Lepanus ustulatus (Lansberge, 1874)

Gunter, Nicole L. & Weir, Thomas A., 2021, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species, Zootaxa 4923 (1), pp. 1-66 : 5-12

publication ID

https://doi.org/10.11646/zootaxa.4923.1.1

publication LSID

lsid:zoobank.org:pub:41D057A7-F855-47C2-A598-8000CFD83A5B

DOI

http://doi.org/10.5281/zenodo.4537919

persistent identifier

https://treatment.plazi.org/id/1A323678-622A-2217-09D9-F88EFA5F6820

treatment provided by

Plazi

scientific name

Lepanus ustulatus
status

 

Lepanus ustulatus   species group

This group includes five species, all of which have been sequenced and form a strongly supported monophylum (see Fig. 1 View FIGURE 1 ).

Diagnosis. The L. ustulatus   species group is identified by their large size (> 3.5 mm) and by having a transverse sinuate fold across the base of the pygidium. The front edge of the protibia is truncate, the apical spur is absent in males and the apical digit is absent in females ( Fig. 2 View FIGURE 2 A–B). All species share the following characteristics. Occasional exceptions are listed in brackets.

Length: 3.5–6.5 mm.

Males. Head: Broad U-shape between clypeal teeth that are not upturned. Margin of head completely bordered, rounded; genal angles slightly prominent, angulate at clypeogenal suture. Mentum thickened anteriorly, reticulate with long setae basally and apically. Dorsal part of eye wide.

Prothorax: Pronotum anterior angles sharp. Basal edge rounded, with a fine margin medially, and without punctures. Lateral margins bordered. Hypomeral surface reticulate. Hypomeral stria absent.

Elytra: Intervals very finely punctate, striae superficial and impunctate. 8th stria slightly curving inwards posteriorly.

Legs: Protibia with tooth on underside near base of tarsus and two or three teeth on the outer edge. Apical digit short and broad. Front edge of protibia truncate; apical spur absent. Metatarsus with 5 tarsomeres with 2nd tarsomere the longest. Basal metatarsomere lobed on inner edge. Tarsal claws small and simple.

Abdomen: Pygidium with a transverse sinuate fold across the base. Abdominal ventrites reticulate for full width. Suture between ventrites 5 and 6 weak.

Pterothorax: Medial lobe of metaventrite broadly margined between mesocoxae. Lateral lobes of metaventrite punctate. Metanepisternum reticulate.

Hind wings: Fully developed.

Females. Protibia with apical spur arising from truncate front edge, apical digit absent. Tooth on the underside of protibia usually smaller than in males (except similar in L. cardwellensis   ). Inner edge of metatibia not crenulate.

Checklist of species

Lepanus cameroni Gunter & Weir   , new species

Lepanus cardwellensis Gunter & Weir   , new species

Lepanus globulus ( Macleay, 1887)  

Lepanus lemannae Gunter & Weir   , new species

Lepanus ustulatus ( Lansberge, 1874)  

Key to species of the Lepanus ustulatus   species group

1 Protibia with 3 teeth on outer edge ( Fig. 2 View FIGURE 2 A–B); elytral surfaces very finely reticulate, at least in part; generally larger species 4.5–6.5 mm .......................................................................................... 2

– Protibia with 2 teeth on outer edge ( Fig. 2C View FIGURE 2 ); elytral surfaces wholly non reticulate, nitid; generally smaller species 3.5–5.0 mm ................................................................................................. ................................................................................................... 3

2 Pygidium with reticulate depressed central area extending under the transverse sinuate fold ( Fig. 4D View FIGURE 4 ); male metatibia with inner edge more or less roundly expanded apically, bearing a defined brush of fine setae ( Fig. 2F View FIGURE 2 ); aedeagus as in Fig. 5D View FIGURE 5 ; north Queensland from Atherton Tableland to central Queensland south of Mackay. .................................................................................................. Lepanus lemannae Gunter & Weir   , new species

– Pygidium with reticulate depressed central area not extending under the transverse sinuate fold, separated from it by an impressed line ( Fig. 4E View FIGURE 4 ); male metatibia with inner edge not expanded apically, without a defined brush of fine setae ( Fig. 2D View FIGURE 2 ); aedeagus as in Fig. 5E View FIGURE 5 ; from central Queensland south of Gladstone to central New South Wales around Taree................................................................................. Lepanus ustulatus ( Lansberge, 1874)  

3 Eye canthus not dividing eye, leaving a gap of about 4 facets ( Fig. 2J View FIGURE 2 ); ridge on underside of protibia with coarse crenulations in the middle ( Fig. 2G View FIGURE 2 ); aedeagus as in Fig. 5A View FIGURE 5 ; north Queensland Iron Range................................................................................................... Lepanus cameroni Gunter & Weir   , new species

– Eye canthus dividing or nearly dividing eye, leaving a gap of not more than 1 facet ( Fig. 2L View FIGURE 2 ); ridge on underside of protibia more finely crenulated or without crenulations ( Figs. 2 View FIGURE 2 H–I).................................................... 4

4 Ridge on underside of protibia distinctly crenulated in the middle ( Fig. 2H View FIGURE 2 ); head usually finely punctate; posterolateral corners of pronotum distinct ( Fig. 2K View FIGURE 2 ); pygidium with reticulations of central depressed area obvious ( Fig. 4C View FIGURE 4 ); male protibia with apical digit broad ( Fig. 2C View FIGURE 2 ); aedeagus as in Fig. 5C View FIGURE 5 ; north Queensland Wet Tropics from south of Cooktown to Atherton Tableland................................................................... Lepanus globulus (Macleay, 1888)  

– Ridge on underside of protibia without crenulations but with a distinct flange in the middle ( Fig. 2I View FIGURE 2 ); head very finely punctate; posterolateral corners of pronotum rounded ( Fig. 2M View FIGURE 2 ); pygidium with reticulations of central depressed area less obvious, area more nitid ( Fig. 4B View FIGURE 4 ); male protibia with apical digit narrower; aedeagus as in Fig. 5B View FIGURE 5 ; north Queensland Cardwell Range area ......................................................... Lepanus cardwellensis Gunter & Weir   , new species

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus

Loc

Lepanus ustulatus

Gunter, Nicole L. & Weir, Thomas A. 2021
2021
Loc

Lepanus cameroni

Gunter & Weir 2021
2021
Loc

Lepanus cardwellensis

Gunter & Weir 2021
2021
Loc

Lepanus lemannae

Gunter & Weir 2021
2021
Loc

Lepanus globulus ( Macleay, 1887 )

DET 1971
1971