Lepanus nitidus Matthews, 1974,

Gunter, Nicole L. & Weir, Thomas A., 2021, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species, Zootaxa 4923 (1), pp. 1-66: 46-50

publication ID

https://doi.org/10.11646/zootaxa.4923.1.1

publication LSID

lsid:zoobank.org:pub:41D057A7-F855-47C2-A598-8000CFD83A5B

DOI

http://doi.org/10.5281/zenodo.4537976

persistent identifier

http://treatment.plazi.org/id/1A323678-6203-222D-09D9-FF2AFB496FDD

treatment provided by

Plazi

scientific name

Lepanus nitidus Matthews, 1974
status

 

Lepanus nitidus Matthews, 1974 

( Figs. 13View FIGURE 13 A–B, G, I, M; 14D; 15D; 16D; 18A, C)

Lepanus nitidus Matthews, 1974: 110  .

Type material examined. Holotype: ♂ “The Boulders” via Babinda , Q. 4.iv.68 E. Matthews ” [17°20’00”S, 145°51’00”E] / “ Lepanus nitidus Det 1972  M, E.G. Matthews HOLOTYPE ” / “ ANIC Database No. 25 057920” (25-057920 1 specimen in ANIC)GoogleMaps  ; Paratypes: “The Boulders” via Babinda , Q. 4.iv.68 E. Matthews ” [17°20’00”S, 145°51’00”E] / “ Lepanus nitidus Det 1972  M, E.G. MatthewsGoogleMaps  PARATYPE ” or “ Lepanus nitidus Matth.  PARATYPE” (25-057921 23 specimens in ANIC); “ Daintree , N.Q. Upper Stewart Ck. 9.x.1969 R.J. Huppatz ”[16°17’00”S, 145°19’00”E], / “ Lepanus nitidus Matth  GoogleMaps  . PARATYPE ” (25-057923 1 specimen in ANIC); “ Kuranda N.Q. 1/71 GB; Black Mt. Rd. Kuranda NQ. 15.1.1971 GJ Brooks, leaf litter” [16°46’00”S, 145°36’00”E],GoogleMaps  / “ PARATYPE ” / “EM. 152” (25-057925 1 specimen in ANIC); “ Cairns N. Queensl ” [16°55’00”S, 145°46’00”E], / “ ( RIGHT) Lepanus nitidus  ♀ Det 1971 M, E.G. Matthews PARATYPE ” /GoogleMaps  “ Lectotype of Lepanus globulus (Macl.)  on same card removed T. Weir 1977” / “On permanent loan from MACLEAY MUSEUM University of Sydney” (25-057926 1 specimen in ANIC); “ Gillies H’way, N.Q. Heales Lookout 7.v.1969, Bournemissza–Ferrar” [17°13’00”S, 145°42’00”E], / “ Lepanus nitidus Matth  GoogleMaps  . PARATYPE ” (25-057922 3 specimens in ANIC); “ Forest, Innisfail 7/9/65” [17°31’00”S, 146°02’00”E] / “ Lepanus nitidus Matth  GoogleMaps  . PARATYPE ” (25-057924 1 specimen in ANIC)  .

Other material examined: Over 800 specimens from over 120 localities were also examined.

