Ayacucho inermis (Roewer, 1957) Benedetti & Pinto-da-Rocha, 2022

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

publication ID


publication LSID


persistent identifier


treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Ayacucho inermis (Roewer, 1957)

comb. nov.

3.13. Ayacucho inermis (Roewer, 1957) comb. nov.

Figs 2C View Figure 2 , 7E, F View Figure 7 , 12C, D View Figure 12 , 28 View Figure 28

Pinocchio inermis Roewer, 1957: 70 (desc.), fig. 3 (male coxa-patella IV).

Palcares inermis : Kury 2003: 145 (cat., syst.).


MALE: Measurements (n =1) DSW: 2.8; DSL: 3.2; CL: 1.0. FIVL: 2.0. ChL: 1.2. Coloration (in ethanol): Uniformly yellowish. In the original description: rusty-yellow body, dorsally darker than ventrally; legs rusty-yellow, slightly blackish. Dorsum: (Fig. 2C View Figure 2 ) Alpha-type DSS, with long coda. Anterior margin densely covered with granules, with median elevation. Ocularium with a pair of spines; densely granulate. Carapace densely granulate at lateral and posterior regions. Areas I-IV densely covered with granules; I unarmed; II-IV with two pairs of small paramedian tubercles, slightly larger than the granules. Posterior margin of dorsal scutum with a row of acuminate and scattered tiny tubercles. Free tergites I-III with a row of large acuminate tubercles, much larger than tubercles of the areas of dorsal scutum and interspersed with small tubercles. Lateral margins of dorsal scutum densely covered with granules. Chelicerae: (Fig. 2C View Figure 2 ) Similar to the female. Segment I densely granulate. Segment II with granules with a much lower density than on segment I; four teeth on finger. Segment III with two teeth. Pedipalpus : Trochanter with a ventroapical tubercle. Femur with a ventrobasal tubercle; a row of three small ventral tubercles, smaller than the ventrobasal tubercle. Tibia: prolateral IIi, retrolateral iIi. Tarsus: prolateral IIi, retrolateral iIii. Venter: Coxa I with a middle row of four small tubercles. Coxae II-IV densely covered with granules. The area between coxae II-III and III-IV with a small tubercle at the apex. Smooth genital area. Free sternites with a row of small granules. Anal operculum covered with granules. Legs: (Figs 2C View Figure 2 , 7E, F View Figure 7 ) Coxae I-II with a prolateral apophyses; III unarmed; IV densely granulate throughout its length, with an apical spiniform apophysis. Trochanters I-IV granulate; IV with a retroapical acuminate tubercle. Femora I-III with granules scattered throughout their extension; IV densely granulate; a retroventral row of 15 large acuminate tubercles throughout its length except at the base and apex; a proventral row of 10-12 large acuminate tubercles (tiny at base) throughout its length, with variation in number of tubercles between right and left legs in the same specimen. Patellae I-IV densely granulate. Tibiae I-IV densely granulate. Tarsal segmentation (n =1) 3, 8, 5-6, 6. Penis: (Fig. 12C, D View Figure 12 ) VP subrectangular with sides diverging towards the apex; distal margin straight; slightly sinuous on lateral view. MS C1-C3 subapical short and straight; MS A1 short and straight; placed more dorsally than MS C. Lateral sacs long, robust and apically blunt; with short T3-like microsetae. Stylus slightly thick, with long apex and small apical projections. Dorsal process present. - FEMALE: Measurements (n =1) DSW: 3.2; DSL: 3.6; CL: 1.2. FIVL: 2.3. ChL: 1.5. Female very similar to male. Chelicerae of similar size. Femur IV with a retroventral and a proventral rows of eight and 11 tubercles respectively, being much smaller than the tubercles in males and tubercles in proventral row larger than that of the retroventral row. Tarsal segmentation: (n =1) 5, 8, 6, 6.


Resembles Ayacucho spiniger comb. nov. because two ventral rows of tubercles in femur IV (Fig. 7F View Figure 7 ). Differs from A. spiniger comb. nov. because femur IV with spiniform tubercles (Fig. 7F View Figure 7 ); ventral plate of penis without lateral projections; VP with short macrosetae; lateral sacs with short microsetae; dorsal process shorter than the stylus (Fig. 12C, D View Figure 12 ).


(Fig. 28 View Figure 28 ) PERU. La Libertad. Huamachuco.

Material examined.

Type material: Holotype ♂, ' PERU, La Libertad, Huamachuco , 3,200m a.s.l. without date, Weyrauch leg. (SMF RII 11393 View Materials /23) - Paratype ♀, ‘ditto’ (SMF RII 11393 View Materials /23) .














Ayacucho inermis (Roewer, 1957)

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo 2022

Pinocchio inermis

Benedetti & Pinto-da-Rocha 2022

Palcares inermis

Benedetti & Pinto-da-Rocha 2022