Maechidius paupianus Heller, 1910

Telnov, Dmitry, 2020, A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line, European Journal of Taxonomy 721, pp. 1-210 : 90-93

publication ID	https://doi.org/ 10.5852/ejt.2020.721.1127
publication LSID	lsid:zoobank.org:pub:89E62EF8-2E45-4C59-94B7-6A5603E8939B
DOI	https://doi.org/10.5281/zenodo.4344253
persistent identifier	https://treatment.plazi.org/id/1A3787F6-9D45-9207-FDD0-F8DBCBE42911
treatment provided by	Valdenar
scientific name	Maechidius paupianus Heller, 1910
status

Treatment

Maechidius paupianus Heller, 1910 View in CoL

Figs 67–68 View Figs 66–69 , 158–159 View Figs 157–168 , 242 View Figs 242–258 , 274 View Figs 266–277 , 291 View Figs 286–301 , 405, 504, 536, 748–750

Maechidius arrowi Frey, 1969: 501 View in CoL syn. nov.

Type material

Lectotype of M. paupianus (herewith designated)

PAPUA NEW GUINEA • ♀; “ Kais.Wilhelmsland Paup Dr.Schlaginhaufen [p, blue label]// 1910 [p] 1 [h] [blue label]// [upper side of label] typus [p] [underside of label] paupianus Heller [h, red label]// Staatl. Museum für Tierkunde.Dresden [p]”; SNSD .

Holotype of Maechidius arrowi

PAPUA NEW GUINEA • ♂; “ Holotype [p, label circular, red frame] // Brit.Mus. 1957-693. [p] // TERR. PAPUA & NEW GUINEA : Maprik. 28.x.1957. J.Smart. [p]// TYPE [p, red label]// TYPE Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH.

Paratypes of Maechidius arrowi

PAPUA NEW GUINEA • 1 spec.; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA : Maprik. 16.x.1957. J.Smart. [p] // Brit.Mus. 1957-693. [p] // PARATYPE [p, red label] //A. TYPE Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 2 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 16.x.1957. J.Smart. [p] // Brit.Mus. 1957-693. [p] //PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 3 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 17.x.1957. J.Smart. [p]// Brit.Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 2 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 18.x.1957. J.Smart. [p] // Brit.Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 2 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 24.x.1957. J.Smart. [p]// Brit. Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 4 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 26.x.1957. J.Smart. [p]// Brit.Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 1 spec.; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 28.x.1957. J.Smart. [p] // Brit.Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 2 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 29.x.1957. J.Smart. [p]// Brit. Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 3 specs; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 16.x.1957. J.Smart. [p]// Brit.Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 1 spec.; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 16.x.1957. J.Smart. [p] // Brit.Mus. 1957-693. [p] // PARATYPE [p, red label] // Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH • 1 spec.; “ Paratype [p, label circular, red frame] // TERR. PAPUA & NEW GUINEA: Maprik. 26.x.1957. J.Smart. [p]// Brit. Mus. 1957-693. [p] // PARATYPE [p, red label]// Maechidius arrowi n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH .

New material

INDONESIA • 1 ♀; “INDONESIA, Papua, Jayapura Distr.; 5 km NE of Sentani S slopes of Cyclops Mts. , Gn. Ifar 02°32.8’S, 140°33.2E, 315 m J.Hájek & J.Šumpich leg., 30.I.2015 // coll.general National Museum Prague, Czech Republic”; NMPC GoogleMaps .

PAPUA NEW GUINEA • 2 specs; “ PAPUA: Kokoda. 1,200 ft. vi.1933. L.E.Cheesman. B.M.1933- 456.”; BMNH • 1 spec.; “at light// PAPUA: Kokoda. 1,200 ft. vi.1933. L.E.Cheesman. B.M.1933-456.”; BMNH • 8 specs; “ PAPUA: Kokoda. 1,200 ft. vii.1933. L.E.Cheesman. B.M.1933-492.”; BMNH • 50 specs; “ PAPUA: Kokoda. 1,200ft. viii.1933. L.E.Cheesman. B.M.1933-577.”; BMNH • 2 specs; “ PAPUA: Kokoda. 1,200ft. ix.1933. L.E.Cheesman. B.M.1933-577.”; BMNH • 1 spec.; “ PAPUA: Mafulu. 4,000 ft. xii.1933. L.E.Cheesman. B.M.1934-321.”; BMNH • 1 ♂; “ PAPUA: Mafulu. 4,000 ft. i.1934. L.E.Cheesman. B.M.1934-321. L.E.Cheesman. B.M.1933-577. // ♂ // Maechidius paupianus, Hell. Determined from description. G.J.A.”; BMNH • 1 ♀; “Str. No. 46. [p] // NEW GUINEA: Madang Dist. , Finisterre Mts. Damanti 3,550 ft. 2-11.x.1964 [p]// M.E.Bacchus. B.M.1965-120 [p] //M2) prope M. arrowi [h] det. G.Frey,1967/68 [p]”; BMNH • 1 ♀; “NEUGUINEA Wau 6.6.1965 PYKA leg.”; SMNS • 1 ♀; “NEUGUINEA Boana , 1000 m 22.11.1965 H. Pyka leg.”; SMNS .