Queensland: Big Tableland , NE corner, 15°42’39”S, 145°17’12”E, 20.xii.1990 – 8.i.1991 ( T55764View Materials T55767View Materials 4 specimens in QM)GoogleMaps  ; Big Tableland , 15°43’05”S, 145°16’41”E, 21.xii.1990 – 9.i.1991 ( T55768View Materials T55771View Materials 4 specimens in QM)GoogleMaps  ; Big Tableland , 15°43’05”S, 145°16’41”E, 20.xii.1990 ( T55772View Materials 1 specimen in QM)GoogleMaps  ; Big Tableland , 15°43’05”S, 145°16’41”E, 20.xii.1990 – 8.i.1991 ( T55773View Materials T55776View Materials 4 specimens in QM)GoogleMaps  ; Mount Hartley , summit, 15°46’28”S, 145°19’24”E, 8.xi.1995 – 16.i.1996 ( T55669View Materials T55714View Materials 46 specimens in QM)GoogleMaps  ; Mount Hartley , summit, 15°46’28”S, 145°19’24”E, 8.xi.1995 ( T55715View Materials T55719View Materials 5 specimens in QM)GoogleMaps  ; Moses Creek, 4 km N by E Mount Finnigan [15°47’00”S, 145°17’00”E], 14–16.x.1980, T.A. Weir (25-058229 6 specimens in ANIC)GoogleMaps  ; Shiptons Flat [15°47’00”S, 145°14’00”E], 16–18.v.1981, J.E. Feehan (25-058227 1specimen in ANIC)GoogleMaps  ; Shiptons Flat [15°47’00”S, 145°14’00”E], 17–19.x.1980, T.A. Weir (25-058231 1 specimen in ANIC, 25-058232 4 specimens in ANIC)GoogleMaps  ; Shiptons Flat [15°47’00”S, 145°14’00”E], 16–18.v.1981, A. Calder & J.E. Feehan (25-058228, 25-058302 2 specimens in ANIC)GoogleMaps  ; Shiptons Flat , 15°47’28”S, 145°13’59”E, 275 m, 29–30.i.2012, G.B. Monteith (25-053899 8 specimens in ANIC also as COL2394 and COL2395)GoogleMaps  ; Shiptons Flat [15°47’33”S, 145°13’51”E], 22.iv.1982 ( T55782View Materials T55787View Materials 6 specimens in QM)GoogleMaps  ; Shiptons Flat [15°47’33”S, 145°13’51”E], 20–27.vii.1974 ( T49938View Materials T50008, T50080 T50100View Materials 92 specimens in QM)GoogleMaps  ; Bloomfield Road, via Helensvale, ABRS Camp , 15°47’45”S, 145°18’17”E, 2.x.1974 ( T50074View Materials T50079View Materials 6 specimens in QM)GoogleMaps  ; Bloomfield Road, via Helensvale, ABRS Camp , 15°47’45”S, 145°18’17”E, 20–27.vii.1974 ( T50012View Materials T50073View Materials 62 specimens in QM)GoogleMaps  ; Mount Hartley , 2.5 km SW, 15°47’49”S, 145°19’34”E, 8.xii.1993 – 2.ii.1994 ( T55720View Materials T55724View Materials 5 specimens in QM)GoogleMaps  ; Mount Hartley , 2.5 km SW, 15°47’49”S, 145°19’34”E, 1.i–5.iii.1994 ( T55725View Materials T55734View Materials 10 specimens in QM)GoogleMaps  ; Mount Sampson , 600–790 m, 15°48’31”S, 145°11’20”E, 27.xii.1990 – 18.i.1991 ( T55788View Materials T55802View Materials 15 specimens in QM)GoogleMaps  ; Mount Sampson , 600–790 m, 15°48’31”S, 145°11’20”E, 27.xii.1990 – 19.i.1991 ( T55827View Materials specimens in QM)GoogleMaps  ; Mount Finnigan , camp, 15°48’53”S, 145°16’28”E, 20–27.vii.1974 ( T50010View Materials T50011View Materials 2 specimens in QM)GoogleMaps  ; Mount Finnigan S1, 30km S of Cooktown [15°49’00”S, 145°17’00”E], 400 m, 2.vii.1982, S. Peck & J. Peck (25-058221 2 specimens in ANIC, 25-058220 5 specimens in ANIC)GoogleMaps  ; Mount Finnigan S1, 30 km S of Cooktown [15°49’00”S, 145°17’00”E], 400 m, 1.vii.1982, S. Peck & J. Peck (25-058219 1 specimen in ANIC)GoogleMaps  ; Annan River at The Little Forks [15°49’00”S, 145°14’00”E], 17–19.x.1980, T.A. Weir (25-058230 6 specimens in ANIC)GoogleMaps  ; Mount Misery , ascent road, 15°52’39”S, 145°12’58”E, 6.xii.1990 – 17.i.1991 ( T60076View Materials T60081View Materials 6 specimens in QM)GoogleMaps  ; Mount Misery summit, 15 °52’46”S, 145°13’32”E, 6.xii.1990 – 17.i.1991 ( T55777View Materials T55781, T60088 6 specimens in QM)GoogleMaps  ; Mount Boolbun South (summit), 15°56’50”S, 145°08’14”E, 5.xi.1995 – 11.i.1996 ( T55484View Materials T55505View Materials 22 specimens in QM)GoogleMaps  ; Mount Boolbun South (camp), 15°57’10”S, 145°08’23”E, 4–6.xi.1995 ( T55506View Materials T55527View Materials 22 specimens in QM)GoogleMaps  ; Mount Boolbun South , 15°57’10”S, 145°08’17”E, 5.xi.1995 – 11.i.1996 ( T55528View Materials T55564View Materials 37 specimens in QM)GoogleMaps  ; Emmagen Creek , 16°02’24”S, 145°27’32”E, 10 m, 29–30.v.2012, G.B. Monteith (25-057627 1 specimen in ANIC)GoogleMaps  ; Cape Tribulation area [16°04’00”S, 145°28’00”E], 21–28.iii.1984, A. Calder & T.A. Weir (25-058233–25-058234 2 specimen in ANIC)GoogleMaps  ; Cape Tribulation transect Site 1, 16°04’12”S, 145°27’54”E, 20–23.iv.1983 ( T55604View Materials T55607View Materials 4 specimens in QM)GoogleMaps  ; Cape Tribulation transect Site 2, 16°04’28”S, 145°27’33”E, 20–23.iv.1983 ( T55598View Materials T55603View Materials 6 specimens in QM)GoogleMaps  ; Mount Sorrow track, 16°04’37”S, 145°27’58”E, 30 m, 28–30.v.2012, G.B. Monteith (25-057630 3 specimens, 25- 057631 1 specimen, 25-057632 1 specimen in ANIC)GoogleMaps  ; Cape Tribulation transect Site 5, 16°04’41”S, 145°27’13”E, 1–31.i.1983 ( T55565View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 5, 16°04’41”S, 145°27’13”E, 5–9.i.1983 ( T55566View Materials T55572View Materials 7 specimens in QM)GoogleMaps  ; Cape Tribulation transect Site 5, 16°04’41”S, 145°27’13”E, 20–23.iv.1983 ( T55578View Materials T55588View Materials 11 specimens in QM)GoogleMaps  ; Cape Tribulation transect Site 5A, 16°04’41”S, 145°27’0”E, 20–23.iv.1983 ( T55589View Materials T55591View Materials 3 specimens in QM)GoogleMaps  ; Cape Tribulation transect Site 4, 16°04’45”S, 145°27’20”E, 20–23.iv.1983 ( T55592View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 3, 16°04’45”S, 145°27’33”E, 20–23.iv.1983 ( T55593View Materials T55597View Materials 5 specimens in QM)GoogleMaps  ; Cape Tribulation , [16°05’00”S, 145°29’00”E], 10 m, 15.vii.1982, S. Peck & J. Peck (25-058212 1 specimen, 25-058213 2 specimens in ANIC)GoogleMaps  ; Cape Tribulation [16°05’00”S, 145°29’00”E], 10 m, 7.i.1982, A. Walford-Huggins & M. Walford-Huggins (CMNEN-00030215 1 specimen in CMNC)GoogleMaps  ; Cape Tribulation , 40 km N of Daintree [16°05’00”S, 145°29’00”E], 10 m, 12–15.vii.1982, S. Peck & J. Peck, (25-058214 2 specimens, 25-058223 4 specimens, 25-058211 2 specimens in ANIC)GoogleMaps  ; Cape Tribulation [16°05’00”S, 145°27’00”E], 20 m, 9.v.1997, C. Reid (25-058252 1 specimen in ANIC)GoogleMaps  ; 5.0 km W of Cape Tribulation [16°05’00”S, 145°26’00”E], 780 m, 20–23.iv.1983, G.B Monteith & D.K. Yeates (25-058266 2 specimens in ANIC)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 11.ix–25.x.1998, ( T88756View Materials 1 specimen in QM)GoogleMaps  ; Thompson Creek , 16°06’31”S, 145°26’25”E, 15.iii–07.v.1998, ( T88757View Materials 1 specimen in QM)GoogleMaps  ; Noah Creek , 7 km ENE Thornton Peak [16°08’00”S, 145°26’00”E], 27.iii.1984, A. Calder & T.A. Weir (25-058235 1 specimen in ANIC)GoogleMaps  ; Oliver Creek , 16°08’10”S, 145°26’28”E, 15 m, 29–30.v.2012, G.B. Monteith (25-057628 1 specimen in ANIC)GoogleMaps  ; Oliver Creek via Daintree [16°08’19”S, 145°26’25”E], 6.iv–6.vi.1974, D. Cook (CMNEN-00030351 1 specimen in CMNC)GoogleMaps  ; Oliver Creek, via Daintree [16°08’22”S, 145°26’24”E], 