddish­brown; elytra darker, except apex; metasternum conspicuously darker; mouthparts and appendages reddish­brown; antenna uniformly yellowish red­brown; coxae, femora, mesosternum, and prosternum paler, somewhat testaceous.

Head large, broad, subquadrate in shape; head narrowed behind eye, temporum short (figs. 153, 154) (about three­fifths length of eye), defined only by supraocular carina extending from posterior end of temporum to frontoclypeal suture; OHW/MDL = 1.5; OHW/PHW = about 1.0; dorsal surface with sparse, irregularly distributed, weakly impressed, small setigerous punctures, surface between with sparse, small, irregularly distributed punctules (fig. 149); eye large; frontoclypeal suture very finely impressed, nearly absent, clypeal region made distinct only by subcuticular darkened lines; surface of clypeus as on frons; without larger setae at posterolateral corners of clypeal region; labrum small, transverse, very shallowly emarginate medially; antennomere III about as long as IV + V + VI; length II:III = 1:2.2; VII very slightly wider than VIII, nearly equal in size; width VII:VIII:IX= 1:1:2.4. Pronotum broad, convex; PNW/PNL = 1.6; PNL/PNH = 1.4; PNW/PNH = 2.2; punctation similar to that of head, a little more faintly impressed (fig. 150). Elytra very wide; length little more than combined width; only slightly attenuate at apex; humeri not angulate, broadly obtuse; punctures much larger than those of head and pronotum, more coarsely impressed, irregularly distributed and moderately dense, distance between punctures about 0.5–1.5 diameter of single puncture, surface between smooth, shiny, with microscopic, nearly invisible and very sparse micropunctules and random faint curving lines (fig. 151). Mesosternum divid­ ed into anterior and posterior parts; anterior part with longitudinal median carina and fine alutaceous sculpturing; with oblique carinae and lateral, longitudinal carinae. Metasternum finely alutaceous, with laterally curved, concentric, rugose ridges, surface covered by fine setae. MSL/MTL = 0.7; MTW/MTL = 0.3.

Male tarsi 5–5–4; with pro­ and mesobasotarsomeres moderately laterally expanded; left mandible not modified; metafemur with conspicuous, broad, nearly apical tooth with small median protruding point; metasternal fovea in large anteromedial oval depression, with long fine setae (fig. 152). Aedeagus with median lobe elongate, robust, ventrally curved, recurved dorsally subapically, apical one­fifth dorsoventrally compressed and laterally narrowed before apex; apex of median lobe narrowed, slender to more or less arrowhead­shaped, with fanlike arrangement of ducts apically (fig. 156); operculum deeply bisected into two pieces, each nearly parallelsided or slightly expanded apically and truncate or very broadly rounded at apex, with apical concentration of pores (fig. 156); lateral lobes long, gradually narrowed before apex that is slightly enlarged, especially in

ventral view, with pair of subapical setae and irregularly distributed pores (figs. 155, 156); endophallic armature not extensive.

Female tarsi 5–4–4.

DISTRIBUTION: This species is nearly ubiquitous in California and occurs into western Oregon, though it does not appear to occur extensively outside California (fig. 101).

