Lepanus globulus ( Macleay, 1887 ),

Gunter, Nicole L. & Weir, Thomas A., 2021, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species, Zootaxa 4923 (1), pp. 1-66: 21-28

publication ID

https://doi.org/10.11646/zootaxa.4923.1.1

publication LSID

lsid:zoobank.org:pub:41D057A7-F855-47C2-A598-8000CFD83A5B

DOI

http://doi.org/10.5281/zenodo.4537927

persistent identifier

http://treatment.plazi.org/id/1A323678-623A-2207-09D9-F9E6FD796E49

treatment provided by

Plazi

scientific name

Lepanus globulus ( Macleay, 1887 )
status

 

Lepanus globulus ( Macleay, 1887) 

( Figs. 2C, HView FIGURE 2, K–L; 3C; 4C; 5C; 7A–B)

Epilissus globulus Macleay, 1887: 222  ; Gillet 1911: 38; Lea 1923: 353; Paulian 1934: 282.

Lepanus globulus: Matthews 1974: 106  .

Type material examined. Lectotype: ♂ “Cairns N Queensl” / “ LECTOTYPE Epilissus globulus Macleay  ” [RED] / “( LEFT) Lepanus globulus (Macl.) DET 1971 E. G Matthews  LECTOTYPE ” / “specimen of Lepanus nitidus Math.  on same card removed T. Weir 1977” / “On permanent loan from MACLEAY MUSEUM University of Sydney” / “ Epilissus globulus Macl. Cairns  ” / “ ANIC Database 25 057888” in ANIC.

Other material examined. 1078 other specimens from over 150 unique localities were also examined.