Remarks

Heller (1910: 23) likely based his description of M. paupianus on a single specimen – although not explicitly stated. The single SNSD female specimen labelled “typus” is herewith designated as lectotype in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other syntypes are discovered in the future.

Description

MEASUREMENTS. Total body length 5.90 (lectotype M. paupianus ) to 7.50 mm.

Dorsum uniformly brown, venter and appendages somewhat paler. Head flattened dorsally, glossy dorsally and ventrally. Male labroclypeus ( Fig. 159 View Figs 157–168 ) broadly shallowly emarginate on anterior margin, anterolateral angles obtuse angulate, protruding anteriad. Female labroclypeus ( Fig. 158 View Figs 157–168 ) very broadly emarginate on anterior margin. Lateral margins of labroclypeus sinuous in both dorsal and lateral views. Anterolateral angles raised up at angle of nearly 90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Underside of labroclypeus with sparse long delicate setae along anterior and lateral margins. Canthus strongly obtusely angulate to broadly rounded in dorsal view. Punctures of frons circular to ovoid, annular, large and shallow. Intervening spaces smaller than punctures, glossy to microreticulate. Setae inconspicuous, suberect, each seta rises from anterior margin of corresponding puncture, hardly surpassing length of corresponding puncture to twice as long as it.Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, subopaque dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum gradually widened anteriorly or broadly rounded, very obtusely angulate postmedially, from here constricted towards base or shallowly emarginate, distinctly crenulate all along (crenulae small but with well-defined intervals, not merged) ( Fig. 242 View Figs 242–258 ). Lateral margin of pronotum sinuous in lateral view. Short suberect seta present between every two crenulae. Hypomeron separated from prosternum by low nearly straight carina, with very long setae on its slightly arched anterolateral margin opposite to compound eye ( Fig. 274 View Figs 266–277 ). Antennal pocket shallow. Pronotal punctures larger than those on head, ovoid, annular, rather dense, intervening spaces variably large, opaque and microreticulate. Inconspicuous curved appressed to suberect seta rises from anterior margin of each puncture, generally surpassing its length. Basal margin near posterolateral angles with a fringe of long setae on either side in some specimens. Scutellar shield narrowly rounded apically. Elytron opaque, without tracks of longitudinal carinae. Elytral punctures linear (elongate and narrow), incision-shaped, moderately deep ( Fig. 291 View Figs 286–301 ). Intervening spaces microreticulate. Seta rises from anterior margin of each puncture, suberect, not surpassing length of corresponding incision. Male and female pygidium flattened dorsally (female one with inconspicuous V-shaped dorsal impression), with large shallow annular punctures ( Fig. 504 View Figs 504–521 ). Intervening spaces microreticulate, generally smaller than punctures. Setae of pygidium sparse, suberect, each seta rises from centre of puncture. Male and female protibia widened distally, with delicate dorsal carina and three external teeth (two distal significantly larger, lower tooth inconspicuous in some specimens) (Fig. 405). Male protibial terminal spur short and thickened, female one narrow and pointed. Female metatibial terminal spurs long, unequal in length, distally rounded. Tarsal claws with pulvilli. Spiculum gastrale as in Fig. 536. Aedeagus as in Figs 748–750 View Figs 739–750 .

Sexual dimorphism

Female lamellae of antennal club shorter; labroclypeus comparatively shallower emarginate anteriorly; female pygidium with vague V-shaped impression in apical part.

SNSD	SNSD
BMNH	United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]
SMNS	Germany, Stuttgart, Staatliches Museum fuer Naturkunde
NEW	University of Newcastle
NMPC	National Museum Prague
SMNS	Staatliches Museum fuer Naturkund Stuttgart

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Coleoptera
Family	Scarabaeidae
SubFamily	Melolonthinae
Tribe	Maechidiini
Genus	Maechidius