Lepanus ustulatus ( Lansberge, 1874 ),

Gunter, Nicole L. & Weir, Thomas A., 2021, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species, Zootaxa 4923 (1), pp. 1-66: 12-19

publication ID

https://doi.org/10.11646/zootaxa.4923.1.1

publication LSID

lsid:zoobank.org:pub:41D057A7-F855-47C2-A598-8000CFD83A5B

DOI

http://doi.org/10.5281/zenodo.4537921

persistent identifier

http://treatment.plazi.org/id/1A323678-6225-220E-09D9-FE2EFD326BAD

treatment provided by

Plazi

scientific name

Lepanus ustulatus ( Lansberge, 1874 )
status

 

Lepanus ustulatus ( Lansberge, 1874) 

( Figs. 2View FIGURE 2 A–B, D–E; 3E; 4E; 5E; 6A–B)

Epilissus ustulatus Lansberge, 1874: 7  ; Masters 1886: 10; Gillet 1911: 39; 1925: 5; Paulian 1934: 282.

Panelus ustulatus: Lea 1923: 361  .

Lepanus ustulatus: Matthews 1974: 104  .

Type material digitally examined. Lectotype: “Wide Bay” (handwritten) / “Ex-Musaeo VAN LANSBERGE” (printed) / “MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” (printed) / “ LECTOTYPE ” / “ Lepanus ustulatus (Lansberge)  . LECTOTYPE, Y. CAMBEFORT det. 1981” (hand written) / “ MNHN EC8675” (printed); Paralectotypes: “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)  ” (printed) / “ MNHN EC8676” (printed); “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)  ” (printed) / “ MNHN EC8677” (printed); “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)  ” (printed) / “ MNHN EC8678” (printed); “Brisbane” (handwritten) / “Ex-Musaeo VAN LANSBERGE” (printed) / “R. PAULIAN Vidit” (printed)/ “MUSÉUM PARIS 1952 coll. R. OBERTHÜR” (printed) / “ PARALECTOTYPE ” / “ PARALECTOTYPE Lepanus ustulatus ( Lansberge 1874)  ” (printed) / “ MNHN EC8679” (printed).

Other material examined. 1407 other specimens from over 200 unique localities.