Queensland: Shiptons Flat [15°47’00”S, 145°14’00”E], 16–18.v.1981, A. Calder & J.E. Feehan (25-058135 2GoogleMaps  specimens in ANIC); Shiptons Flat [15°47’00”S, 145°14’00”E], 17–19.x.1980, T.A. Weir (25-058136 3 specimens in ANIC)GoogleMaps  ; Shiptons Flat [15°47’33”S, 145°13’51”E], 6.xii.1990 – 19.i.1991 ( T44598View Materials T44600View Materials 3 specimens in QM)GoogleMaps  ; Shiptons Flat, via Helensvale [15°47’33”S, 145°13’51”E], 20–27.vii.1974 ( T49771View Materials T49779View Materials 9 specimens in QM)GoogleMaps  ; Bloomfield Road, via Helensvale [15°47’45”S, 145°18’17”E], 2.v.1974 ( T49765View Materials 1 specimen in QM)GoogleMaps  ; Bloomfield Road, via Helensvale [15°47’45”S, 145°18’17”E], 20–27.vii.1974 ( T49766View Materials T49770View Materials 5 specimens in QM)GoogleMaps  ; Mount Hartley , 2.5 km SW, 15°47’49”S, 145°19’34”E, 1.i.1994 – 5.iii.1994 ( T44672View Materials T44673View Materials 2 specimens in QM)GoogleMaps  ; Shiptons Flat via Helensvale [15°48’00”S, 145°15’00”E], 240 m, 20–27.vii.1974, G.B. Monteith & D. Cook (25-015662 3 specimens in QM)GoogleMaps  ; Shiptons Flat via Helensvale, ABRS Site 1 [15°48’00”S, 145°15’00”E], 198 m, 20–27.vii.1974, G.B. Monteith & D. Cook (25-015663 6 specimens in QMGoogleMaps  ; Bloomfield Road via Helensvale, ABRS Site 3 [15°48’00”S, 145°19’00”E], 168 m, 2.v.1974, G.B. Monteith (25-015660 1 specimen in QM)GoogleMaps  ; Bloomfield Road via Helensvale, ABRS Site 3 [15°48’00”S, 145°19’00”E], 168 m, 20–27.vii.1974, G.B. Monteith & D. Cook (25- 015661 5 specimens in QM)GoogleMaps  ; Donovan Creek , 16°01’13”S, 145°26’38”E, 14.i–10.ii.1998, S. Grove ( T73176View Materials T73206View Materials 31 specimens in QM)GoogleMaps  ; Donovan Creek , 16°01’13”S, 145°26’38”E, 10.ii–14.iii.1998, S. Grove ( T85552View Materials T85572View Materials 21 specimens in QM)GoogleMaps  ; Donovan Creek , 16°01’13”S, 145°26’38”E, 14.iii–6.v.1998, S. Grove ( T88852View Materials T88854, T88859, T88863, T88866 T88870View Materials 10 specimens in QM)GoogleMaps  ; Donovan Creek , 16°01’13”S, 145°26’38”E, 10.ix–26.x.1998, S. Grove ( T88855View Materials T88858, T88860 T88862, T88864 T88865View Materials 9 specimens in QM)GoogleMaps  ; Cape Tribulation area [16°04’00”S 145°28’00”E], 21–28.iii.1984, A. Calder & T.A. Weir (25-058141 2 specimens in ANIC)GoogleMaps  ; Cape Tribulation [16°05’00”S, 145°27’00”E], 20 m, 9.v.1997, C. Reid (25-058184 1 specimen in ANIC)GoogleMaps  ; Cape Tribulation transect Site 5A, 16°04’41”S, 145°27’0”E, 5–9.i.1983 ( T55573View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 5 [16°04’41”S, 145°27’13”E], 23.ix–7.x.1982 ( T44754View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 5 [16°04’41”S, 145°27’13”E], 20–23.iv.1983 ( T44755View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 4 [16°04’45”S, 145°27’20”E], 5–9.i.1983 ( T44752View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 4 [16°04’45”S, 145°27’20”E], 20–23.iv.1983 ( T44753View Materials 1 specimen in QM)GoogleMaps  ; Pilgrim Sands [16°04’S, 145°28’E], 19.xi–10.xii.1993 ( T44741View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 1 [16°04’12”S, 145°27’54”E], 23.iv.1983 ( T44751View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 1 [16°04’12”S, 145°27’54”E], 20–23.iv.1983 ( T44756View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 1 [16°04’12”S, 145°27’54”E], 23.ix–7.x.1982 ( T55608View Materials T55609View Materials 2 specimens in QM)GoogleMaps  ; Cape Tribulation [16°05’00”S, 145°28’00”E], 10 m, 9.v.1997, C. Reid (25-058183 9 specimens in ANIC)GoogleMaps  ; Cape Tribulation , 40 km N of Daintree [16°05’00”S, 145°29’00”E], 10 m, 12.vii.1982, S. & J. Peck (25-058131 2 specimens in ANIC)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 10.i–12.ii.1998, S. Grove ( T73221View Materials T73225View Materials 5 specimens in QM)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 19.xii.1998 – 16.i.1999, S. Grove ( T85537View Materials 1 specimen in QM)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 12.ii–15.iii.1998, S. Grove, ( T85555, T85542 2 specimens in QM)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 15.iii–7.v.1998, S. Grove ( T88883, T88886 2 specimens in QM)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 11.ix–25.x.1998, S. Grove ( T88884View Materials –88885, T88887View Materials T88888View Materials 4 specimens in QM)GoogleMaps  ; Noah Creek , 7 km ENE Thornton Peak [16°08’00”S 145°26’00”E], 27.iii.1984, A. Calder & T.A. Weir (25-058142 1 specimen in ANIC)GoogleMaps  ; Oliver Creek (= Arsenic Creek ) [16°08’22”S, 145°26’24”E], 5–9.i.1983 ( T55613View Materials 1 specimen in QM)GoogleMaps  ; Noah Beach , 16°09’04”S, 145°26’27”E, 9.ix–10.x.1998, S. Grove ( T73226View Materials T73229View Materials 4 specimens in QM)GoogleMaps  ; Noah Beach , 16°09’04”S, 145°26’27”E, 26.xii.1997 – 7.i.1998 ( T73230View Materials T85548, T85571 20 specimens in QM)GoogleMaps  ; Noah Beach , 16°09’04”S, 145°26’27”E, 9.ii–15.iii.1998, ( T85550View Materials T85567, T85543, T88878 20 specimens in QM)GoogleMaps  ; Noah Beach , 16°09’04”S, 145°26’27”E, 15.iii–7.v.1998 ( T88873View Materials T88874, T88879 3 specimens in QM)GoogleMaps  ; Noah Beach , 16°09’04”S, 145°26’27”E, 9.ix–22.x.1998 ( T88875View Materials 1 specimen in QM)GoogleMaps  ; Noah Beach , 16°09’04”S, 145°26’27”E, collected 15.iii–7.v.1998 ( T88876View Materials T88877View Materials 2 specimens in QM)GoogleMaps  ; Cooper Creek , 16°09’10”S, 145°24’19”E, 11.i–10.ii.1998 ( T73172View Materials T73175View Materials 4 specimens in QM)GoogleMaps  ; Cooper Creek , 16°09’10”S, 145°24’19”E, 10.ii–16.iii.1998 ( T85551View Materials T85562View Materials 12 specimens in QM)GoogleMaps  ; Cooper Creek , 16°09’10”S, 145°24’19”E 16.iii–7.v.1998 ( T88880View Materials 1 specimen in QM)GoogleMaps  ; Cooper Creek , 16°09’10”S, 145°24’19”E, 11.ix–24.x.1998 ( T88881View Materials 1 specimen in QM)GoogleMaps  ; 1.5 km SE Mount Emmett [16°10’00”S, 145°26’00”E], 23–24.iv.1999, C. Reid, (25-058172 1 specimen in ANIC)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 8.i–9.ii.1998, S. Grove ( T73216View Materials T73220View Materials 5 specimens in QM)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 9.ii–13.iii.1998, S. Grove ( T85540View Materials T85568View Materials 29 specimens in QM)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 13.iii–8.v.1998, S. Grove ( T88889View Materials T88890View Materials 2 specimens in QM)GoogleMaps  ; Cow Bay , N of Daintree River [16°12’00”S, 145°28’00”E], 25.x–15.xi.1987, R.I. Storey & Cunningham (25-058182 1 specimen in ANIC)GoogleMaps  ; Cow Bay , N of Daintree River [16°12’00”S, 145°28’00”E], 6.ii–23.iii.1988, R.I. Storey & Cunningham (25-058181 1 specimen in ANIC)GoogleMaps  ; Cow Bay , N of Daintree River [16°12’00”S, 145°28’00”E], 6.i–5.ii.1988, R.I. Storey & I. Cunningham (25-058180 4 specimens in ANIC)GoogleMaps  ; Mangrove Road , 16°12’42”S, 145°25’18”E, 8.i– 9.ii.1998, S. Grove ( T73207View Materials T73214View Materials 8 specimens in QM)GoogleMaps  ; Mangrove Road , 16°12’42”S, 145°25’18”E, 9.ii– 13.iii.1998, S. Grove ( T85546View Materials T85564View Materials 17 specimens in QM)GoogleMaps  ; Mangrove Road , 16°12’42”S, 145°25’18”E, 9.ix– 25.x.1998, S. Grove ( T88891View Materials T88897View Materials 7 specimens in QM)GoogleMaps  ; Mangrove Road , 16°12’42”S, 145°25’18”E, 13.iii–7.v.1998, S. Grove ( T88898View Materials T88900View Materials 3 specimens in QM)GoogleMaps  ; Hutchinson Creek , 16°13’04”S, 145°23’57”E, 13.i–11.ii.1998, S. Grove ( T73215View Materials specimens in QM)GoogleMaps  ; Hutchinson Creek , 16°13’04”S, 145°23’57”E, 14.iii–8.v.1998, S. Grove ( T88882View Materials 1 specimen in QM)GoogleMaps  ; Buchanan Creek , 16°14’39”S, 145°25’54”E, 5.i–11.ii.1998, S. Grove ( T73168View Materials T73171View Materials 4 specimens in QM)GoogleMaps  ; Buchanan Creek , 16°14’39”S, 145°25’54”E, 11.ii–12.iii.1998, S. Grove ( T85545View Materials 1 specimen in QM)GoogleMaps  ; Buchanan Creek , 16°14’39”S, 145°25’54”E, 12.ix–23.x.1998, S. Grove ( T88871View Materials 1 specimen in QM)GoogleMaps  ; Buchanan Creek , 16°14’39”S, 145°25’54”E, 12.iii–8.v.1998, S. Grove ( T88872View Materials 1 specimen in QM)GoogleMaps  ; Fairy Creek , 16°14’16”S, 145°24’57”E, 11.ii–13.iii.1998, S. Grove ( T85558View Materials T85565View Materials 8 specimens in QM)GoogleMaps  ; Fairy Creek , 16°14’16”S, 145°24’57”E, 13.iii–8.v.1998, S. Grove ( T88843View Materials T88851View Materials 9 specimens in QM)GoogleMaps  ; Windsor Tableland, Bargoo Creek Road [16°14’51”S, 145°04’08”E], 15–18.iv.1982 ( T44757View Materials specimens in QM)GoogleMaps  Windsor Tableland , 28 km NNW Mount Carbine, 16°17’05”S, 145°04’40”E, 15–18.iv.1982 ( T44749View Materials 1 specimen in QM)GoogleMaps  ; Wonga , 2 km N, 16°19’51”S, 145°25’05”E, 14–18.v.1998 ( T66805View Materials 1 specimen in QM)GoogleMaps  ; Devils Thumb transect Site 4, 16°23’39”S, 145°19’02”E, 26.xii.1989 – 15.i.1990 ( T44826View Materials T44827View Materials 2 specimens in QM)GoogleMaps  ; Mossman Gorge National Park , [16°28’00”S, 145°21’00”E], 13–14.iv.1973, R.I. Storey, (25-058475 2 specimens in QM)GoogleMaps  ; Rex Creek , site 1, 16°28’15”S, 145°19’30”E, 11.ii.2005 (T132288 1 specimen in QM)GoogleMaps  ; Mossman Bluff Site 1, 16°28’19”S, 145°19’37”E, 1–16.i.1989 ( T44767View Materials T44775View Materials 9 specimens in QM)GoogleMaps  ; Mossman Bluff Site 1, 16°28’19”S, 145°19’37”E 20.xii.1989 – 15.i.1990 ( T44857View Materials T44858View Materials 2 specimens in QM)GoogleMaps  ; Rex Range , summit, 16°32’12”S, 145°22’44”E, 15– 18.v.1998 ( T66803View Materials T66804View Materials 2 specimens in QM)GoogleMaps  ; Rex Creek , site 5, 16°28’23”S, 145°19’14”E, 21.xi.2005 (T132950 1 specimen in QM)GoogleMaps  ; Mossman Bluff Site 4, 16°28’29”S, 145°18’09”E, 1–16.i.1989, ( T44785View Materials T44792View Materials 8 specimens in QM)GoogleMaps  ; Mossman Bluff Site 4, 16°28’29”S, 145°18’09”E, 16–30.xii.1988 ( T44793View Materials T44799View Materials 7 specimens in QM)GoogleMaps  ; Mossman Bluff Site 4, 16°28’29”S, 145°18’09”E, 20.xii–15.i.1990 ( T44828View Materials T44838View Materials 11 specimens in QM)GoogleMaps  ; Mossman Bluff Site 2, 16°28’29”S, 145°19’03”E, 1–16.i.1989 ( T44776View Materials T44784View Materials 8 specimens in QM)GoogleMaps  ; Mossman Bluff Site 2, 16°28’29”S, 145°19’03”E, 16–30.xii.1988 ( T44811View Materials T44818View Materials 8 specimens in QM)GoogleMaps  ; Mossman Bluff Site 2, 16°28’29”S, 145°19’03”E, 20.xii.1989 – 15.i.1990 ( T44851View Materials T44856View Materials 6 specimens in QM)GoogleMaps  ; Mossman Bluff Site 3, 16°28’35”S, 145°18’40”E, 1–16.i.1989 ( T44759View Materials T44766View Materials 8 specimens in QM)GoogleMaps  ; Mossman Bluff Site 3, 16°28’35”S, 145°18’40”E, 16–30.xii.1988 ( T44800View Materials T44810View Materials 11 specimens in QM)GoogleMaps  ; Mossman Bluff Site 3, 16°28’35”S, 145°18’40”E, 20.xii.1989 – 15.i.1990 ( T44839View Materials T44850View Materials 12 specimens in QM)GoogleMaps  ; Mossman Gorge National Park , 6 km SW of Mossman [16°30’00”S, 145°20’00”E], 50 m, 11–18.vii.1982, S. Peck & J. Peck (25-058130 2 specimens in ANIC)GoogleMaps  ; Mount Perseverance [16°32’00”S, 145°20’00”E], 500 m, 4.iv.1997, K. Abbott (25-058185 5 specimens in ANIC)GoogleMaps  ; Rex Range , 16°32’02”S, 145°23’13”E, 430 m, 20–21.ix.2010, G.B. Monteith (25-060874 1 specimen also as COL 1139 in ANIC)GoogleMaps  ; Julatten [16°34’02”S, 145°21’37”E], 8.ii.1983, A. Walford–Huggins (CMNEN-00030070 1 specimen in CMNC)GoogleMaps  ; Clacherty Road via Julatten [16°34’19”S 145°21’52”E, 8.i–2.ii.1987, H. Howden & R.I. Storey (CMNEN-00030062, 00030197 2 specimens in CMNC)GoogleMaps  ; Black Mountain Kuranda Forest Station [16°49’24”S, 145°39’13”E], 30–31.xii.1988, H. Howden & A. Howden (CMNEN-00030066 1 specimen in CMNC)GoogleMaps  ; Julatten [16°37’00”S, 145°20’00”E], 10–29.ix.1987, A. Walford-Huggins (25-058139 2 specimens in ANIC)GoogleMaps  ; Julatten [16°37’00”S, 145°20’00”E], 20–29.x.1987, A. Walford-Huggins (25-058148 1 specimen in ANIC)GoogleMaps  ; Julatten [16°37’00”S, 145°20’00”E], 29.x–21.xi.1987, A. Walford-Huggins (25-058138 1 specimen in ANIC)GoogleMaps  ; Julatten [16°37’00”S, 145°20’00”E], 30.xi–13.xii.1987, A. Walford-Huggins (25-058147 1 specimen in ANIC)GoogleMaps  ; Julatten [16°37’00”S, 145°20’00”E], 5–20.x.1987, A. Walford-Huggins (25-058146 1 specimen in ANIC)GoogleMaps  ; Black Mountain , lower slopes, 16°39’52”S, 