SPECIMENS EXAMINED: UNITED STATES: California: state only (3, MCZC) ; Berkeley , 7 Dec 1919 (3, MCZC) ; Poway (1, CASC) ; Carmel , 5 Jan 1924 (1, CASC) ; Willows , 13 Jun 1966, RP Allen (1, FGAC) ; Santa Barbara , 20 Dec 1980, pan trap, S Miller (4, CUIC) ; Descanso Jct , 31 Mar 1969 (1, CNCI) ; Poway , FC Bowdin (1, MCZC) ; Sequoia Natl Park , 12 May 1979, R Baranowski (7, LUND) ; Yosemite Valley , 8 Jun 1930 (1, CASC) ; San Diego (1, CASC) ; Sacramento , 7 Apr 1931, HH Keifer (1, FGAC) ; Pasadena (8, MCZC) ; Pomona (1, MCZC) ; Pasadena, Mar , A Fenyes (1, CMNH) ; Santa Cruz Mts (2, MCZC) ; Castro Valley , 27 Mar 1938, CT Sierra (1, FGAC) ; Stanford Univ. , 30 Apr 1950, P Bartholomew (1, CASC) ; Pasadena (2, CASC) ; Alameda Co.: Livermore , 11 Apr 1958, JT Doyen (1, EMEC) ; Strawberry Can. , 11 May 1984, J Whitefield (1, CASC) ; Corral Hollow , 10 mi E Livermore, 10 Mar 1976, J Doyen (1, CASC) ; 4 mi SE Livermore, Arroyo Mocho , 24 Mar 1976, FG Andrews (1, FGAC) ; Oakland , 22 Mar 1952, Quercus agrifolia, H Schuster (1, EMEC) ; Alpine Co.: 1.2 mi E Monitor Base , 25 Jun 1983, 8000′, FG Andrews (1, FGAC) ; Woodfords , 18 Oct 1962 (1, FGAC) ; Amado Co.: Pine Grove , 5 Mar 1971, FG Andrews, RF Wilkey (1, FGAC) ; Butte Co.: 4.4 mi SW Rackerby , 24 Mar 1983, A Hardy, F Andrews (1, FGAC) ; 4.4 mi SW Rackerby , 6 Feb 1980, pitfall, AR Hardy (5, FGAC) ; 3 mi E Chico , 17 Jan 1979, Bermuda grass, DS Chandler (1, QDWC) ; Colusa Co.: Rumsey , 1 May 1970, oak litter, FL Blano (1, FGAC) ; Contra Costa Co.: Mt Diablo , 24 Nov 1953 (2, EMEC) ; Antioch , 1 mi E, 24 Jan 1975, J Doyen (1, CASC) ; Vine Hill , 12 Apr 1908, FE Blaisdell (1, CASC) ; Antioch Natl Wildlife Refuge , 13 Mar 1982, pitfall traps, Powell (1, CASC) ; El Dorado Co.: Peavine Ridge Rd 7 mi SW Ice House, 16 Apr 1992, Quercus kellogi litter, FG Andrews (1, FGAC) ; 1 mi E Pacific House , 10 Jun 1994, under bark of dead oak, CB Barr (11, CASC) ; 1.3 mi S Pacific House , 15 Feb 1991, F Andrews, T Eichlin (3, FGAC) ; Blodgett Exp. For. 14 mi E Georgetown, 8 May 1976, J Doyen (1, CASC) ; 2 mi SE Latrobe , 18 Apr 1978, oak duff, Stanley, C Kuba (3, FGAC) ; Blodgett Forest 13 mi E Georgetown, 28 Apr 1976, Quercus kellogi litter, J Doyen (2, CASC) ; 3 mi W Grizzly Flat , 24 Mar 1982, under oak bark, F Andrews (1, FGAC) ; 7.7 mi SE Virner Blodgett Forest , 29 Mar 1984, under bark Pseudotsuga trifolia, FG Andrews (1, FGAC) ; 7.7 mi SE Virner Blodgett Forest , 29 Mar 1984, under bark Ponderosa pine, FG Andrews (2, FGAC) ; 3 mi SW Somerset , 26 Oct 1971, R Wharton (11, FGAC) ; 3.5 mi NW Rescue , 30 Nov 1976, Quercus litter, FG Andrews (1, FGAC) ; Fresno Co.: 8 mi W Coalinga Wartham Canyon , 16 Apr 1980, Andrews, Paddock (27, FGAC) ; Cierro Hills , 16 Apr 1980, CY Kitayama (17, CASC) ; 8 mi W Coalinga Warthan Canyon , 16 Apr 1980, flying at dusk, Andrews (12, FGAC) ; 2 mi N Tollhouse , 28 Nov 1977, Ceanothus duff, A Gilbert (3, FGAC) ; 2 mi NE Auberry , 13 Feb 1981, AJ Gilbert, HN Smith (1, FGAC) ; 9 mi E Coalinga , 20 Feb 1981, Andrews, Gilbert (5, FGAC) ; Dinky Crk , 5 Apr 1970, pine duff, TR Haig (1, FGAC) ; 8 mi W Coalinga, Warthan Canyon , 16 Apr 1980, Andrews, Paddock (1, FGAC) ; 9 mi E Coalinga , 20 Feb 