Queensland: Mount Beecher , 0.5 km NW, 23°54’58”S, 151°10’53”E, 27.ix–15.xii.1999 (T105335 1 specimen in QM)GoogleMaps  ; Eurimbula Creek via Round Hill Head , [24°12’00”S 151°46’48”E], 2–5.v.1975, R.I. Storey, (25-058453 1 specimen in QM)GoogleMaps  ; Bulburin Barracks, 600 m, via Builyan [24°31’39”S, 151°28’01”E], 18.ix–2.xii.1989 ( T80281View Materials T80303View Materials 23 specimens in QM)GoogleMaps  ; Bulburin Barracks, 600 m, via Builyan [24°31’39”S, 151°28’01”E], 2.xii.1989 – 16.i.1990 ( T80304View Materials T80319View Materials 16 specimens in QM)GoogleMaps  ; Bulburin Barracks, 600 m, via Builyan , [24°31’39”S, 151°28’01”E], 7–9.x.1999 ( T89647View Materials T89648View Materials 2 specimens in QM)GoogleMaps  ; Bulburin State Forest , [24°32’00”S 151°23’00”E], 8.xii.1972, R.I. Storey, (25-058565 specimens in QDPC)GoogleMaps  ; Bulburin National Park , 24°31’26”S, 151°28’08”E, 570 m, 11–12.iv.2012, G.B. Monteith, (25-053941 2 specimens in ANIC, 25-053942 4 specimens in ANIC)GoogleMaps  ; Bulburin State Forest , [24°35’00”S 151°35’00”E], 12–14.iv.1974, B.P. Moore, (25-058331 1 specimen in ANIC)GoogleMaps  ; Mount Fort William, via Kalpowar [24°39’06”S, 151°20’24”E], 18.i.1990 ( T50268View Materials T50270,T80268 T80270View Materials 6 specimens in QM)GoogleMaps  ; Mount Fort William, via Kalpowar [24°39’06”S, 151°20’24”E], 2.xii.1989 – 16.i.1990 ( T50271, T80271 2 specimens in QM)GoogleMaps  ; Hurdle Gully , 10.8 km WSW Monto, 24°54’02”S, 151°01’10”E, 20–21.xii.1997 ( T85382View Materials 1 specimen in QM)GoogleMaps  ; Forest Station, Bulburin State Forest, via Many Peaks , [24°32’00”S, 151°23’00”E], 200m, 2– 5.iv.1972, S.R. Monteith, (25-058446 13 specimens in QM)GoogleMaps  ; Ventor Tableland, 21km ESE Monto, Site 1, 24°53’35”S, 151°19’59”E, 475 m, 1.x.2013 – 5.xii.2013, G.B. Monteith, (25-057705 15 specimens in ANIC)GoogleMaps  ; One Tree Hill , 1 km SE, 25°17’46”S, 151°54’36”E, 22–26.i.2001 (T105674–T105676 3 specimens in QM)GoogleMaps  ; One Tree Hill , 1.6km ENE, 25°17’05”S, 151°55’10”E, 14.xii.1999 – 19.iii.2000 ( T95331View Materials 1 specimen in QM)GoogleMaps  ; Yidney Scrub, Fraser Island , [25°19’00”S, 153°11’00”E], 1–4.xii.1975, G. Thompson & J.A. Slater, (25-058450 1 specimen in QM)GoogleMaps  ; Fraser Island, Garawongera Lake , 25°19’54”S, 153°09’14”E, 1–30.xi.1992 ( T80343View Materials 1 specimen in QM)GoogleMaps  ; Mount Woowoonga , 1.4 km SW, 25°26’28”S, 152°05’55”E, 21–22.i.2001 (T105673 1 specimen in QM)GoogleMaps  ; Mount Woowoonga , 1.4 km SW, 25°26’28”S, 152°05’55”E, 20–23.iii.2001 (T105315–T105321 7 specimens in QM)GoogleMaps  ; Fairlies Knob , 0.5 km SE, 25°30’32”S, 152°17’22”E, 20.xii.2000 – 23.iii.2001 (T105685 1 specimen in QM)GoogleMaps  ; Fairlies Knob , 0.5 km SE, 25°30’32”S, 152°17’22”E, 20.x–20.xii.2000 (T105694–T105696 5 specimens in QM)GoogleMaps  ; Wetheron , 3.5 km SW, 25°34’01”S, 151°41’13”E, 19.xii.1998 – 27.i.1999 ( T85224View Materials 1 specimen in QM)GoogleMaps  ; Teddington Weir , 25°39’11”S, 152°40’08”E, 30 m, 4–5.i.2012, G.B. Monteith, (25-053880 1 specimen in ANIC)GoogleMaps  ; Tinana Creek , 25°45’07”S, 152°41’38”E, 35 m, 6–7.xii.2011, G.B. Monteith, (25-053885 1 specimen in ANIC)GoogleMaps  ; Tallegalla Weir Junction , 25°45’25”S, 152°41’49”E, 35 m, 6–7.xii.2011, G.B. Monteith, (25-053886 3 specimens in ANIC)GoogleMaps  ; Stockhaven , 3 km SE, 25°48’03”S, 151°59’05”E, 10.x–19.xii.1998 ( T85323View Materials T85324View Materials 2 specimens in QM)GoogleMaps  ; Yarraman, Cooyar Range , 26°52’08”S, 151°54’10”E, 7.i–2.iii.1992, ( T78237View Materials T78242View Materials 6 specimens in QM)GoogleMaps  ; Yarraman, Cooyar Range , 26°52’08”S, 151°54’10”E, 2.iii–12.iv.1992 ( T78243View Materials T78244View Materials 2 specimens in QM)GoogleMaps  ; Yarraman, Cooyar Range , 26°52’08”S, 151°54’10”E, 1.xii.1991 – 7.i.1992 ( T78245View Materials 1 specimen in QM)GoogleMaps  ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 30.xi.1991 – 8.i.1992 ( T78039View Materials T78053View Materials 15 specimens in QM)GoogleMaps  ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 8.i–3.iii.1992 ( T78076View Materials T78081View Materials 6 specimens in QM)GoogleMaps  ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 3.iii–21.iv.1992 ( T78082View Materials T78098View Materials 17 specimens in QM)GoogleMaps  ; Cooloola, Womalah State Forest , 25°56’58”S, 153°05’31”E, 30.xi.1991 – 8.i.1992 ( T78099View Materials T78110View Materials 12 specimens in QM)GoogleMaps  ; Cooloola 3, 25°57’04”S, 153°06’43”E, 3.iii–21.iv.1992 ( T78054View Materials T78068View Materials 15 specimens in QM)GoogleMaps  ; Cooloola 3, 25°57’04”S, 153°06’43”E, 30.xi.1991 – 8.i.1992 ( T78069View Materials T78074View Materials 6 specimens in QM)GoogleMaps  ; Bymien, Cooloola , 25°57’14”S, 153°06’18”E, 150 m, 3–4.ii.2013, G.B. Monteith, (25-057693 2 specimens, 25-057694, 25-057695 3 specimens, 25-057698 1 specimen in ANIC)GoogleMaps  ; Freshwater Road, Cooloola , [25°58’00”S, 153°09’00”E], 9.xi.1974, G.B. Monteith, (25-058462 1 specimen in QM)GoogleMaps  ; Cooloola, via Gympie , [25°58’00”S, 153°09’00”E], 25–27.iv.1975, R.I. Storey, (25-058448 7 specimens in QM)GoogleMaps  ; Gunalda 3.6 km NE, [25°58’00”S 152°35’00”E], 140 m, 21–26.i.2001, D.J. Cook, (25-060844 2 specimens also as COL582, COL 583 specimens in ANIC)GoogleMaps  ; Gunalda , 3.6 km NE, 25°58’26”S, 152°35’07”E, 18–21.xii.2007 (T154645–T154650 6 specimens in QM)GoogleMaps  ; Gunalda , 3.6 km NE, 25°58’26”S, 152°35’07”E, 15–17.ii.2008 (T154651 1 specimen in QM)GoogleMaps  ; Gunalda , 3.6 km NE, 25°58’26”S, 152°35’07”E, 21– 26.i.2001 (T105677–T105678 2 specimens in QM)GoogleMaps  ; 2 km SE of Camp Milo, Cooloola National Park , [26°00’00”S, 153°05’00”E], 17–18.iv.1982, A. Calder, (25-057974 1 specimen in ANIC)GoogleMaps  ; Mudlo National Park , top, [26°00’40”S 152°13’52”E], 