145°30’04”E, 2.xii.1997 – 6.ii.1998 ( T57475View Materials T57478View Materials 4 specimens in QM)GoogleMaps  ; Quaid Road, Mareeba , 16°44’11”S, 145°28’37”E, 17.iii–14.iv.1998, De Faveri & K.H. Halfpapp (25-058563 3 specimens in QDPC)GoogleMaps  ; 5 km NNW of Kuranda [16°45’00”S, 145°37’00”E], 15.i–20.ii.1985, R.I. Storey & K.H. Halfpapp (25- 058164 1 specimen in ANIC)GoogleMaps  ; 4 km NNW of Kuranda [16°47’00”S, 145°37’00”E], 10.xii.1984 – 15.i.1985, R.I. Storey & K.H. Halfpapp (25-058156 2 specimens in ANIC)GoogleMaps  ; Kuranda St. Forest , 3 km N Kuranda [16°49’00”S, 145°38’00”E], 360 m, 25.vi–3.viii.1982, S. Peck & J. Peck (25-058129 3 specimens in ANIC)GoogleMaps  ; Saddle Mountain , summit, 16°49’11”S, 145°39’42”E, 10.xii.1995 – 2.ii.1996 ( T44544View Materials T44548View Materials 5 specimens in QM)GoogleMaps  ; Saddle Mountain , summit, 16°49’11”S, 145°39’42”E, 3.xi–10.xii.1995 ( T44549View Materials T44550View Materials 2 specimens in QM)GoogleMaps  ; 8 km SW Kuranda [16°52’00”S, 145°34’00”E], iii.1993, G. O’Reilly (25-060765 1 specimen also as COL 741 in AM)GoogleMaps  ; 6 km SW of Kuranda , [16°52’00”S, 145°37’00”E], 10.xii.1984 – 15.i.1985, R.I. Storey & K.H. Halfpapp (25-058157 3 specimens in ANIC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 15.i–20.ii.1985, R.I. Storey & K.H. Halfpapp (25-058155 4 specimens in ANIC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 20.ii–20.iii.1985, R.I. Storey & K.H. Halfpapp (25-032566 1 specimen in QDPC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 20.ix– 6.xi.1984, R.I. Storey & K.H. Halfpapp (25-032565 1 specimen in QDPC, 25-058168 1 specimen in ANIC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 6.xi–10.xii.1984, R.I. Storey & K.H. Halfpapp (25-032564 1 specimen in QDPC)GoogleMaps  ; Mount Williams summit, 16°54’53”S, 145°39’43”E, 27.xi.1997 – 10.ii.1998 ( T57468View Materials T57474View Materials 7 specimens in QM)GoogleMaps  ; Mount Williams summit, 16°54’53”S, 145°39’43”E, 27–28.xi.1997 ( T57482View Materials T57483View Materials 2 specimens in QM)GoogleMaps  ; Cedar Pocket , [16°56’00”S, 145°36’00”E], 440 m, 17–19.iv.1999, C. Reid & I. Reid (25-058174 1 specimen in ANIC)GoogleMaps  ; Whitfield Range , 16°57’33”S, 145°42’31”E, 28.viii–19.x.1991 ( T44824View Materials 1 specimen in QM)GoogleMaps  ; Pine Creek Tower , 16°59’36”S, 145°50’07”E, 12.ix–20.x.1991 ( T44739View Materials T44740View Materials 2 specimens in QM)GoogleMaps  ; Lambs Head , 20 km SW Cairns, 17°01’23”S, 145°38’33”E, 4–13.xii.1988 ( T44671View Materials 1 specimen in QM)GoogleMaps  ; Davies Creek Road, via Mareeba , 17°02’33”S, 145°36’51”E, 4–13.xii.1988 ( T44696View Materials T44710View Materials 15 specimens in QM)GoogleMaps  ; Mount Haig [17°06’00”S, 145°36’00”E], 1150 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-058179 1 specimen in ANIC)GoogleMaps  ; Robson Creek, Atherton [17°07’00”S, 145°37’00”E], 18.i.1996, R.J. Kitching & H. Mitchell (25-058150 1 specimen in ANIC)GoogleMaps  ; Robson Creek Atherton [17°07’00”S, 145°37’07”E], 17.i.1996, R.J. Kitching & H. Mitchell (25-058186 1 specimen in ANIC)GoogleMaps  ; Robson Creek Atherton [17°07’00”S, 145°37’07”E], 18.i.1996, R.J. Kitching & H. Mitchell (25-058187 1 specimen in ANIC)GoogleMaps  ; Danbulla State Forest, Kauri Creek Trail, Tinaroo Lake , 17°08’03”S, 145°35’52”E, 625 m, 24.i.2000, R.A. Anderson (CMNEN-00030059 1 specimen in CMNC)GoogleMaps  ; Tinaroo Dam Drive , 17°08’31”S, 145°35’24”E, 3–4.xii.2010, G.B. Monteith (25-060830 1 specimen in ANIC aslo as COL1462)GoogleMaps  ; 17 km NE Yungaburra [17°09’40”S, 145°37’56”E], 21.xii.1986, H. Howden & A. Howden (CMNEN-00030051, 00030053 2 specimens in CMNC)GoogleMaps  ; Lake Euramoo , 17°09’46”S, 145°37’38”E, 10–11.ii.1999 ( T76207View Materials 1 specimen in QM)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve] [17°10’00”S, 145°38’00”E], 13.ii–6.iii.1987, R.I. Storey & De Faveri (25-032574 1 specimen in QDPC, 25-058165 1 specimen in ANIC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve] [17°10’00”S, 145°38’00”E], 20.xii.1986 – 13.i.1987, R.I. Storey & De Faveri (25-032573 1 specimen in QDPC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve] [17°10’00”S, 145°38’00”E], 23.iv–27.v.1987, R.I. Storey & De Faveri (25-058166 1 specimen in ANIC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve] [17°10’00”S, 145°38’00”E], 23.vi–28.vii.1987, R.I. Storey & De Faveri (25-058167 1 specimen in ANIC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve] [17°10’00”S, 145°38’00”E], 26.iii–23.iv.1987, R.I. Storey & De Faveri (25-032575 1 specimen in QDPC)GoogleMaps  ; Boar Pocket Road 3.9 m. N, [17°10’00”S, 145°39’00”E], 2500 feet, 4.viii.1974, J.G. Brooks (25-057895 1 specimen in ANIC)GoogleMaps  ; upper Mulgrave River [17°10’00”S, 145°53’00”E], 2.iv.1968, E. Matthews (25-057890 1 specimen in ANIC)GoogleMaps  ; Cathedral Tree , 17°10’37”S, 145°39’36”E, 3–4.xii.2010, G.B. Monteith, (25-057622 7 specimens in ANIC)GoogleMaps  ; Cathedral Tree , 17°10’37”S, 145°39’36”E, 4–5.xii.2010, G.B. Monteith (25-057621 2 specimens in ANIC)GoogleMaps  ; Cathedral Fig , 17°10’52”S, 145°39’26”E, 7.ii.1996 ( T44588View Materials T44589View Materials 2 specimens in QM)GoogleMaps  ; Danbulla Scientific Reserve , 17°11’09”S, 145°39’36”E, 10.xii.1995 – 7.ii.1996 ( T44601View Materials T44606, T44670, T47307 T47311View Materials 13 specimens in QM)GoogleMaps  ; Boar Pocket Road [17°12’00”S, 145°40’09”E], xi.1969, J.G. Brooks (CMNEN-00030063 1 specimen in CMNC)GoogleMaps  ; Boar Pocket Road [17°12’00”S, 145°40’09”E], 20.x.1974, J.G. Brooks (CMNEN-00030064–00030065 2 specimens in CMNC)GoogleMaps  ; Boar Pocket Road [17°12’00”S, 145°40’09”E], 27.i.1975, J.G. Brooks (CMNEN-00030071–00030072 2 specimens in CMNC)GoogleMaps  ; Boar Pocket Road [17°12’00”S, 145°40’09”E], viii.1974, J.G. Brooks (CMNEN-00030068 1 specimen in CMNC)GoogleMaps  ; Boar Pocket Road [17°12’00”S, 145°42’00”E], 2540 feet, 8.xii.1974, J.G. Brooks (25-057893 1 specimen in ANIC)GoogleMaps  ; Boar Pocket Road [17°12’00”S, 145°42’00”E], 21.ii.1970, J.G. Brooks (25-057896 1 specimen in ANIC)GoogleMaps  ; Gillies Highway [17°12’00”S, 145°42’00”E], 2760 feet, 24.xi.1974, J.G. Brooks (25-057892 1 specimen in ANIC)GoogleMaps  ; Gillies Road , 16 km NE Yungaburra [17°12’00”S, 145°42’00”E], 800 m, 24.vi–3.viii.1982, S. Peck & J. Peck (25-058128 1 specimen in ANIC)GoogleMaps  ; Tolga Scrub [17°14’00”S, 145°28’00”E], 760 m, 17–19.iv.1999, C. Reid & I. Reid (25-058171 2 specimens in ANIC)GoogleMaps  ; Tolga Scrub [17°14’42”S, 145°28’45”E], 23–24.ii.2001 (T106785–T106786 2 specimens in QM)GoogleMaps  ; Tolga , 17°14’06”S, 145°28’45”E, 9.xii.1989 – 14.i.1990 ( T44699View Materials T44702View Materials 4 specimens in QM)GoogleMaps  ; Tolga Scrub [17°14’42”S, 145°28’45”E], 3–6.ii.1999 ( T76202View Materials T76214View Materials 13 specimens in QM)GoogleMaps  ; Lake Barrine , 17°14’46”S, 145°38’24”E, 750 m, 17.xi.2009, G.B. Monteith & F. Turco (25-060784 1 specimen in ANIC also as COL774)GoogleMaps  ; Barrine National Park [17°15’00”S, 145°38’00”E], 760 m, 21.iii.1975, R.W. Taylor (25-057891 1 specimen in ANIC)GoogleMaps  ; Gadgarra State Forest , 10 km E Lake Eacham, 17°15’30”S, 145°41’16”E, 9–31.xii.1989 ( T44746View Materials T44747View Materials 2 specimens in QM)GoogleMaps  ; Yungaburra [17°16’00”S, 145°35’00”E], 7.v.1969, G. Bornemissza & P. Ferrar (25-057889 3 specimens in ANIC)GoogleMaps  ; Wrights Creek, Lake Eacham [17°16’00”S, 145°37’00”E], 750 m, 31.vii.1982, S. Peck & J. Peck (25-058132 1 specimen in ANIC)GoogleMaps  ; Mulgrave River Road, 7 km WbyS Bellenden Ker [17°16’00”S, 145°47’00”E], 60 m, 2.iv.1984, A.Calder & T.A.Weir (25-058140 3 specimens in ANIC)GoogleMaps  ; Bellenden Ker Range, Cableway Base Station [17°16’00”S, 145°54’00”E], 100 m, 17.x–9.xi.1981, Earthwatch, Queensland Museum (25-058133 1 specimen in ANIC)GoogleMaps  ; Bellenden Ker Range , 0.5 km S Cable Tower 7 [17°16’00”S, 145°55’00”E], 500 m, 25–31.x.1981, Earthwatch, Queensland Museum (25-058134 1 specimen in ANIC)GoogleMaps  ; 2 mi. S of Yungaburra [17°17’00”S, 145°33’00”E], 16– 18.iv.1973, R.I. Storey (25-058474 4 specimens in QM)GoogleMaps  ; Curtain Fig [17°17’00”S, 145°34’00”E], 700 m, 20– 21.iv.1999, C. Reid & I. Reid (25-058173 1 specimen in ANIC)GoogleMaps  ; Lake Eacham , 17°16’59”S, 145°37’28”E, 9.xii.1989 – 14.i.1990 ( T44819View Materials T44821View Materials 3 specimens in QM)GoogleMaps  ; Peeramon Scrub , 17°18’43”S, 145°37’34”E, 2.xi– 10.xii.1995 ( T44607View Materials T44609View Materials 3 specimens in QM)GoogleMaps  ; Peeramon Scrub , 17°18’43”S, 145°37’34”E, 10.xii.1995 – 7.ii.1996 ( T44610View Materials T44620View Materials 11 specimens in QM)GoogleMaps  ; Lake Eacham via Yungaburra , [17°17’00”S, 145°36’00”E], 14– 15.iv.1973, R.I. Storey (25-058472 2 specimens in QM)GoogleMaps  ; Lake Eacham [17°17’00”S, 145°37’00”E], 14.xi.1998 (25-057902 1 specimen in ANIC)GoogleMaps  ; Lake Eacham National Park [17°17’00”S, 145°38’00”E], 2–5.ii.1998, T.A. Weir (25-057901 2 specimens in ANIC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 1.xii.1983 – 9.i.1984, R.I. Storey & Brown (25-032555 2 specimens in QDPC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 1.xii.1983 – 9.i.1984, R.I. Storey & Brown (25-032561 1 specimen in QDPC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 1.v–21.vi.1984, R.I. Storey & Brown (25- 032560 1 specimen in QDPC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 10.ii– 13.iii.1984, R.I. Storey & Brown (25-032556 2 specimens in QDPC, 25-032558 2 specimens in QDPC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 10.xi–1.xii.1983, R.I. Storey & Brown (25-032582 1 specimen in QDPC, 25-032584 1 specimen in QDPC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 13.iii–1.v.1984, R.I. Storey & Brown (25-032559 1 specimen in QDPC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 3.ix–1.xi.1984, R.I. Storey & Brown (25-058149 4 specimens in ANIC, 25-058169 2 specimens in ANIC)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton [17°18’00”S, 145°31’00”E], 9.i–10.ii.1984, R.I. Storey & Brown (25-032557 2 specimens in QDPC, 25-032583 1 specimen in QDPC)GoogleMaps  ; Peeramon Scrub via Yungaburra [17°19’00”S, 145°36’00”E], 14–17.iv.1973, R.I. Storey (25-058473 1 specimen in QM)GoogleMaps  ; Wongabel State Forest , N end, 17°19’07”S, 145°29’37”E, 23–24.ii.2001 (T106782–T106784 3 specimens in QM)GoogleMaps  ; Wongabel State Forest , 17°19’29”S, 145°29’50”E, 5–14.xii.1988 ( T44693View Materials T44695View Materials 3 specimens in QM)GoogleMaps  ; Wongabel State Forest , 17°19’52”S, 145°30’04”E, 770 m, 22–23.i.2012, G.B. Monteith (25-053844, 25-055606 2 specimens in ANIC)GoogleMaps  ; Wongabel State Forest [17°20’02”S, 145°30’00”E], 760 m, 18–19.ix.2010, G.B. Monteith (25-060873 1 specimen in ANIC also as COL1138)GoogleMaps  ; Malanda Falls [17°21’00”S, 145°35’00”E], 14.v.1974, R.I. Storey (25-032587 1 specimen in QDPC)GoogleMaps  ; Malanda Falls , 17°21’22”S, 145°35’10”E, 750 m, 15–16.ix.2010, G.B. Monteith (25-060871 1 specimen in ANIC also as COL1136)GoogleMaps  ; Malanda Falls , 17°21’22”S, 145°35’10”E, 750 m, 23–24.i.2012, G.B. Monteith, (25-053846, 25-055610 2 specimens in ANIC)GoogleMaps  ; Bartle Frere , west base, 17°22’47”S, 145°45’51”E, 25.xi.1994 – 10.i.1995 ( T44859View Materials T44860, T47357 3 specimens in QM)GoogleMaps  ; Bartle Frere , west base, 17°22’47”S, 145°45’51”E, 7.iii–15.v.1995 ( T44859View Materials T44867, T74356 10 specimens in QM)GoogleMaps  ; Bartle Frere , west base, 17°22’47”S, 145°45’51”E, 8.xii.1988 ( T44733View Materials 1 specimen in QM)GoogleMaps  ; 2 km NNE Tarzali [17°24’00”S, 145°36’00”E], 740 m, 18–20.iv.1999, C. Reid & I. Reid (25-058176 1 specimen in ANIC)GoogleMaps  ; Topaz, Pei Road, ( Frith house), 17°24’23”S, 145°41’05”E, 6.xii.1993 – 25.ii.1994 ( T44675View Materials T44682View Materials 6 specimens in QM)GoogleMaps  ; Topaz, Westcott Road ( Cooper house), 17°24’49”S, 145°41’59”E, 6.xii.1993 – 25.ii.1994 ( T44822, T47349 T47350View Materials 3 specimens in QM)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton [17°25’00”S, 145°28’00”E], 12.x–5.xi.1983, R.I. Storey & Brown (25-032585 1 specimen in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton [17°25’00”S, 145°28’00”E], 13.iii–1.v.1984, R.I. Storey & Brown (25- 032577 2 specimens in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton [17°25’00”S, 145°28’00”E], 5.xi– 1.xii.1983, R.I. Storey & Brown (25-032576 