1981, pitfall, Andrews, Gilbert (4, FGAC) ; 15 mi E Fresno , 16 Mar 1984, oak litter, AJ Gilbert (2, FGAC) ; 11 mi W Coalinga , 28 Oct 1971 (11, FGAC) ; Humboldt Co.: Eureka , 27 May 1981, salix litter, TR Haig (1, FGAC) ; Kern Co.: 1 mi N McKittrick , 24 Mar 1981, pitfall, RL Aalbu (2, FGAC) ; Kern River , 6 mi E Tupman, 21 Mar 1975, Atriplex litter, J Doyen (1, CASC) ; Frazier Park , 13 Jan 1961, EL Sleeper (1, FGAC) ; Arvin Hills , 9 Mar 1986, under rock, J Schuh (1, MCZC) ; 12 mi NW Taft, 9 Apr , DB Wahl (1, AMNH) ; 1.4 mi S Valley Acres , 30 Jan 1979, Andrews, Hardy (1, FGAC) ; Kings Co.: 1 mi SW Kettleman City , 4 Mar 1988, pitfall trap in sand, RL Aalbu (1, CNCI) ; 8 mi SSW Avenal , 29 Apr 1980, Andrews, Kuba, Paddock (1, FGAC) ; 8 mi SSW Avenal , 17 Sep 1980, litter at base of oak, Andrews, Kuba, Paddock (4, FGAC) ; 1 mi SW Kettleman City , 4 Mar 1988, pitfall, RL Aalbu (1, FGAC) ; 1 mi SW Kettleman City , 4 Mar 1988, pitfall trap, RL Aalbu (2, FGAC) ; Tar Canyon SE Avenal , 8 Mar 1973, pitfall, Gilbert, Bookout (1, FGAC) ; 10 mi S Avenal, Sunflower Velley , 9 Jan 1980, oak, pitfall, A Gilbert (2, FGAC) ; Tar Canyon SE Avenal , 8 Mar 1975, pitfall trap, Gilbert and Bookout (1, FGAC) ; 10 mi SW Avenal , 6 Feb 1980, oak juniper duff, A Gilbert (2, FGAC) ; 8 mi SSW Avenal , 17 Apr 1980, base of oak, Andrews, Kuba, Paddock (1, FGAC) ; Lake Co.: Butte Canyon 10 mi SE Middletown, 3 Apr 1981, Hardy, Andrews (1, FGAC) ; Lassen Co.: Ravendale , 16 Apr 1975, FG Andrews (1, FGAC) ; Los Angeles Co.: San Gabriel Canyon, 9 Feb 1961, FG Andrews (1, FGAC) ; Pine Canyon 6 mi W Lake Hughs, 27 Feb 1978, A Hardy, F Andrews (1, FGAC) ; Madera Co.: 32 mi NE Fresno USDA Exp. Station , 25 Jan 1980, woodrat nest, N Smith (1, FGAC) ; Marin Co.: Pt Reyes Station , 11 Mar 1976, oak duff, F Andrews, KS Corwin (1, FGAC) ; McClure’s Beach , 17 Dec 1976, rock crevices, upper intertidal, J Doyen (1, CASC) ; Taylor State Park , 20 Apr 1947, HB Ross (1, CASC) ; Samuel P Taylor State Park , 31 Mar 1976, FG Andrews (1, FGAC) ; Samuel P Tayler State Park , 31 Mar 1976, FG Andrews, TD Eichlin (2, FGAC) ; Novato , 22 Nov 1969, swimming pool, T Haig (2, FGAC) ; Pt Reys Mt Vinson , 25 Feb 1982, on myxomycete, FG Andrews (1, FGAC) ; Taylor State Park , 20 Jun 1947, ES Ross (1, CASC) ; Mariposa Co.: Tioga Rd , 1.9 mi E Crane Flat, 18 May 1976, 6600 ′, conifer forest litter, A Newton, M Thayer (1, MCZC) ; Mendocino Co.: 18 Mar 1938, J Helfer (1, CASC) ; 2 Jul 1926 (1, CASC) ; 29 Jul 1952 (1, EMEC) ; Hopland Field Sta. HQ Lake , 15 May 1977, J Powell (1, CASC) ; Hopland Filed Sta. HQ Lake , 15 May 1979, J Powell (1, CASC) ; Merced Co.: Los Banos Creek , 1 Apr 1987, 400′, pitfall, D Giuliani (2, FGAC) ; Modoc Co.: 5 mi NE Altura , 20 Mar 1980, oak duff, TR Haig (3, FGAC) ; 5 mi NE Altura , 20 May 1980, Juniperus occidentalis duff, TR Haig (4, FGAC) ; Mono Co.: 6 mi SW Toms Place , 8 Aug 1969, 9000 ′, A Smetana (1, CNCI) ; Monterey Co.: 19.4 mi E San Lucas , 16 Nov 1971, Heteromeles arbutifolia, FG Andrews (1, FGAC) ; 19.4 mi E San Lucas , 16 Nov 1971, oak duff, FG Andrews (1, FGAC) ; Preist Valley , 20 Mar 1981, pitfall, Gilbert, Andrews (1, FGAC) ; Arroyo Seco , 9 Feb 1982, under oak bark, F Andrews (1, FGAC) ; 