320 m, 20–21.xii.2011, G.B. Monteith, (25-053882 1 specimen in ANIC)GoogleMaps  ; Mudlo Gap , top site, 26°00’48”S, 152°13’46”E, 25.i–7.vi.1999 ( T95330, T85211 T85213View Materials 4 specimens in QM)GoogleMaps  ; Mudlo Gap , top site, 26°00’48”S, 152°13’46”E, 17.xii.1998 – 25.i.1999 ( T85214View Materials T85215View Materials 2 specimens in QM)GoogleMaps  ; Mudlo Gap , top site, 26°00’48”S, 152°13’46”E, 9.x–17.xii.1998 ( T85216View Materials 1 specimen in QM)GoogleMaps  ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 17.xii.1998 – 25.i.1999 ( T85209View Materials T85210View Materials 2 specimens in QM)GoogleMaps  ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 9.x–17.xii.1998 ( T85217View Materials T85221View Materials 5 specimens in QM)GoogleMaps  ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 25.i.1999 ( T85222View Materials 1 specimen in QM)GoogleMaps  ; Mudlo Gap , bottom site, 26°01’27”S, 152°13’57”E, 25.i– 7.vi.1999 ( T85223View Materials 1 specimen in QM)GoogleMaps  ; Oakview State Forest , barracks, 26°06’47”S, 152°19’36”E, 15–16.i.2002 (T122782 1 specimen in QM)GoogleMaps  ; Nangur State Forest , 1st site, 26°07’45”S, 151°58’15”E, 24.xi.1995 – 3.ii.1996 ( T78290View Materials T85397View Materials 8 specimens in QM)GoogleMaps  ; Nangur State Forest , 2nd site, 26°07’55”S, 151°58’43”E, 24.xi.1995 – 3.ii.1996 ( T78296View Materials T85398View Materials 3 specimens in QM)GoogleMaps  ; Jack Smith Environmental Park on Boat Mountain , 26°09’17”S, 151°54’34”E, 15.xii.1994 – 26.i.1995 ( T80226View Materials T80227View Materials 2 specimens in QM)GoogleMaps  ; Oakview State Forest , summit, 26°09’40”S, 152°19’43”E, 15.i–27.v.2002 (T111420–T111429 10 specimens in QM)GoogleMaps  ; Oakview State Forest , summit, 26°09’40”S, 152°19’43”E, 25.ix.2001 – 15.i.2002 (T116837–T116846 10 specimens in QM)GoogleMaps  ; Oakview State Forest , 26°08’32”S, 152°20’12”E, 12.xii.1997 – 26.iv.1998 ( T85311View Materials T85315View Materials 5 specimens in QM)GoogleMaps  ; Gympie , 11 km ESE, 26°14’24”S, 152°45’36”E, 1.i–28.ii.2002 (T111395 1 specimen in QM)GoogleMaps  ; Marys Creek, Gympie , 26°15’44”S, 152°32’40”E, 1974–1975 (T105313 1 specimen in QM)GoogleMaps  ; Cooran Tableland , BS68, 26°15’49”S, 152°49’43”E, 16.i– 27.v.2002 (T111401–T111406 6 specimens in QM)GoogleMaps  ; Cooran Tableland , 26°16’25”S, 152°50’05”E, 30.xi.1991 – 8.i.1992 ( T77869View Materials T77911View Materials 3 specimens in QM)GoogleMaps  ; Cooran Tableland , 26°16’25”S, 152°50’05”E, 8.i–2.iii.1992 ( T77912View Materials T77952View Materials 41 specimens in QM)GoogleMaps  ; Cooran Tableland , 26°16’25”S, 152°50’05”E, 2.iii–17.iv.1992 ( T77953View Materials T77969View Materials 17 specimens in QM)GoogleMaps  ; Cooran Barracks , 0.5 km N, 26°17’01”S, 152°49’57”E, 16.i–27.v.2002 (T111396– T111400 5 specimens in QM)GoogleMaps  ; Cooran Barracks , 0.5 km N, 26°17’01”S, 152°49’57”E, 25.ix.2001 – 16.i.2002 (T116859 1 specimen in QM)GoogleMaps  ; Cooran Plateau, 1300 feet via Gympie , [26°21’00”S, 152°47’00”E], 396 m, 3.x– 3.xi.1973, R.I. Storey, (CMNEN-00030167 specimens in CMNC)GoogleMaps  ; Cooran Plateau via Gympie , [26°21’00”S, 152°47’00”E], 1300 m, 10–11.iii.1973, R.I. Storey, (25-058447 20 specimens in QM, 25-058564 2 specimens in QDPC)GoogleMaps  ; Amamoor Creek , 26°23’55”S, 152°36’00”E, 15.i–27.v.2002 (T111410–T111417 8 specimens in QM)GoogleMaps  ; Amamoor Creek , 26°23’55”S, 152°36’00”E, 24.ix.2001 – 15.i.2002 (T116847–T116851 5 specimens in QM)GoogleMaps  ; Mount Kandanga , 26°27’07”S, 152°34’40”E, 15.i–27.v.2002 (T111418–T111419 2 specimens in QM)GoogleMaps  ; Mount Kandanga , summit, 26°27’07”S, 152°34’40”E, 24.ix.2001 – 15.i.2002 (T116852 specimens in QM)GoogleMaps  ; Dwyer Creek, Imbil , 26°27’53”S, 152°38’09”E, 15.i–27.v.2002 (T111407–T111409 3 specimens in QM)GoogleMaps  ; Dwyer Creek, Imbil , 26°27’53”S, 152°38’09”E, 24.ix.2001 – 15.i.2002 (T116871–T116876 6 specimens in QM)GoogleMaps  ; Perry’s Knob , 4 km NNE Rosewood, 27°36’11”S, 152°36’17”E, 11.xi.1998 – 13.i.1999 ( T85290View Materials 1 specimen in QM)GoogleMaps  ; Perry’s Knob , 4 km NNE Rosewood, 27°36’11”S, 152°36’17”E, 13.i–16.v.1999 ( T85291View Materials T85309View Materials 19 specimens in QM)GoogleMaps  ; 1 mi N Nambour , [26°37’08”S, 152°58’00”E], 27.v–2.vi.1975, R.I. Storey (25-058452 7 specimens in QM)GoogleMaps  ; Little Yabba Creek , 26°37’22”S, 152°41’26”E, 2.iii–12.iv.1992 ( T77845View Materials T77855View Materials 11 specimens in QM)GoogleMaps  ; Little Yabba Creek , 26°37’22”S, 152°41’26”E, 7.i.1992 – 2.iii.1992 ( T77856View Materials T77863View Materials 8 specimens in QM)GoogleMaps  ; Little Yabba Creek , 26°37’22”S, 152°41’26”E, 30.xi.1991 – 7.i.1992 ( T77864View Materials T77868View Materials 5 specimens in QM)GoogleMaps  ; Mapleton Falls National Park , 26°37’49”S, 152°50’21”E, 8.i–03.iii.1992 ( T78298View Materials T78316View Materials 17 specimens in QM)GoogleMaps  ; Mapleton Falls National Park , 26°37’49”S, 152°50’21”E, 30.xi.1991 – 8.i.1992 ( T78317View Materials 1 specimen in QM)GoogleMaps  ; Mapleton Falls National Park , 26°37’49”S, 152°50’21”E, 3.iii–17.iv.1992 ( T80131View Materials T80147View Materials 17 specimens in QM)GoogleMaps  ; Jimna , 26°38’37”S, 152°27’12”E, 29.xi.1991 – 7.i.1992 ( T78142View Materials T78147View Materials 6 specimens in QM)GoogleMaps  ; Jimna , 26°38’37”S, 152°27’12”E, 7.i– 02.iii.1992 ( T78148View Materials T78156View Materials 9 specimens in QM)GoogleMaps  ; Jimna , 26°38’37”S, 152°27’12”E, 02.iii–12.iv.1992 ( T78157View Materials T78184View Materials 28 specimens in QM)GoogleMaps  ; 20 km NW Maleny, Booloumba Creek , [26°38’44”S, 152°39’03”E], 150 m, 26– 29.iii.1993, C. Carlton, (CMNEN-00030170 1 specimen in CMNC)GoogleMaps  ; Tungi Creek, Jimna , 26°38’57”S, 152°28’21”E, 18.xii.1996 – 20.i.1997 ( T78185View Materials T78188View