1 specimen, 25-032586 1 specimen in QDPC)GoogleMaps  ; The Crater National Park [17°25’00”S, 145°29’00”E], 16–18.iv.1973, R.I. Storey (25-058471 10 specimens in QM)GoogleMaps  ; The Crater , 17°25’43”S, 145°29’13”E, 26.xi.1992 – 15.iv.1993 (T106771–T106781 11 specimens in QM)GoogleMaps  ; Crater National Park (= Mout Hypipamee National Park ) [17°26’00”S, 145°29’00”E], 25.i.1972, J.G. Brooks (25-057897 1 specimen in ANIC)GoogleMaps  ; Topaz, Hughes Road ( Breeden house), 17°26’04”S, 145°42’02”E, 6.xii.1993 – 25.ii.1994 ( T44687View Materials T44690View Materials 4 specimens in QM)GoogleMaps  ; Stone Creek , 17°27’40”S, 146°01’04”E, 1.xi.1995 – 6.ii.1996, J. Hasenpusch ( T44732View Materials 1 specimen in QM)GoogleMaps  ; Tower S of Crater National Park, 17°27’23”S, 145°29’12”E, 10.i–31.iii.1995 ( T44674View Materials 1 specimen in QM)GoogleMaps  ; Tower S of Crater National Park, 17°27’23”S, 145°29’12”E, 25.xi.1994 – 10.i.1995 ( T47344View Materials 1 specimen in QM)GoogleMaps  ; Longlands Gap [17°28’00”S, 145°29’00”E], 1150 m, 3.i–4.ii.1995, P. Zborowski (25-058178 1 specimen in ANIC)GoogleMaps  ; Kenny Road, Atherton Tablelands , 17°28’13”S, 145°32’04”E, 10.i–31.iii.1995 ( T44683View Materials T44709View Materials 17 specimens in QM)GoogleMaps  ; Kenny Road, Atherton Tablelands , 17°28’13”S, 145°32’04”E, 25.xi.1994 – 10.i.1995 ( T47338View Materials T47343View Materials 6 specimens in QM)GoogleMaps  ; Millaa Millaa Falls , 17°29’53”S, 145°36’32”E, 5.v.1983 ( T44758View Materials 1 specimen in QM)GoogleMaps  ; Millaa Millaa Falls , [17°30’00”S, 145°36’00”E], 780 m, 23–25.i.2012, G.B. Monteith (25-055611 1 specimen in ANIC)GoogleMaps  ; above Millaa Millaa Falls , 17°29’53”S, 145°36’47”E, 830 m, 23–25.i.2012, G.B. Monteith (25-053845 2 specimens in ANIC, 25-055608 2 specimens in ANIC)GoogleMaps  ; Millaa Millaa Falls (above), [17°30’00”S, 145°36’00”E], 800 m, 18– 20.iv.1999, C. Reid & I. Reid (25-058175 1 specimen in ANIC)GoogleMaps  ; Millaa Millaa Falls , [17°30’00”S, 145°37’00”E], 760 m, 11.v.1974, R.I. Storey (25-032579 1 specimen in QDPC)GoogleMaps  ; Millaa Millaa Falls [17°30’00”S, 145°37’00”E], 760 m, 13.vii–15.viii.1990, R.I. Storey & K.H. Halfpapp (25-032553 1 specimen in QDPC)GoogleMaps  ; Millaa Millaa Falls [17°30’00”S, 145°37’00”E], 760 m, 13.iii–10.iv.1990, R.I. Storey & K.H. Halfpapp (25-032552 3 specimens in QDPCGoogleMaps  ; Millaa Millaa Falls [17°30’00”S, 145°37’00”E], 760 m, 15.viii–12.ix.1990, R.I. Storey & K.H. Halfpapp (25-032554 1 specimen in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’00”S, 145°37’00”E], 760 m, 4.1– 7.ii.1990, R.I. Storey & K.H. Halfpapp (25-032549 7 specimens in QDPC)GoogleMaps  ; Millaa Millaa Falls [17°30’00”S, 145°37’00”E], 760 m, 7.ii–13.iii.1990, R.I. Storey & K.H. Halfpapp (25-032550–032551 2 specimens in QDPC)GoogleMaps  ; Mount Hugh Nelson, S base, 17°31’00”S, 145°33’15”E, 7–9.ii.1999 ( T76218View Materials 1 specimen in QM)GoogleMaps  ; Mount Hugh Nelson S base[17°31’00”S, 145°33’00”E], 1080 m, 7–9.ii.1999, G.B. Monteith & D. Cook (25-060845 1 specimen also as COL 584 in ANIC)GoogleMaps  ; Tom Carrick Road , 17°31’55”S, 145°30’50”E, 1070 m, 18–19.ix.2010, G.B. Monteith (25-060872 1 specimen in ANIC also as COL1137)GoogleMaps  ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 7.iii–15.v.1995 ( T44621View Materials T44632View Materials 12 specimens in QM)GoogleMaps  ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 10.i–31.iii.1995 ( T44633View Materials T44660, T55897 28 specimens in QM)GoogleMaps  ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 25.xi.1994 – 10.i.1995 ( T44661View Materials T44666, T47301 T47306View Materials 13 specimens in QM)GoogleMaps  ; The Millstream , 10 km NNE Ravenshoe, 17°31’57”S, 145°30’45”E, 6.xii.1998 – 5.ii.1999 ( T76208View Materials T76209View Materials 2 specimens in QM)GoogleMaps  ; Sluice Creek , 17°32’24”S, 145°31’56”E, 1.xii.1993 – 25.ii.1994 ( T44667View Materials T44669View Materials 3 specimens in QM)GoogleMaps  ; Sluice Creek , 17°32’24”S, 145°31’56”E, 5.ii–15.iv.1999 ( T79114View Materials 1 specimen in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 1.xii.1993 – 25.ii.1994 ( T44691View Materials T44692View Materials 2 specimens in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 27.iv.1982 ( T44703View Materials 1 specimen in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 8.xii.1989 ( T54145View Materials 1 specimen in QM)GoogleMaps  ; Palmerston National Park , west boundary, 17°33’36”S, 145°41’15”E, 1.xii.1993 – 25.ii.1994 ( T44744View Materials T44745View Materials 2 specimens in QM)GoogleMaps  ; Beatrice River , 2 km S, 17°34’35”S, 145°41’25”E, 1.xii.1993 – 25.ii.1994 ( T44823View Materials 1 specimen in QM)GoogleMaps  ; Palmerston Forestry turnoff, 17°35’20”S, 145°42’14”E, 700 m, 15–16.ix.2010, G.B. Monteith (25-060868 2 specimens in ANIC also as COL1128–29)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 7.iii–15.v.1995 ( T44551View Materials T44562View Materials 12 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995 ( T44563View Materials T44573View Materials 11 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 25.xi.1994 – 10.i.1995 ( T44574View Materials T44587, T47345 15 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995 ( T47346View Materials T47348View Materials 3 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 6–9.ii.1999 ( T76216View Materials T76217View Materials 2 specimens in QM)GoogleMaps  ; Mount Father Clancy , 2 km W, 17°35’30”S, 145°36’45”E, 25.vii–26.xi.1992 (T111993 1 specimen in QM)GoogleMaps  ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, 7.iii–15.v.1995 ( T44485View Materials T44513View Materials 19 specimens in QM)GoogleMaps  ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, 10.i–7.iii.1995 ( T44514View Materials T44527, T47312 T47325View Materials 28 specimens in QM)GoogleMaps  ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, 26.xi.1994 – 10.i.1995 ( T44528View Materials T44543, T47326 T47337View Materials 28 specimens in QM)GoogleMaps  ; Henrietta Creek, Palmerston National Park , 17°35’58”S, 145°45’28”E, 30.xi.1992 – 15.iv.1993 (T111990–T111992 3 specimens in QM)GoogleMaps  ; Mount Father Clancy , 800 m, 17°35’58”S, 