Maechidius esau Heller, 1914

Telnov, Dmitry, 2020, A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line, European Journal of Taxonomy 721, pp. 1-210 : 45-46

publication ID

https://doi.org/ 10.5852/ejt.2020.721.1127

publication LSID

lsid:zoobank.org:pub:89E62EF8-2E45-4C59-94B7-6A5603E8939B

DOI

https://doi.org/10.5281/zenodo.4344389

persistent identifier

https://treatment.plazi.org/id/1A3787F6-9D30-9276-FDF4-FCA5C9E82F8A

treatment provided by

Valdenar

scientific name

Maechidius esau Heller, 1914
status

 

Maechidius esau Heller, 1914 View in CoL

Figs 2–3 View Figs 2–7 , 36–37 View Figs 35–37 , 118–119 View Figs 109–120 , 213 View Figs 208–225 , 303 View Figs 302–316 , 376, 470, 561, 614–616

Maechidius setosellus Frey, 1969: 500 View in CoL syn. nov.

Maechidius setosus Moser, 1920: 17 View in CoL syn. nov.

Type material

Lectotype of M. esau (herewith designated)

PAPUA NEW GUINEA • ♂; “ N.Guinea [p, blue label]// Gehr.W.Müller Vermächt. 1909 [p, blue label]// Typus! [p] esau Heller [h, red label] // Staatl. Museum für Tierkunde.Dresden [p]”; SNSD .

Lectotype of M. setosus (herewith designated)

PAPUA NEW GUINEA • ♂; “ Maechidius setosus Mos. Type [h] // D. Neu-Guinea [p] Stephansort// SYNTYPE Maechidius setosus Moser, 1920 labelled by MFNB 2019 View Materials [p, red label]”; ZMHB .

Holotype of M. setosellus

PAPUA NEW GUINEA • ♂; “ Holotype [p, label circular, red frame] // Stn. No. 28. [p] // NEW GUINEA: Madang Dist. R. Buru, c. 90 m. N.W.Lae 1200 ft. 1.x.1964 [p]// Type Maechidius setosellus n. sp. [h] det. G.Frey,1967//86 [p]// M.E.Bacchus. B.M. 1965-120. [p] // TYPUS [p, red label]”; BMNH.

Paratype of M. setosellus

PAPUA NEW GUINEA • ♂; “ Paratype [p, label circular, yellow border] // NEW GUINEA: Madang Dist. R. Buru, c. 90 m. N.W.Lae 1200 ft. 1.x.1964 [p] // M.E.Bacchus. B.M. 1965-120. [p] // PARATYPE [p, red label]// Maechidius setosellus n. sp. [h] det. G.Frey,1967/68 [p]”; BMNH .

New material

PAPUA NEW GUINEA • 2 specs; “ TERR. PAPUA & NEW GUINEA : Maprik. 17.x.1957. J.Smart // Brit.Mus.1957-693.”; BMNH 2 specs; “ TERR. PAPUA & NEW GUINEA : Maprik. 18.x.1957. J.Smart // Brit.Mus.1957-693.”; BMNH 1 spec.; “ TERR. PAPUA & NEW GUINEA : Maprik. 19.x.1957. J.Smart // Brit.Mus.1957-693.”; BMNH 1 spec.; “ TERR. PAPUA & NEW GUINEA : Maprik. 24.x.1957. J.Smart // Brit.Mus.1957-693.”; BMNH 5 specs; “ TERR. PAPUA & NEW GUINEA : Maprik. 26.x.1957. J.Smart // Brit.Mus.1957-693.”; BMNH 5 specs; “ TERR. PAPUA & NEW GUINEA : Maprik. 28.x.1957. J.Smart // Brit.Mus.1957-693.”; BMNH 22 specs; “Stn. No. 28.// NEW GUINEA : Madang Dist., R. Buru , c. 90 m. N.W.Lae 1200 ft. I. x.1964 // M.E. Bacchus. B.M. 1965-120.”; BMNH 1 ♀; “ Papua New Guinea Wau 1500 m Aug.1983 M. Mastaller leg.”; SMNS .