Materials 4 specimens in QM)GoogleMaps  ; Tungi Creek, Jimna , 26°38’57”S, 152°28’21”E, 20.i–9.iii.1997 ( T78189View Materials 1 specimen in QM)GoogleMaps  ; Tungi Creek, Jimna , 26°38’57”S, 152°28’21”E, 9.iii–15.v.1997 ( T78190View Materials T78196View Materials 7 specimens in QM)GoogleMaps  ; Booloumba Creek , 26°39’09”S, 15802602 °38’32”E, 29.xi.1991 – 8.i.1992 ( T77757View Materials T77778View Materials 22 specimens in QM)  ; Booloumba Creek , 26°39’09”S, 152°38’32”E, 8.i–2.iii.1992 ( T77779View Materials T77806View Materials 28 specimens in QM)GoogleMaps  ; Booloumba Creek , 26°39’09”S, 152°38’32”E, 2.iii–12.iv.1992 ( T77807View Materials T77826View Materials 20 specimens in QM)GoogleMaps  ; Sunday Creek, Conondale Range 26°42’39”S, 152°33’49”E, 7.i–2.iii.1992 ( T80176View Materials 1 specimen in QM)GoogleMaps  ; Sunday Creek, Conondale Range , 26°42’39”S, 152°33’49”E, 29.xi.1991 – 7.i.1992 ( T80177View Materials 1 specimen in QM)GoogleMaps  ; Mary Cairncross Park , 26°46’52”S, 152°52’56”E, 29.xi.1991 – 8.i.1992 ( T80251View Materials T80259View Materials 9 specimens in QM)GoogleMaps  ; Mary Cairncross Park , 26°46’52”S, 152°52’56”E, 8.i–3.iii.1992 ( T80260View Materials T80261View Materials 2 specimens in QM)GoogleMaps  ; Mary Cairncross Park , 26°46’52”S, 152°52’56”E, 3.iii–12.iv.1992 ( T80262View Materials T80263View Materials 2 specimens in QM)GoogleMaps  ; Dularcha National Park , 26°47’41”S, 152°58’04”E, 1–3.iv.2004 (T122767 1 specimen in QM)GoogleMaps  ; Landsborough , 2.5 km NE, 26°47’31”S, 152°58’40”E, 25–27.iii.2004 (T122762–T122766 5 specimens in QM)GoogleMaps  ; Yarraman , 2.2 km SE, 26°51’35”S, 151°59’36”E, 14–15.i.1998 ( T78203View Materials 1 specimen in QM)GoogleMaps  ; Yarraman , 26°51’38”S, 152°00’34”E, 2.iii– 12.iv.1992 ( T78246View Materials T78248View Materials 3 specimens in QM)GoogleMaps  ; Yarraman , 26°51’38”S, 152°00’34”E, 7.i–2.iii.1992 ( T78249View Materials 1 specimen in QM)GoogleMaps  ; Saddle Tree Creek , 26°51’37”S, 151°38’06”E, 1.iii–12.iv.1992 ( T80248View Materials T80250View Materials 3 specimens in QM)GoogleMaps  ; Mellum Creek Bridge, 26°51’51”S, 153°01’02”E, 22.ii–14.iii.2008 (T156697 1 specimen in QM)GoogleMaps  ; Saddle Tree Creek, Bunya Mountains , 26°52’09”S, 151°37’26”E, 1.i.1975 – 31.xii.1976 (T105312 1 specimen in QM)GoogleMaps  ; Top of Blackbutt Range , 26°52’38”S, 152°11’32”E, 24.xi.1995 – 3.ii.1996 (T116877, T85400View Materials 2 specimens in QM)GoogleMaps  ; Blackbutt Range, 6 mi E of Blackbutt, [26°53’00”S, 152°11’00”E], 22–23.xii.1973, R.I. Storey, (25-058451 1 specimen in QM)GoogleMaps  ; Neurum Creek Conservation Park, 26°58’20”S, 152°40’54”E, 25–27.iii.2004 (T122761 1 specimen in QM)GoogleMaps  ; Brisbane, Forest Park, Scrub Creek Road, [27°01’00”S 152°26’00”E], 18.ix–29.x.1998, P. Bouchard, (25-058458 17 specimens in QM)GoogleMaps  ; Mount Glorious , [27°20’00”S, 152°46’00”E], 630 m, 14.xi.1986 – 30.i.1987, T. Hiller, (25-057981 5 specimens in ANIC)GoogleMaps  ; Mount Glorious , [27°20’00”S, 152°46’00”E], 13–20.iv.1975, R.I. Storey, (25-058449 1 specimen in QM)GoogleMaps  ; Mount Glorious , [27°20’00”S, 152°46’00”E], 10.x.1999, P. Bouchard, K. Evans, & J. Skevington (25-058464 4 specimens in QM)GoogleMaps  ; Mount Delaney , 0.9 km SSW, 27°01’14”S, 152°41’58”E, 10–11.ii.2001 (T105679 1 specimen in QM)GoogleMaps  ; Mount Mee , 27°02’53”S, 152°40’49”E, 29.xi.1991 – 8.i.1992 ( T80148View Materials T80153View Materials 6 specimens in QM)GoogleMaps  ; Mount Mee , 27°02’53”S, 152°40’49”E, 3.iii–12.iv.1992 ( T80154View Materials T80164View Materials 11 specimens in QM)GoogleMaps  ; Mount Mee , 27°02’53”S, 152°40’49”E, 8.i–3.iii.1992 ( T80165View Materials T80175View Materials 11 specimens in QM)GoogleMaps  ; Mount Deongwar , 2 km SE, 27°12’53”S, 152°16’45”E, 14.x–31.xii.1998 ( T85322View Materials 1 specimen in QM)GoogleMaps  ; Mount Deongwar, 3 km S, Site 2, 27°13’49”S, 152°15’25”E, 30.xii.1998 – 26.iii.1999 ( T85316View Materials T85321View Materials 5 specimens in QM)GoogleMaps  ; Mount Glorious , barracks, 27°19’54”S, 152°45’12”E, 16–28.iv.2007 (T148364–T148365 2 specimens in QM)GoogleMaps  ; Mount Glorious , 27°20’20”S, 152°45’30”E, 1.i–31.iii.1982 ( T78255View Materials T78256View Materials 2 specimens in QM)GoogleMaps  ; Stony Creek, via Samford , 27°20’20”S, 152°47’52”E, 22.x.1994 – 2.ii.1995 ( T78215View Materials T78232View Materials 18 specimens in QM)GoogleMaps  ; Stony Creek, via Samford , 27°20’20”S, 152°47’52”E, 2.ii–8.iv.1995 ( T78233View Materials T78236, T87587 5 specimens in QM)GoogleMaps  ; Cashs Crossing , [27°21’00”S, 152°58’00”E], 16.iii.1959, A. Diatloff, (25-058457 1 specimen in QM)GoogleMaps  ; Ravensbourne National Park , 27°22’09”S, 152°10’44”E, 1.iii–11.iv.1992 ( T77829View Materials T77835View Materials 7 specimens in QM)GoogleMaps  ; Ravensbourne National Park , 27°22’09”S, 152°10’44”E, 7.i–1.iii.1992 ( T77836View Materials T77844View Materials 9 specimens in QM)GoogleMaps  ; Boombana National Park , site 1, 27°24’08”S, 152°47’22”E, 15–16.ii.2004 (T122060–T122062 3 specimens in QM)GoogleMaps  ; Enoggera Reservoir , site 8, 27°26’37”S, 152°54’14”E, 15.iii–18.v.2000 (T105338 1 specimen in QM)GoogleMaps  ; Enoggera Reservoir , site 2, 27°26’28”S, 152°55’16”E, 27.i–15.iii.2000 (T105339 1 specimen in QM)GoogleMaps  ; Enoggera Reservoir , site 2, 27°26’28”S, 152°55’16”E, 16.x–21.xii.1999 (T105322–T105323 2 specimens in QM)GoogleMaps  ; Enoggera Reservoir , site 3, 27°26’44”S, 152°55’19”E, 16.x–21.xii.1999 (T105324–T105326 3 specimens in QM)GoogleMaps  ; Enoggera Reservoir , site 3, 27°26’44”S, 152°55’19”E, 27.i–15.iii.2000 (T105331–T105332 2 specimens in QM)GoogleMaps  ; Enoggera Reservoir , site 3, 27°26’44”S, 152°55’19”E, 10.xi.1999 (T118119–T118120 2 specimens in QM)GoogleMaps  ; Enoggera Reservoir , site 5, 27°26’44”S, 152°55’19”E, 16.x–21.xii.1999 (T105327–T105328 2 specimens in QM)GoogleMaps  ; Enoggera Reservoir , site 5, 27°26’44”S, 152°55’19”E, 21.xii.1999 – 27.i.2000 (T105329 1 specimen in QM)GoogleMaps  ; Enoggera Reservoir , site 5, 27°26’44”S, 152°55’19”E, 15.iii–18.v.2000 (T105337 1 specimen in QM)GoogleMaps  ; Enoggera Reservoir , site 1, 27°26’31”S, 152°55’27”E, 27.i–15.iii.2000 (T105330 1 specimen in QM)GoogleMaps  ; Enoggera Reservoir , site 1, 27°26’31”S, 152°55’27”E, 15.iii–18.v.2000 (T105336 1 specimen in QM)GoogleMaps  ; Jones Road, Brookfield , 27°28’08”S, 152°54’40”E, 12–13.ii.2011, G.B. Monteith, (25-053874–25-05376 3 specimens also as COL1673, COL1674, COL 1675 specimens in ANIC)GoogleMaps  ; Upper Brookfield Road , 27°28’21”S, 152°50’06”E, 12–13.ii.2011, G.B. Monteith, (25-053873, also as COL71672View Materials 1 specimen in ANIC)GoogleMaps  ; Slaughter Falls , 27°28’44”S, 152°57’34”E, 6–7.ii.2001 (T105680–T105684 5 specimens in QM)GoogleMaps  ; Upper Brookfield Road, Brisbane , [27°29’00”S, 152°52’00”E], 22.ix–4.x.1975, R.I. Storey, (25-058459 3 specimens in QM)GoogleMaps  ; Upper Brookfield Road, Brisbane , [27°29’00”S, 152°52’00”E], 24.viii–7.ix.1975, R.I. Storey, (25-058460 7 specimens in QM)GoogleMaps  ; Mount Coot–tha , [27°29’00”S, 152°57’00”E], 15–20.iii.1971, G.B. Monteith, (25-058456 1 specimen in QM)GoogleMaps  ; Moggill Creek Brisbane, 27°30’25”S, 152°55’52”E, 22–23.i.2011, G.B. Monteith, (25-060829 1 specimen in ANIC)GoogleMaps  ; The Knobby , 4 km W Glamorgan Vale, 27°30’27”S, 152°35’18”E, 13.i–16.v.1999 ( T85310View Materials 1 specimen in QM)GoogleMaps  ; Cameron’s Scrub , 27°30’32”S, 152°43’48”E, 75 m, 30.iv–1.v.2012, G.B. Monteith, (25-057576 2 specimens in ANIC)GoogleMaps  ; Cameron’s Scrub , top ridge, 27°30’29”S, 152°43’54”E, 13.i–16.v.1999 ( T85231View Materials T85234View Materials 4 specimens in QM)GoogleMaps  ; Cameron’s Scrub , gully, 27°30’35”S, 152°43’54”E, 13.i–16.v.1999 ( T85235View Materials T85259View Materials 25 specimens in QM)GoogleMaps  ; Cameron’s Scrub , gully, 27°30’35”S, 152°43’54”E, 11.xi.1998 – 13.i.1999 ( T85260View Materials T85286View Materials 27 specimens in QM)GoogleMaps  ; Cameron’s Scrub, Snig track, 27°30’49”S, 152°43’43”E, 13.i–16.v.1999 ( T85225View Materials T85230View Materials 6 specimens in QM)GoogleMaps  ; Cameron’s Scrub, Snig track, 27°30’49”S, 152°43’43”E, 11.xi.1998 – 13.i.1999 ( T85287View Materials T85289View Materials 3 specimens in QM)GoogleMaps  ; Belmont Hill , top, 27°30’48”S, 153°06’59”E, 12.xii.1997 – 7.v.1998 ( T78197View Materials T78199, T89630 T89646View Materials 20 specimens in QM)GoogleMaps  ; Belmont Hills Bushlands , 27°30’47”S, 153°07’05”E, 25–27.ii.2004 (T122063 1 specimen in QM)GoogleMaps  ; Belmont Hill , gully, 27°30’44”S, 153°07’06”E, 12.xii.1997 – 7.v.1998 ( T85325View Materials - T85338View Materials 14 specimens in QM)GoogleMaps  ; Pine Mountain (Flinton Hill in QM)  , 27°31’23”S, 152°44’45”E, 1975–1976 ( T78257View Materials T78258View Materials 2 specimens in QM)GoogleMaps  ; Redwood Park, Toowoomba , [27°35’00”S 151°59’00”E], 8.xi.1988, E.S. Nielsen & M. Horak, (25-057983 1 specimen in ANIC)GoogleMaps  ; Buhot Creek, Burbank , 27°35’27”S, 153°10’19”E, 6.xi.2003 (T122059 1 specimen in QM)GoogleMaps  ; Mount Cotton , 27°36’58”S, 153°12’21”E, 3.ix–12.xii.1997 ( T78205View Materials T78209View Materials 5 specimens in QM)GoogleMaps  ; Mount Cotton, Scott’s Dam , 27°36’58”S, 153°12’21”E, 12.xii.1997 – 7.v.1998 ( T85374View Materials T85381View Materials 8 specimens in QM)GoogleMaps  ; Mount Cotton , upper gully, 27°36’58”S, 153°12’32”E, 12.xii.1997 – 7.v.1998 ( T78200View Materials T78202, T85356 –T785373 21 specimens in QM)GoogleMaps  ; Perry’s Knob , 27°36’00”S, 152°36E, 13.i–16.v.1999 ( T98837View Materials 1 specimen in QM)GoogleMaps  ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 21.i–9.iii.1992 ( T77622View Materials T77647View Materials 26 specimens in QM)GoogleMaps  ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 10.xii.1991 – 21.i.1992 ( T77648View Materials T77670View Materials 23 specimens in QM)GoogleMaps  ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 9.iii–30.iv.1992 ( T77671View Materials T77682View Materials 12 specimens in QM)GoogleMaps  ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 10.xii.1991 – 21.i.1992 ( T77706View Materials T77726View Materials 27 specimens in QM)GoogleMaps  ; Bahr’s Scrub , 27°45’31”S, 153°10’06”E, 21.i–9.iii.1992 ( T77727View Materials T77729View Materials 3 specimens in QM)GoogleMaps  ; Mount Tamborine National Park , [27°53’11”S, 153°10’51”E], 24.vi–6.vii.1978, S. Peck & J. Peck (CMNEN- 00030168 1 specimen in CMNC)GoogleMaps  ; Tamborine Mountain (general), [27°55’00”S, 153°10’00”E], 7.xii.1967, E. Matthews, (25-057973 1 specimen in ANIC)GoogleMaps  ; Tamborine Mountain (general), [27°55’00”S, 153°10’00”E], 26.xii.1972, R.I. Storey, (25-058455 1 specimen in QM)GoogleMaps  ; Palm Grove, Tamborine [Tamborine Mountain], [27°55’00”S, 153°10’00”E], 28.ix–12.x.1975, R.I. Storey, (25-058461 2 specimens in QM)GoogleMaps  ; North Tamborine , 27°55’13”S, 153°11’50”E, 9.iii–25.iv.1992 (T105311 1 specimen in QM)GoogleMaps  ; North Tamborine , 27°55’13”S, 153°11’50”E, 11.i– 9.iii.1992 ( T78259View Materials T78277View Materials 19 specimens in QM)GoogleMaps  ; North Tamborine , 27°55’13”S, 153°11’50”E, 9.iii–25.iv.1992 ( T78278View Materials T78289View Materials 12 specimens in QM)GoogleMaps  ; Tamborine, Guanaba shelf, 27°55’19”S, 153°13’14”E, 1.xii.1992 – 31.i.1993 ( T80345View Materials T80346View Materials 2 specimens in QM)GoogleMaps  ; Mount French National Park , 28°00’07”S, 152°37’00”E, 2.xii.1991 – 6.i.1992 ( T80237View Materials T80240View Materials 4 specimens in QM)GoogleMaps  ; Mount French National Park , 28°00’07”S, 152°37’00”E, 6.i–29.ii.1992 ( T80241View Materials T80244View Materials 4 specimens in QM)GoogleMaps  ; Mount French National Park , 28°00’07”S, 152°37’00”E, 29.ii–11.iv.1992 ( T80245View Materials T80247View Materials 3 specimens in QM)GoogleMaps  ; Glen Witheren , main scrub, 28°02’48”S, 153°07’12”E, 30.xii.1991 – 24.v.1992 ( T80187View Materials T80195View Materials 9 specimens in QM)GoogleMaps  ; Glen Witheren , main scrub, 28°02’48”S, 153°07’12”E, 1.ii– 30.iv.1991 ( T80209View Materials T80210View Materials 2 specimens in QM)GoogleMaps  ; Glen Witheren , shelf scrub, 28°02’58”S, 153°06’58”E, 30.x– 30.xii.1991 ( T80196View Materials T80208View Materials 13 specimens in QM)GoogleMaps  ; Burleigh Headland National Park , 28°05’45”S, 153°27’25”E, 1.i–4.iii.1992 ( T77683View Materials T77695, T77741 T77742View Materials 15 specimens in QM)GoogleMaps  ; Burleigh Headland National Park , 28°05’45”S, 153°27’25”E, 9.xii.1991 – 1.i.1992 ( T77696View Materials T77705View Materials 10 specimens in QM)GoogleMaps  ; Burleigh Headland National Park , 28°05’45”S, 153°27’25”E, 4.iii–2.iv.1992 ( T77743View Materials T77756View Materials 14 specimens in QM)GoogleMaps  ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 2.xii.2000 – 13.v.2001 (T105663–T105672, T106763–T106764 12 specimens in QM)GoogleMaps  ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 6.x–3.xii.2000 (T105686–T105693 8 specimens in QM)GoogleMaps  ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 2.xii.2000 – 13.v.2001 (T105341–T105342 2 specimens in QM)GoogleMaps  ; Black Rock Scrub , 28°07’32”S, 152°39’48”E, 6.x–3.xii.2000 (T105333–T105334 2 specimens in QM)GoogleMaps  ; Environment   Centre, Numinbah Valley , 28°07’24”S, 153°14’10”E, 1978–1979 (T105314 1 specimen in QM)GoogleMaps  ; Bonogin Road , 0.5 km W Smith Road, 28°07’50”S, 153°21’11”E, 3–7.iii.2008 (T156691–T156692 2 specimens in QM)GoogleMaps  ; Smith Road , gully rainforest, 28°08’47”S, 153°21’36”E, 3–7.iii.2008 (T156693–T156696 4 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 13–23.i.2007 (T145225 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 16–26.x.2006 (T145229, T145234 2 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 21–26.x.2006 (T145238 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–D, 28°08’31.2”S, 153°07’58.8”E, 18–23.i.2007 (T145243 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–A, 28°08’52.8”S, 153°08’13.2”E, 13–23.i.2007 (T145222 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–A, 28°08’52.8”S, 153°08’13.2”E, 16–26.x.2006 (T145232 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 13–23.i.2007 (T145224 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 16–26.x.2006 (T145231 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 13–18.i.2007 (T145240 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–C, 28°09’3.6”S, 153°08’16.8”E, 18–23.i.2007 (T145242 1 specimen in QM)GoogleMaps  ; Beechmont Road , 28°09’22”S, 153°11’42”E, 580 m, 29.ii.2012 –iii, 2012, G.B. Monteith, (25-053910 1 specimen in ANIC)GoogleMaps  ; Springbrook , north end, 28°09’47”S, 153°15’42”E, 30.viii–31.x.1997 ( T78204View Materials 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 13–23.i.2007 (T145223 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 16–26.x.2006 (T145230–T145235 6 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 21–26.x.2006 (T145237 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E 13–18.i.2007 (T145239 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–300–B, 28°09’18”S, 153°08’20.4”E, 18–23.i.2007 (T145241 1 specimen in QM)GoogleMaps  ; Gwinganna , 6 km SW by S of Tallebudgera, 28°11’00”S, 153°23’00”E, 18–23.iv.1994, D. Rentz, W. Lee & M. Upton, (25-057982 1 specimen in ANIC)GoogleMaps  ; Nichol’s Scrub , 28°11’26”S, 153°25’33”E, 1.i–4.iii.1992 ( T78111View Materials T78123View Materials 13 specimens in QM)GoogleMaps  ; Nichol’s Scrub , 28°11’26”S, 153°25’33”E, 4.iii–29.iv.1992 ( T78124View Materials T78134View Materials 11 specimens in QM)GoogleMaps  ; Nichol’s Scrub , 28°11’26”S, 153°25’33”E, 9.xii.1991 – 1.i.1992 ( T78135View Materials T78141View Materials 7 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–500–C, 28°12’36”S, 153°08’20.4”E, 22.iii–1.iv.2007 (T108394 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–500–C, 28°12’36”S, 153°08’20.4”E, 15–25.i.2007 (T145228 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–700–D, 28°12’14.4”S, 153°07’44.4”E, 14–24.i.2007 (T145226 1 specimen in QM)GoogleMaps  ; Lower Coomera , 28°12’20”S, 153°10’46”E, 9.i.1995 – 6.iv.1995 (T126027–T126029, T80219View Materials T80223View Materials 5 specimens in QM)GoogleMaps  ; Lower Coomera , 28°12’20”S, 153°10’46”E, 3.xii.1994 – 9.i.1995 ( T80215View Materials T80218, T80344 5 specimens in QM)GoogleMaps  ; Tullawallal, Binna Burra , 28°12’36”S, 153°11’22”E, 3.xii.1994 – 9.i.1995 ( T80224View Materials T80225View Materials 2 specimens in QM)GoogleMaps  ; Mount Gannon , 1.5 km SE, 28°12’39”S, 153°19’27”E, 9–12.ii.2006 (T135162–T135166 5 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 15–25.i.2007 (T145227 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 18–28.x.2006 (T145233 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 18–23.x.2006 (T145236 1 specimen in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–500–B, 28°12’43.2”S, 153°08’27.6”E, 20–25.i.2007 (T145244 1 specimen in QM)GoogleMaps  ; Mount Tallebudgera , 0.4 km NE, 28°12’55”S, 153°21’35”E, 31.i–3.ii.1997 ( T85388View Materials T85395View Materials 8 specimens in QM)GoogleMaps  ; Orams Dump , 28°13’24”S, 153°18’59”E, 26.i–2.ii.2006 (T133766–T133767 2 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–900–D, 28°13’37.2”S, 153°07’51.6”E, 20.i.2008 (T155890 1 specimen in QM)GoogleMaps  ; Mount Cougal , W Peak, 1 km NNW, 28°13’42”S, 153°18’34”E, 26.i–2.ii.2006 (T133768 1 specimen in QM)GoogleMaps  ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 19.x.2003 (T124780 