145°38’04”E, 6–14.xii.1988 ( T44704View Materials T44708View Materials 5 specimens in QM)GoogleMaps  ; Mount Father Clancy , 1000 m, 17°35’25”S, 145°38’04”E, 6.xii.1988 ( T44825View Materials 1 specimen in QM)GoogleMaps  ; South Johnstone Research Station [17°36’00”S, 145°58’00”E], 11.x–29.xi.1990, K.H. Halfpapp (25-032562 1 specimen in QDPC)GoogleMaps  ; South Johnstone Research Station [17°36’00”S, 145°58’00”E], 29.xi.1989 – 14.i.1990, K.H. Halfpapp (25-032563 1 specimen in QDPC)GoogleMaps  ; Palmerston National Park , east margin, 17°36’40”S, 145°45’48”E, 10.xii.1995 – 7.ii.1996 ( T44742View Materials T44743View Materials 2 specimens in QM)GoogleMaps  ; Wooroonooran National Park, Patterson Sector, Tchupala Falls Trail, 17°36’25”S, 145°46’44”E, 22.i.2000, R.A. Anderson (CMNEN-00030060–00030061 2 specimens in CMNC)GoogleMaps  ; Wooroonooran National Park Patterson Sector, Crawfords Lookout , 17°36’46”S, 145°46’36”E, 550 m, 22.i.2000, R.A.Anderson (CMNEN-00030054–00030058 5 specimens in CMNC)GoogleMaps  ; Massey Creek [17°37’00”S, 145°34’00”E], 1000 m, 1.xii.1994 – 3.i.1995, P. Zborowski, (25-058177 4 specimens in ANIC)GoogleMaps  ; South Johnstone Forestry Camp , 5 km W, 17°38’12”S, 145°40’15”E, 1.xii.1993 – 25.ii.1994 ( T44590View Materials T44597View Materials 8 specimens in QM)GoogleMaps  ; Vine Creek, Majors Mountain , 17°40’58”S, 145°32’02”E, 15–18.iv.1999 ( T79115View Materials 1 specimen in QM)GoogleMaps  ; Charmillin Creek crossing [17°42’03”S, 145°31’21”E], 8.xii.1989 – 5.i.1990 ( T47353View Materials T47355View Materials 3 specimens in QM)GoogleMaps  ; Charmillin Creek crossing [17°42’03”S, 145°31’21”E], 1.xii.1997 ( T57479View Materials 1 specimen in QM)GoogleMaps  ; Charmillin Creek crossing [17°42’03”S, 145°31’21”E], 5–11.ii.1998 ( T57480View Materials T57481, T60417 3 specimens in QM)GoogleMaps  ; Charmillin Creek crossing [17°42’03”S, 145°31’21”E], 2–5.xii.1998 ( T70928View Materials 1 specimen in QM)GoogleMaps  ; Charmillin Creek , 17°42’54”S, 145°31’34”E, 880 m, 18–19.ix.2010, G.B. Monteith (25-060875 1 specimen in ANIC also as 1140)GoogleMaps  ; Ravenshoe State Forest, Tully Falls Road [17°43’00”S, 145°33’00”E], 10.x–15.xi.1987, A. Walford-Huggins (25-057898 5 specimens in ANIC, 25- 058145 1 specimen in ANIC)GoogleMaps  ; Ravenshoe State Forest, Tully Falls Road [17°43’00”S, 145°33’00”E], 15.xi– 12.xii.1987, A. Walford-Huggins (25-057899 11 specimens in ANIC, 25-058137 2 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest , 18 km SSW Ravenshoe [17°46’00”S, 145°31’00”E], 730 m, 1.x–5.xi.1987, R.I. Storey & Dickinson (25-058163 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest , 18 km SSW Ravenshoe [17°46’00”S, 145°31’00”E], 730 m, 20.iv–25.v.1988, R.I. Storey & Dickinson (25-058151 4 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest , 18 km SSW Ravenshoe [17°46’00”S, 145°31’00”E], 730 m, 3.iii–20.iv.1988, R.I. Storey & Dickinson (25-032571 1 specimen in QDPC)GoogleMaps  ; Tully Falls State Forest , 18 km SSW Ravenshoe [17°46’00”S, 145°31’00”E], 730 m, 5.i– 7.ii.1988, R.I. Storey & Dickinson (25-032569 1 specimen in QDPC, 25-058158 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest , 18 km SSW Ravenshoe [17°46’00”S, 145°31’00”E], 730 m, 7.xii.1987 – 7.i.1988, R.I. Storey & Dickinson (25-058154 3specimens in ANIC)GoogleMaps  ; Tully Falls State Forest , 18 km SSW Ravenshoe [17°46’00”S, 145°31’00”E], 730 m, 7.xi–7.xii.1987, R.I. Storey & Dickinson (25-058153 2 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest , 11 km SSW Ravenshoe [17°47’00”S, 145°34’00”E], 900 m, 1.x–5.xi.1987, R.I. Storey & Dickinson (25-032567 2 specimens in QDPC)GoogleMaps  ; Tully Falls State Forest , 11 km SSW Ravenshoe [17°47’00”S, 145°34’00”E], 900 m, 5.i–7.ii.1988, R.I. Storey & Dickinson (25-058161 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest , 11 km SSW Ravenshoe [17°47’00”S, 145°34’00”E], 900 m, 5.xi–7.xii.1987, R.I. Storey & Dickinson (25-058159 1 specimen in ANIC, 25- 032568 2 specimens in QDPC)GoogleMaps  ; Tully Falls State Forest , 11 km SSW Ravenshoe [17°47’00”S, 145°34’00”E], 900 m, 7.xii.1987 – 7.i.1988, R.I. Storey & Dickinson (25-058162 1 specimen in ANIC)GoogleMaps  ; Tully Falls , 30 mi. NE of Tully, [17°42’03”S, 145°59’19”E], 2500 feet, 5.vi.1972, G. Monteith & S.R. Monteith (CMNEN–00030069 1 specimen in CMNC)GoogleMaps  ; Tully Falls State Forest , 9.5 km SSW Ravenshoe [17°47’00”S, 145°34’00”E], 1000 m, 5.xi–7.xii.1987, R.I. Storey & Dickinson (25-058160 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest , 9.5 km SSW Ravenshoe [17°47’00”S, 145°34’00”E], 1000 m, 7.i–9.ii.1988, R.I. Storey & Dickinson (25-058152 3 specimens in ANIC)GoogleMaps  ; Tully Falls via Ravenshoe [17°47’00”S, 145°34’00”E], 20–21.iv.1973, R.I. Storey (25-032588 1 specimen in QDPC, 25-058470 3 specimens in QM)GoogleMaps  ; Tully Falls , 30 mi NE Tully [17°47’00”S, 145°34’00”E], 5.vi.1972, G.B & S.R. Monteith (25-058469 5 specimens in QM)GoogleMaps  ; Tully Falls , 17°47’00”S, 145°33’27”E, 8.xii.1989 – 5.i.1990 ( T44711View Materials T44717View Materials 7 specimens in QM)GoogleMaps  ; Red Road turnoff on Tully Falls Road , 17°49’19”S, 145°32’42”E, 8.xii.1989 – 5.i.1990 ( T44734View Materials T44736View Materials 3 specimens in QM)GoogleMaps  ; Upper Boulder Creek , top camp, 17°49’44”S, 145°54’02”E, 7.xii.1989 ( T44738View Materials 1 specimen in QM)GoogleMaps  ; Upper Boulder Creek , lower camp, 17°49’44”S, 145°54’02”E, 27.x.1983 ( T44748View Materials 1 specimen in QM)GoogleMaps  ; Koombooloomba Township , 2.5 km WSW, 17°50’28”S, 145°34’27”E, 5.xii.1998 – 4.ii.1999 ( T76215View Materials 1 specimen in QM)GoogleMaps  ; Koombooloomba Township , 2.5 km WSW, 17°50’28”S, 145°34’27”E, 4.ii–15.iv.1999 ( T79113View Materials 1 specimen in QM)GoogleMaps  ; Tully River crossing, 1.5 km N, 17°54’15”S, 145°37’18”E, 8.xii.1989 – 5.i.1990 ( T44750View Materials T47351View Materials 2 specimens in QM)GoogleMaps  ; Tully River crossing, 10 km S Koombooloomba Dam, 17°55’32”S, 145°37’38”E, 8.xii.1989 – 4.i.1990, ( T44737, T47352 2 specimens in QM)GoogleMaps  ; Kirrama Range, Douglas Creek , 18°13’30”S, 145°48’13”E, 10.xii.1986 – 11.i.1987 ( T44718View Materials T44731View Materials 14 specimens in QM)GoogleMaps  .