17 mi E San Lucas , 20 Mar 1981, pitfall, Gilbert, Andrews (2, FGAC) ; Mt Palo Exctio , Jun 1972, oak duff, D Giuliani (1, FGAC) ; Carmel , 1915 (1, MCZC) ; 6.7 mi N Hwy 1, 23 Feb 1984, oak litter, berlese, AJ Gilbert (1, FGAC) ; Cast Rd along Sierra Ck 2.4 mi SE Bixby landing, 21 Feb 1989, F Andrews, T Eichlin (1, FGAC) ; Napa Co.: 2 mi N St Helena White’s Cave entrance, 26 Apr 1981, pitfall, R Aalbu (1, FGAC) ; 3 mi W Cordelia , 1 Oct 1971, oak duff, WW Wiard (7, FGAC) ; 2 mi ENE St Helena , 24 Jan 1981, RL Aalbu (2, FGAC) ; Nevada Co.: 4 mi S Rough and Ready , 5 May 1980, under bark of Quercus kellogi, JT Doyen (38, CASC) ; Orange Co.: Buena Park , 16 Feb 1967 (1, FGAC) ; Riverside Co.: 0.5 mi SE Aguanga Chaparral Scrub , 7 Jan 1979, pitfall, RL Aalbu (2, FGAC) ; Menifee Valley , 28 May 1980, JD Pinto (1, FGAC) ; Sacramento Co.: Sacramento , 5 Apr 1977, TR Haig (1, FGAC) ; Nimbus, Alder Creek at US Hwy 50, 2 Mar 1990, under bark, CB Barr, WD Shepard (1, CASC) ; 9 Sep 1930, blue oak (1, FGAC) ; 1 mi E Folsom Humberg Gr , 8 Sep 1978, pitfall, S Kuba (1, FGAC) ; Folsom , 1 Jan 1938, Q Tomich (1, EMEC) ; San Benito Co.: 17 mi N New Idris , 9 Apr 1982, ptifall, A Gilbert, N Smith (5, FGAC) ; 14 mi S San Benito , 16 Nov 1971 (2, FGAC) ; 1.8 mi SW Idria , 25 Mar 1981, pitfall, AJ Gilbert, N Smith (5, FGAC) ; 1.4 mi N New Idris , 8 Jun 1981, manzanita, AJ Gilbert N Smith (1, FGAC) ; 18.4 mi NE Idris , 25 Apr 1981, AJ Gilbert, HN Smith (1, FGAC) ; 8.2 mi W on Panochee Rd , 1 Jan 1979, AJ Gilbert (1, FGAC) ; San Bernardino Co.: 3 mi E Summit , 2 May 1978, KW Cooper (2, FGAC) ; 2 mi E Mentone , 15 May 1978, KW Cooper (1, FGAC) ; 6 mi W Wrightwood , 14 Apr 1986, under bark, F Andrews, TD Eichlin, A Hardy (2, FGAC) ; 1.4 mi N New Idris , 8 Jun 1981, manzanita, pitfall, AJ Gilbert, N Smith (1, FGAC) ; 3 mi E Summit , 2 May 1978, Neotoma nest, KW Cooper (1, FGAC) ; Seven Oaks , 26 Jan 1963, oak duff, FG Andrews (2, FGAC) ; 3 mi E Summit , 2 May 1978, Neotoma nest, KW Cooper (9, FGAC) ; Lake Fulmore , Nov 1967, 6000′, tullgren funnel, D Hagstrum (1, FGAC) ; San Diego Co.: Pine Hills, William Heise Co. Park, 19 Mar 1979, poison oak, berlese, KW Cooper (1, FGAC) ; 4 mi E Campo , 26 Mar 1961, EE Lindquist (2, EMEC) ; 7.5 mi E Julian, Banner Crk. , 2 May 1981, 2700′, L Herman (1, AMNH) ; Borego , 28 Apr 1955 (1, EMEC) ; San Luis Obispo Co.: Cambria , 29 Oct 1971, oak duff, FG Andrews (1, FGAC) ; Arroyo Grande , 7 Aug 1987, pitfall, J McLaughlin (1, FGAC) ; 7.5 mi W Simmler , 30 Jan 1979, pitfall, Andrews, Hardy (12, FGAC) ; Santa Barbara Co.: Carpinteria , 19 Jan 1989, J McLaughlin (1, FGAC) ; Refugio , 7 Apr 1972, D Teiman (1, FGAC) ; Santa Clara Co.: Saratoga Gap , 19 Apr 1976, FG Andrews (1, FGAC) ; 24 Apr 1923, ER Leach (1, CMNH) ; Sveadal , 27 Apr 1968, A and A Gillogly (1, FGAC) ; Santa Cruz Co.: 7.5 mi S Los Gatos, Santa Cruz Mts , 23 Nov 1963, Irwin, Westcott (1, FGAC) ; Shasta Co.: Redding , 1 Mar 1980, TR Haig (5, FGAC) ; 4 mi NE Montgomery Ck , 19 Apr 1984, oak duff, berlese, TR Haig (4, FGAC) ; Redding , 1 Apr 1979, oak duff, TR Haig (4, FGAC) ; Redding , 1 Apr 1980, TR Haig (2, FGAC) ; 4 mi NE Montgomery Ck , 9 Feb 1986, oak duff, berlese, TR Haig (1, FGAC) ; Sierra