1 specimen in QM)GoogleMaps  ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 1.v.2005 (T132352 1 specimen in QM)GoogleMaps  ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 29.x.2004 (T132353 1 specimen in QM)GoogleMaps  ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 26.ix.1998 – 21.v.2000 (T116858 1 specimen in QM)GoogleMaps  ; Tallebudgera Valley , falls area, 28°13’40”S, 153°18’46”E, 11.xi.1997 – 28.i.1998 ( T85383View Materials T85387View Materials 4 specimens in QM)GoogleMaps  ; Bilbrough Lookout , 1.3 km ENE, 28°13’47”S, 153°17’59”E, 4– 11.ii.2006 (T135160–T135161 2 specimens in QM)GoogleMaps  ; Springbrook Track , 300 m, 28°13’36”S, 153°18’30”E, 25–26.xi.2001 (T116836 1 specimen in QM)GoogleMaps  ; Tallebudgera Valley , 420 m, 28°13’57”S, 153°18’13”E, 4–28.iv.1997 ( T78075View Materials 1 specimen in QM)GoogleMaps  ; Numinbah Arch , 28°13’51”S, 153°14’37”E, 1.i–29.ii.1992 ( T77598View Materials T77621, T77737 T77740View Materials 28 specimens in QM)GoogleMaps  ; Numinbah Arch , 28°13’51”S, 153°14’37”E, 29.ii–11.iv.1992 ( T77730View Materials T77735, T77827 T77828View Materials 8 specimens in QM)GoogleMaps  ; Upper Tallebudgera 330 m, 28°14’00”S, 153°18’32”E, 2.i–17.ii.1995 ( T78250View Materials T78254View Materials 5 specimens in QM)GoogleMaps  ; Tallebudgera Valley , 28°14’07”S, 153°17’59”E, 28.xii.1991 – 6.iii.1992 ( T65288View Materials T65291View Materials 4 specimens in QM)GoogleMaps  ; Tallebudgera Valley , 28°14’07”S, 153°17’59”E, 6.iii–28.iv.1992 ( T65292View Materials T65296View Materials 5 specimens in QM)GoogleMaps  ; Tallebudgera Valley , 28°14’07”S, 153°17’59”E, 18.xi–28.xii.1991 ( T65297View Materials T65298View Materials 2 specimens in QM)GoogleMaps  ; Lamington National Park, IBISCA Site IQ–900–A, 28°14’2.4”S, 153°08’27.6”E, 21–26.i.2007 (T145245 1 specimen in QM)GoogleMaps  ; Numinbah Valley Natural Arch National Park ., [28°14’19”S, 153°14’17”E], 300 m, 7.vii.1978, S. Peck & J. Peck (CMNEN-00030169 1 specimen in CMNC)GoogleMaps  ; Levers Plateau , 28°18’57”S, 152°50’53”E, 6.i–29.ii.1992 ( T65299View Materials T65312View Materials 14 specimens in QM)GoogleMaps  ; Levers Plateau , 28°18’57”S, 152°50’53”E, 2.xii.1991 – 6.i.1992 ( T65313View Materials T65316View Materials 4 specimens in QM)GoogleMaps  ; Levers Plateau , 28°18’57”S, 152°50’53”E, 29.ii–11.iv.1992 ( T65317View Materials T65323View Materials 7 specimens in QM)GoogleMaps  ; Levers Plateau via Rathdowney , [28°20’00”S, 152°52’00”E], 640 m, 1.xi.1975, I.D. Naumann, (25-058465 1 specimen in QM)GoogleMaps  ; Boyds Butte , 2 km ESE,, 28°14’21”S, 153°21’17”E 13.vii.2005 (T132354–T132355 2 specimens in QM)GoogleMaps  ; New South Wales: Wiangaree State Forest Isaksson Ridge, [28°23’00”S, 153°06’00”E], 1050 m, 29.ii–3.iii.1980, A. Newton & M. Thayer, (25-057978 2 specimens in ANIC)GoogleMaps  ; Wiangaree State Forest, Sheepstation Creek , [28°23’00”S, 153°06’00”E], 600 m, 29.ii–3.iii.1980, A. Newton & M. Thayer, (25-057975–7 3 specimens in ANIC)GoogleMaps  ; Unumgar State Forest near Woodenbong, Pole Bridge road 788, [28°24’00”S, 152°40’00”E], 430 m, 2–11.i.1987, A. Newton & M. Thayer, (25-058330 8 specimens in ANIC)GoogleMaps  ; Byrrill Creek , campground, 28°26’55”S, 153°11’35”E, 30.i–1.ii.2008 (T154644 1 specimen in QM)GoogleMaps  ; Byrrill Creek , lower creek, 28°26’51”S, 153°11’39”E, 30.i–1.ii.2008 (T154642 1 specimen in QM)GoogleMaps  ; Beaury National Forest, Koreelah Range 1.8 km from Tooloom Lookout along Tucker box Road , 28°28’23”S, 152°24’04”E, 23.iii–9.v.1999, S. Lassar & C. Lemann, (25-060773 also as COL 753 specimens in AM)GoogleMaps  ; Tooloom Plateau , 14 km W Urbenville, [28°29’00”S, 152°24’00”E], 15–21.ii.1984, I.D. Naumann, (25-057980 1 specimen in ANIC)GoogleMaps  ; Whian Whian State Forest , gate, 28°33’29”S, 153°20’09”E, 29–31.i.2008 (T154636 1 specimen in QM)GoogleMaps  ; The Channon , 8 km NNE, 28°36’08”S, 153°17’48”E, 30.i–2.ii.2008 (T154643 1 specimen in QM)GoogleMaps  ; Minyon Falls , 3.5 km S, 28°38’27”S, 153°21’39”E, 29–31.i.2008 (T154637 1 specimen in QM)GoogleMaps  ; Boat harbour, site 2, 28°46’42”S, 153°19’45”E, 29.i– 31.i.2008 (T154638 1 specimen in QM)GoogleMaps  ; Lion’s Park, via Boatharbour , [28°47’00”S, 153°20’00”E], 19–20.i.1974, R.I. Storey, (25-058454 1 specimen in QM)GoogleMaps  ; Rotary Park, Lismore , site 2, 28°48’41”S, 153°17’46”E, 29–31.i.2008 (T154641 1 specimen in QM)GoogleMaps  ; Wilson Nature Reserve Lismore, [28°49’00”S, 153°16’00”E], 17.xii.1998, G. Williams & B. Williams, (25-058111 1 specimen in GWC)GoogleMaps  ; Wilson Park, Lismore , 28°50’02”S, 153°17’25”E, 29– 31.i.2008 (T154639 1 specimen in QM)GoogleMaps  ; Davis Scrub , site 2, 28°52’02”S, 153°24’20”E, 29–31.i.2008 (T154640 1 specimen in QM)GoogleMaps  ; Victoria Park, via Alstonville , [28°54’00”S, 153°24’00”E], 16.xi.1974, G.B. Monteith, (25- 058463 1 specimen in QM)GoogleMaps  ; Victoria Park near Alstonville [28°54’00”S, 153°24’00”E], 6.ii.1980, (1 specimen in AM)GoogleMaps  ; Huonbrook [28°32’00”S, 152°21’00”E], 2.iii.1964, (2 specimens in AM)GoogleMaps  ; Yabbra Scrub, Yabbra State Forest [28°31’00”S, 152°34’00”E], 14.xii.1988 (1 specimen in AM)GoogleMaps  ; Lorien, Lansdowne , [31°52’00”S, 152°29’00”E], 7.ii.1981, G. Williams & T. Williams, (25-058112 1 specimen in GWC)GoogleMaps  ; Lorien Wildlife Refuge 3km N Lansdowne, 31°45’00”S, 152°32’00”E, 16.ix.2010, G. Williams & T. Williams, (25-060880 also as COL1145 1 specimen in ANIC)GoogleMaps  ; 4 km W. Lansdowne, Lorien , [31°45’00”S, 152°32’00”E], 4.i.1987, H. Howden & A. Howden (CMNEN- 00030171 1 specimen in CMNC)GoogleMaps  ; Newbys Lane, Lansdowne , [31°47’00”S, 152°32’00”E], 14.i.1981, G. Williams & T. Williams, (25-057979 3 specimens in ANIC)GoogleMaps  ; Swans Crossing, Kerewong State Forest , 12 km WNW of Kendall, 31°36”S, 152°34”E, 24.x.1993 (1 specimen in GWC)  .