Remarks

Heller (1914: 627) likely based his description of M. esau on a single specimen, although not explicitly stated. The single SNSD specimen labelled “Typus♂” is herewith designated as lectotype in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other syntypes are discovered in the future. Moser (1920: 17) based his description of M. setosus on an unstated number of specimens. The single ZMHB syntype is therefore designated as lectotype. The single SNSD specimen labelled “Typus♂” is herewith designated as lectotype in order to enhance the stability of nomenclature and fix the specimen I based my redescription and new synonymy on, in case other syntypes are discovered in the future. Frey (1969: 500) described M. setosellus from two specimens captured near locality that M. setosus was described from.

Description

MEASUREMENTS. Total body length 8.70 (lectotype M. esau ) to 10.10 mm (lectotype M. setosus ).

Dorsum and venter uniformly black or brown. Head flattened dorsally, glossy to subopaque dorsally and ventrally. Male and female labroclypeus very broadly V-shaped emarginate anteriorly ( Figs 36– 37 View Figs 35–37 ), its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles broadly rounded, protruding, raised up at angle of nearly 90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Punctures of frons irregularly circular, moderately large to large, rather deep, their backgrounds shagreened. Intervening spaces microreticulate, variably large. Setae long, suberect. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy to subopaque dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum slightly gradually widened in anterior half, slightly constricted postmedially, nearly straight at base, crenulate all along ( Fig. 213 View Figs 208–225 ). Extraordinarily long erect seta present between every two crenulae, at least as long as longitudinal diameter of compound eye in fresh specimens. Lateral margin of pronotum sinuous in lateral view. Punctures of pronotal disc generally sparser than those on head, circular to annular (in basal third of disc), intervening spaces glossy, generally larger than punctures. Hypomeron separated from prosternum by low straight carina, with moderately long setae on anterolateral margin opposite to compound eye ( Figs 2–3 View Figs 2–7 ). Pronotal setation as on head, stronger appressed. Scutellar shield rounded apically. Elytron glossy to subopaque, with vague tracks of longitudinal carinae. Punctures of elytral disc double ( Fig. 303 View Figs 302–316 ). Ordinary punctures of disc shallow and rather dense, irregular in size and shape. Each ordinary puncture supplemented with a shallow linear (narrow and long) incision-shaped puncture. Intervening spaces in part microreticulate, in part glossy, generally larger than ordinary punctures. Elytral setae moderately long, erect to suberect. Some setae longer and stronger erect, arranged in poorly defined longitudinal rows. Lateral margin of elytra with extraordinarily long erect setae. Male and female pygidium slightly convex dorsally, with large shallow annular punctures ( Fig. 470 View Figs 469–486 ). Intervening spaces glossy, variably large. Setae of pygidium long, erect to suberect; each seta rises from either anterior margin or centre of corresponding puncture. Protibia widened distally, dorsally with delicate longitudinal carina, with two large acute distal external teeth and much smaller basal one (Fig. 376). Male protibial terminal spur not present. Male metatibial terminal spurs long, nearly equally long. Distal margin of metatibia in both sexes with projection over basal metatarsomere. Tarsal claws with pulvilli. Spiculum gastrale as in Fig. 561 View Figs 545–569 . Aedeagus as in Figs 614–616 View Figs 601–616 .

Sexual dimorphism

Female lamellae of antennal club distinctly shorter; female labroclypeus comparatively shallower emarginate anteriorly.

SNSD

SNSD

ZMHB

Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet

BMNH

United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]

SMNS

Germany, Stuttgart, Staatliches Museum fuer Naturkunde

NEW

University of Newcastle

R

Departamento de Geologia, Universidad de Chile

SMNS

Staatliches Museum fuer Naturkund Stuttgart

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Maechidius

Loc

Maechidius esau Heller, 1914

Telnov, Dmitry 2020
2020
Loc

Maechidius setosellus

Frey G. 1969: 500
1969
Loc

Maechidius setosus

Moser J. 1920: 17
1920
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