Co.: Gold Lake , 14 Jul 1921 (1, CASC) ; Slano Co.: Travis AFB, 6 Apr 1971 (3, FGAC) ; Sonoma Co.: Trinity , 30 Nov 1933 (1, CASC) ; Armstrong Park , 14 Mar 1954, J Helfer (1, EMEC) ; Sta Clara Co.: Guadalupe Res , 1 Apr 1973, under bark blue oak, CY Kitayama (5, CASC) ; Sta Cruz Co.: S end Sunset Bch , 4 May 1973, J Doyen (1, CASC) ; Stanislaus Co.: Knights Ferry , 25 Mar 1987, meadow, Andrews, Eichlin, Hard (1, FGAC) ; La Grange , Oct 1975, RP Allen (1, FGAC) ; La Grange , 5 Oct 1970, at light, RP Allen (11, FGAC) ; Tehama Co.: 1 mi SW Dales , 13 Feb 1971, DS Chandler (5, CASC) ; 15 mi W Red Bluff , 14 Feb 1971, DS Chandler (3, CASC) ; Red Bluff , 16 Feb 1973, TR Haig (1, FGAC) ; S Fk Battle Creek , 25 Dec 1970, oak lit­ ter, DS Chandler (3, CASC) ; 2 mi NE Dales , 13 Feb 1971, DS Chandler (2, EMEC) ; Trinity Co.: Junction City , 12 Mar 1981, TR Haig (1, FGAC) ; Junction City , 04 Jan 1990, madrone duff, TR Haig (1, FGAC) ; Weaverville , 15 Apr 1981, litter, TR Haig (1, FGAC) ; Del Loma , 5 Jan 1985, Alnus duff, TR Haig (10, FGAC) ; Del Loma , 23 Jan 1980, TR Haig (10, FGAC) ; Big Bar , 3 Aug 1976, leaf litter, FG Andrews (1, FGAC) ; 5 mi W Coffee Ck Ranch , 19 Jun 1972, pine duff, DS Chandler, DP Levin (2, CASC) ; Tulare Co.: Ash Mt nr Three Rivers , 11 Feb 1983, flume debris trap, R Haines (1, FGAC) ; Ash Mt , 17 Mar 1984, at flume forebay, R Haines (22, FGAC) ; Wood L., 19 Dec 1947, rotary trap, NW Frazier (4, EMEC) ; Kaweah , 16 Nov 1971, oak duff, FG Andrews (1, FGAC) ; 1 Jun 1933, FT Scott (5, FGAC) ; Ashe Mt HQ, 28 Apr 1979, Douglas fir litter, JT Doyen (9, CASC) ; Camp Wishon 10 mi NE Springville, 1 May 1979, J Doyen (1, CASC) ; S FK Camp 13 mi SE Three Rivers , 29 Apr 1979, Arctostaphylos litter, JT Doyen (1, CASC) ; 10 mi SE Three Rivers , S FK Kaweah R., 29 Apr 1979, Quercus litter, JT Doyen (1, CASC) ; Sequoia Nat. Pk. Huckleberry Meadow, 28 May 1984, 6000′, R Baranowski (1, LUND) ; Tuolumne Co.: Strawberry , 23 Jun 1953, JG Rozen (1, EMEC) ; Ventura Co.: Moorpark , 23 Mar 1971, oats and barley, Anderson, Johnson (1, FGAC) ; Yolo Co.: 2 mi W Rumsey, Cache Crk , 8 Nov 1971, Bermuda grass, sweeping, DS Chandler (2, CASC) ; Davis , 15 Nov 1956, Pyrecantha litter, RO Shuster (4, CASC) ; Yuba Co.: Sierra Foothill Feild Sta 6 mi N Smartville, 4 May 1980, under bark Pinus sabiniana, JA Powell (4, CASC) ; Dry Crk 4 mi NW Smartville, 3 May 1980, under bark Pinus sabiniana, JT Doyen (3, CASC) ; 15 mi E Marysville , 6 Feb 1980, pitfall, AR Hardy (16, FGAC) ; 19 Apr 1913 (1, MCZC) ; Spencerville Wildlife Area, 9 Apr 1978, F Andrews (7, FGAC). Oregon : Clatsop Co.: 5 mi N 7 mi W Elsie, 13 Aug 1973, 700′, duff, E Benedict (1, QDWC) ; Douglas Co.: 16 km E Steamboat , 16 Oct 1972, 3200 ′, duff, E Benedict (1, QDWC) ; 5.5 mi NE Idleyld Pk. , 1 Apr 1972, 1200 ′, duff, E Benedict (1, QDWC) ; Jackson Co.: 4 mi E Eagle Point, Kanutcham Ck , 22 Jan 1972, 1400′, madrone oak duff, E Benedict (2, QDWC) ; 9 mi N Central Point , 22 Jan 1972, 1400′, oak duff, E Benedict (4, QDWC) ; Josephine Co.: 1 mi S, 5 mi W O’Brien, Hwy 99, 18 Dec 1971, 1400′, debris, E Benedict (1, PECK) ; Linn Co.: Swamp Mt Rd 2 mi E Cascadia, 29 Apr 1972, maple duff and wood, EM Benedict (1, PECK) .