Description. Lectotype predominately black, humeri and apical edges of elytra reddish brown. Most other specimens also usually have red elytral humeri. Apical antennomere (3 rd) of antennal clubs white, basal 2 antennomeres (1 st, 2 nd) grey.

Total length: 5.0– 6.5 mm. Lectotype measurements (taken from photograph): total length 6.5 mm, elytral width 4.35 mm.

Male. Head: Width to length ratio 45: 32. Surface very finely reticulate with fine punctation. Dorsal part of eyes wide, separated by an interocular space approximately 8 times eye width (65: 8); eye canthus not dividing eye.

Prothorax: Pronotum with posterolateral angles distinct, lateral margins bordered. Surface finely punctate and finely reticulate in anterior half, finely punctate and nitid in posterior half. Pronotum width to length ratio 78: 47. Hypomeral surface finely reticulate, hypomeral stria absent.

Elytra: Surface reticulate. All striae on disc equal width, impunctate. Striae 6 and 7 not extending to the elytral base. 8th stria sometimes with a few punctures where it curves inwards posteriorly. Ratio of length of the elytra along suture to maximum elytral width 85: 100.

Legs: Protibia with ventral ridge not crenulate. Protibia with 3 teeth on the outer edge. Front edge truncate, apical digit broad and blunt apically, and apical spur absent. Mesotibia with a brush of setae apically on inner edge. Metatibia slightly bent with variously crenulated inner edge, and no round expansion apically. Metafemur slightly angulate basally on inner edge. Basal metatarsomere lobed on inner edge.

Abdomen: Pygidium with a transverse sinuate fold across base that is curved and impressed medially. Pygidial disc with a large flat, reticulate central area that occupies more than half the disc not extending under the transverse sinuate fold and separated from it by an impressed line. Ventrite 6 very finely punctate.

Pterothorax: Mesometaventral suture arched. Medial lobe of metaventrite very finely reticulate and virtually impunctate, broadly margined between mesocoxae. Lateral lobe of metaventrite very finely reticulate, finely punctate and setose. Mesoventrite very finely reticulate and notched anteriorly. Mesepimeron very finely reticulate. Metanepisternum reticulate.

Female. Protibia with front edge truncate and apical spur arising directly from front edge, apical digit absent ( Fig. 2BView FIGURE 2). Inner edge of metatibia not crenulate and inner edge of metafemur not angulate ( Fig. 2EView FIGURE 2). Mesotibia without a brush of setae apically.

Distribution. Known from rainforests, wet and dry sclerophyll forest from south of Gladstone, Queensland to Lansdowne, New South Wales ( Fig. 6AView FIGURE 6). It has been recorded from the following Interim Biogeographic Regionalisations for Australia (IBRA) bioregions: South Eastern Queensland (SEQ), the most northerly extents of New South Wales North Coast (NNC) and New England Tablelands (NET), Sydney Basin (SYB), the most easterly extents of Brigalow Belt South (BBS). Notably, there is a large gap in the distribution that extends across most of the NNC bioregion, separating records north of 29°S from those south of 31°36”S. BIOCLIM modelling suggests that most of the NNC bioregion, except for a small gap at ~ 30°S, has suitable climatic conditions for the species ( Fig. 6BView FIGURE 6). The region has been well sampled and the gap in the distribution likely reflects true absence.

Comments. Lepanus ustulatus  is a generalist species, collected in traps with a variety of baits including macropod or human dung, mushroom, and carrion (chicken or squid). Also collected at light, in berlesed leaf litter, flight intercept and window traps. In a food preference study that tested attraction to 10 different baits including a variety of dung, carrion, fermenting fruits and mushroom lures, seven specimens of L. ustulatus  were collected including six at domestic fowl dung and one specimen at vertebrate carrion ( Ebert et al. 2019).

In southern Queensland and northern New South Wales, L. ustulatus  appears to be part of a mimicry ring involving several species of scarabs and hydrophilids that are attracted to dung and other decaying organic resources ( Fikacek et al. 2014). The species listed in the potential mimicry complex also includes Onthophagus arrilla Matthews 1974  ( Coleoptera  : Scarabaeidae  : Scarabaeinae  : Onthophagini), O. sydneyensis ( Lansberge, 1874)  , Liparochrus nanus Paulian, 1980  ( Hybosoridae  ), and Austrotypus nothofagi Fikacek, Minoshima & Newton, 2014  ( Hydrophilidae  ) and are all superficially similar, most notably with distinct reddish-brown elytral humeri on the otherwise black body. Other recently described Lepanus  that occur in the region, L. burnetti Gunter & Weir, 2019  and L. meierae  , may also be part of this mimcry ring. Beyond the region, a number of other Lepanus  species also have this colour pattern including most species in the L. ustulatus  species group, although the reddish-brown humeri are less defined than in L. ustulatus  .

Lepanus ustulatus  is one of only seven species in the genus that has been assessed for the IUCN redlist and was classified as a species of least concern due to its wide distribution and expected lack of significant threats to its populations ( Weir 2013). This broad distribution of L. ustulatus  from Gladstone to Landsdowne, especially the absence of records in much of the New South Wales North Coast Bioregion, is interesting to consider. There are no consistent morphological differences that support splitting the northern and southern populations of this species and sequence data from specimens from SYB and SEQ IBRA bioregions is congruent with expected levels of intraspecific variation. The disjunction within the distribution could represent an extinction of populations in the New South Wales North Coast bioregion.

Matthews (1974) assigned a lectotype for L. ustulatus  from the series of MNHN specimens, formerly of the van Lansberge collection, and selected a male labelled “Wide Bay”. Lepanus ustulatus  is the only species within this species group that occurs in southern Queensland including the region around Wide Bay. Specimens listed in Matthews (1974) from Paluma (Wet Tropics), Eungella and near Mackay (central Queensland) represent the morphologically similar and cryptic species, Lepanus lemannae  described below. The two specimens of L. ustulatus  reported by Aristophanous (2014) from the Atherton region were not examined but based on geographic range of these superficially similar species they are probably Lepanus lemannae  .

Dorsal, lateral, ventral views, a close up of the pygidium and all labels of the lectotype of L. ustulatus  , and the dorsal habitus, pygidia, and label images of the four additional paralectotypes are published on the Muséum national d’Histoire naturelle, Paris ( France) online database.

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8675;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8676;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8677;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8678;

http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec8679;

MNHN

Museum National d'Histoire Naturelle

QM

Queensland Museum

QDPC

Queensland Primary Industries Insect Collection

ANIC

Australian National Insect Collection

AM

Australian Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus

Loc

Lepanus ustulatus ( Lansberge, 1874 )

Gunter, Nicole L. & Weir, Thomas A. 2021
2021
Loc

Lepanus ustulatus:

Matthews, E. G. 1974: 104
1974
Loc

Panelus ustulatus:

Lea, A. M. 1923: 361
1923
Loc

Epilissus ustulatus

Paulian, R. 1934: 282
Gillet, J. J. E. 1925: 5
Gillet, J. J. E. 1911: 39
Masters, G. 1886: 10
Lansberge, G. van 1874: 7
1874