Description. Head and thorax dark brown to black, elytra predominately light brown to dark brown with humeri and apical edges reddish brown. Antennal clubs with apical antennomere (3 rd) white, basal 2 antennomeres (1 st and 2 nd) grey.

Total length: 3.5–5.0 mm. Lectotype measurements: Total length 3.8 mm; elytral width 2.7 mm.

Male. Head: Width to length ratio 37: 26. Surface smooth and nitid, with fine punctation. Dorsal part of eyes wide, separated by an interocular space approximately 9 times eye width (54: 6); eye canthus dividing or nearly dividing eye.

Prothorax: Pronotum with posterolateral angles distinct, lateral margins bordered. Pronotum surface smooth, nitid, finely punctate. Hypomeral surface very finely reticulate, hypomeral stria absent. Pronotum width to length ratio 60: 37.

Elytra: Surface smooth, nitid. Striae superficial, impunctate. Stria 6 extending to the elytral base, and stria 7 not extending to the elytral base. 8th stria sometimes punctate where it curves inwards posteriorly. Ratio of length of the elytra along suture to maximum elytral width 70: 75.

Legs: Protibia with a distinct crenulate ridge on the underside. Protibia with 2 teeth on outer edge, front edge truncate, apical digit broad, apical spur absent. Mesotibia with a brush of setae apically on inner edge. Metatibia with inner edge finely crenulate, nearly straight. Basal metatarsomere with slight lobe on inner edge.

Abdomen: Pygidium with a transverse sinuate fold across the base that is curved and not impressed medially. Pygidial disc with a large flat, reticulate central area that occupies more than half the disc. Ventrite 6 punctate.

Pterothorax: Mesometaventral suture arched. Medial lobe of metaventrite smooth, nitid, virtually impunctate, broadly margined beside mesocoxae. Lateral lobe of metaventrite finely punctate, setose. Mesoventrite smooth, nitid and notched anteriorly. Mesepimeron very finely reticulate. Metanepisternum reticulate.

Female. Protibia with front edge truncate and apical spur arising directly from front edge. Apical digit absent. Metatibia with inner edge not crenulate. Mesotibia without a brush of setae apically.

Distribution. Known from rainforests of the Australian Wet Tropics (WET) IBRA bioregion [Zones 1, 2, 3, 4, 6, 7, 8, 9, 10, 12, and 13], with occurrences recorded from south of Cooktown to west of Innisfail ( Fig. 7AView FIGURE 7). Specimens have been collected over a wide elevational gradient from sea level to 1150 m. BIOCLIM modelling suggests that climatic conditions are more suitable at higher elevations ( Fig. 7BView FIGURE 7).

Comments. The extensive collection of additional material since Matthews (1974) revision provided new insights into morphological variation between the Cape York specimens and L. globulus  that is restricted to only the Australian Wet Tropics. Molecular sequence data from multiple specimens sampled from across the geographic range of L. globulus  have been generated to confirm the distribution, its placement in the L. ustulatus  species group and sister relationship to L. cameroni  (see Fig. 1View FIGURE 1).

The majority of L. globulus  specimens have been collected in flight intercept traps but it has also collected in pitfall traps baited with dung or mushrooms, berlesed leaf and log litter, yellow pan traps, malaise traps, and at light. A recent study of food preference of Australian dung beetles reported observations for L. globulus  exposed to ten different baits: macropod dung, fowl dung, vertebrate carrion, three types of invertebrate carrion, and decaying banana, cucumber, pineapple, and mushroom ( Ebert et al. 2019). Forty–eight specimens were collected from dung (21 at kangaroo dung and 15 at fowl dung) and carrion (8 at vertebrate carrion, 3 at invertebrate carrion), while none were collected from decaying fruit and mushrooms.

Aristophanous (2014) report observations of L. globulus  across elevational gradients at three sites in the Atherton, Carbine, and Windsor subregions of the Australian Wet Tropics. The coordinates reported for each of these transects are in close proximity to known occurrences of L. globulus  and distant from that of the superficially similar species L. cardwellensis  , thus likely comprise records of only L. globulus  across these transects. Within the Atherton transect, the abundance of L. globulus  increased with elevation with 11 specimens collected at 100 m, 26 collected at 200 m, 24 collected at 400 m, 34 collected at 600 m, 64 collected at 800 m, and 169 collected at 1000 m ( Aristophanous 2014). While in the Carbine transect, L. globulus  was most abundant at the 600 m site (n = 47), with 12 specimens collected at 100 m, 28 collected at 200 m, 17 collected at 400 m, 3 collected at 800 m, and 11 collected at 1000 m ( Aristophanous 2014). Only three specimens were recorded at the 900 m site of the Windsor transect, with no occurrences at 1000 or 1100 m ( Aristophanous 2014). These results are broadly congruent with observational records of specimens examined here which show that L. globulus  is common at high elevations in the southern part of its range, but is only found below 600 m in the northern half. These observations are also congruent with the results of Monteith (1985a, 1985b) with L. globulus  only being recorded up to 200 m along a Cape Tribulation transect.

Species distribution models suggest that climatic conditions of higher-elevation habitats are more suitable for L. globulus  . Its absence in apparently suitable high-elevation habitat in the northern half of its range may be a result of competitive exclusion by other dung beetle species that dominate at high elevation. Storey & Monteith (2000) compared the distribution of apterous species of Aptenocanthon Matthews, 1974  and Temnoplectron Westwood, 1892  in the Wet Tropics noting a similar pattern of absence of Aptenocanthon  at high elevations in otherwise ammenable habitat and hypothesise the competive exclusion by flightless Temnoplectron  species. Competition from hyper-abundant upland endemic dung beetles is discussed in Aristophanous (2014) as the elevational transect data supports the occurance of competitive exclusion.

ANIC

Australian National Insect Collection

QM

Queensland Museum

QDPC

Queensland Primary Industries Insect Collection

AM

Australian Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus

Loc

Lepanus globulus ( Macleay, 1887 )

Gunter, Nicole L. & Weir, Thomas A. 2021
2021
Loc

Lepanus globulus:

Matthews, E. G. 1974: 106
1974
Loc

Epilissus globulus

Paulian, R. 1934: 282
Lea, A. M. 1923: 353
Gillet, J. J. E. 1911: 38
Macleay, W. 1887: 222
1887