DISCUSSION: The holotype of A. hebetatum Hatch (USNM) was examined and found to be a synonym of A. virile based on similarity

in punctation and shape of the median lobe of the aedeagus. This specimen is well within the variation of typical members of A. virile .

Agathidium virile is a common and morphologically variable species in the far western region of the United States. Specimens assigned to this species are diverse in punctation and, to a certain extent, in the form of the aedeagus. The elytra are densely punctate, but the punctures vary from rather feebly impressed to very deep and coarsely impressed. Among those with coarse elytral punctation, punctures of the head and especially the pronotum may be somewhat small­ er but similarly strikingly coarse and deep ranging to rather small and decidedly weakly impressed. The apex of the median lobe varies from slender and gradually narrowed to more broadly expanded. The branches of the operculum vary from very slender and elongate to distinctly broader, but generally the branches are narrowed medially with the apices broadly expanded and truncate.

It is possible that additional species exists within what we regard as A. virile . However, no correlations were found among these variable traits that suggest a reasonable inference of species status for any of the included variable specimens, and we regard the species as simply polymorphic. A male from Stanford, California (CASC), has a relatively broad median lobe apex, gradually broadened and apically spatulate ventral opercula, in combination with rather weakly impressed pronotal and elytral punctation. In this sense it is somewhat intermediate in condition with A. angustoperculum . Considering the known variation within A. virile , however, this specimen is most reasonably assigned here for the present. Another specimen from Santa Clara County, California (CNMH), and one from Linn County, Oregon (PECK), are tentatively assigned to this species and have the apical portion of the median lobe much more slen­ der than in most specimens, with the lateral margins nearly parallel.

A specimen from McClure’s Beach, Marine County, California (CASC), was collected from ‘‘rock crevices, upper intertidal’’, though this must certainly be an accidental occurrence for this species.

Figures

Figs. 101–103. Geographic distribution of Agathidium revolvens group species: 101, A. virile Fall;

Figs. 149–154. Agathidium virile Fall: Figs. 149–151, punctation detail: 149, head; 150, pronotum; 151, elytron. 152. Male metasternal fovea. Figs. 153, 154, habitus: 153, dorsal; 154, lateral. Bars = 0.5 mm.

Figs. 155, 156. Agathidium virile Fall aedeagus: 155, lateral; 156, ventral.