4–7.vi.1974 ( T50009View Materials 1 specimen in QM)GoogleMaps  ; Cooper Creek , 16°09’10”S, 145°24’19”E, 16.iii–7.v.1998 ( T88767View Materials T88768View Materials 2 specimens in QM)GoogleMaps  ; Cooper Creek , 16°09’10”S, 145°24’19”E, 10.ii–16.iii.1998 ( T85685, T85688 2 specimens in QM)GoogleMaps  ; 1.5 km SE Mount Emmett [16°10’00”S, 145°26’00”E], 23–24.iv.1999, C.Reid (25-058241 2 specimens in ANIC)GoogleMaps  ; Candlenut Road, Daintree area , 16°11’02”S, 145°24’54”E, 20 m, 5.xii.2012, T.A. Weir (25-057666 2 specimens in ANIC)GoogleMaps  ; Daintree–section Cape Tribulation [16°12’127”S, 145°04’124”E], 85 m, 29.xii.2004 – 7.i.2005, V. Maly (CMNEN-00030219 1 specimen in CMNC)  ; Alexandra Bay , [16°12’00”S, 145°26’00”E], 50 m, 24.vi.1971, R.W. Taylor & J.E. Feehan (25-023717 specimens in ANIC)GoogleMaps  ; Lync–Haven, Daintree area , 16°12’07”S, 145°24’32”E, 35 m, 6–9.xii.2012, G.B. Monteith, (25-057650 4, 25-057651 1 specimen in ANIC)GoogleMaps  ; Lync–Haven, Daintree area , 16°12’07”S, 145°24’32”E, 35 m, 2–10.xii.2012, Weir & Monteith (25-057657 1, 25-057659 5, 25-057660 1 specimen in ANIC)GoogleMaps  ; Windsor Tableland , 46 km from main road [Mount Windsor Tableland] [16°13’00”S, 145°00’00”E], 28.xi–20.xii.1985, R.I. Storey & Brown (25-032653 1 specimen in QDPC)GoogleMaps  ; Windsor Tableland , 46 km from main road [Mount Windsor Tableland] [16°13’00”S, 145°00’00”E], 20.xii.1985 – 15.i.1986, R.I. Storey & Brown, (25-058267 3 specimens in ANIC)GoogleMaps  ; Windsor Tableland , 46 km from main road [Mount Windsor Tableland] [16°13’00”S, 145°00’00”E], 15.i–5.iii.1986, R.I. Storey & Brown (25- 058268 1 specimen in ANIC)GoogleMaps  ; Thornton Range near Daintree [16°14’00”S, 145°24’00”E], 200 m, 12–18.vii.1982, S. Peck & J. Peck (25-058222 4 specimens in ANIC)GoogleMaps  ; Alexandra Range top, 16°14’46”S, 145°26’10”E, 200 m, 28–30.v.2012, G.B. Monteith (25-057629 1 specimen in ANIC)GoogleMaps  ; Windsor Tableland via Mount Carbine [Mount Windsor Tableland] [16°15’00”S, 145°02’00”E], 25.xi.1976, R.I. Storey (25-032654 3 specimens in QDPC)GoogleMaps  ; Windsor Tableland [Mount Windsor Tableland] [16°15’00”S, 145°02’00”E], 27.xii.1976, A. Walford–Huggins &. M. Walford–Huggins (25-032655 1 specimen in QDPC)GoogleMaps  ; Mount Windsor Tableland [16°17’00”S, 145°05’00”E], 900 m, 5.xi.1976, R.W. Taylor & T.A. Weir (25-058236 1 specimen in ANIC)GoogleMaps  ; Rex Range , 16°32’02”S, 145°23’13”E, 430 m, 20–21.ix.2010, G.B. Monteith (25-060863 2 specimens in ANIC also as COL1111)GoogleMaps  ; Rex Range , 12 km S Mossman [16°34’00”S, 145°22’00”E], 400 m, 10–18.vii.1982, S. Peck & J. Peck (25-058216 1 specimen in ANIC)GoogleMaps  ; 7.5 km NNW of Kuranda [16°46’00”S, 145°37’00”E], 20.ii–20.iii.1985, R.I. Storey & K.H. Halfpapp (25-032652 1 specimen in QDPC)GoogleMaps  ; Kuranda, Black Mount Road [16°46’00”S, 145°36’00”E], 390 m, 22.vi.1971, R.W. Taylor, & J.E. Feehan (25-058239 2 specimens in ANIC)GoogleMaps  ; Black Mountain Road, 6 miles N Kuranda [16°45’22”S, 145°36’45”E], 10.ii.1971, A. Walford–Huggins & M. Walford–Huggins (CMNEN–00030213 1 specimen in CMNC)GoogleMaps  ; 7.5 km NNW of Kuranda [16°46’00”S, 145°37’00”E], 15.i–20.ii.1985, R.I. Storey & K.H. Halfpapp (25- 058253 5 specimens in ANIC)GoogleMaps  ; 7.5 km NNW of Kuranda [16°46’00”S, 145°37’00”E], 20.iii–1.v.1985, R.I. Storey & Titmarsh (25-058255 1 specimen in ANIC)GoogleMaps  ; 7.5 km NNW of Kuranda [16°46’00”S, 145°37’00”E], 20.xii.1984 – 15.i.1985, R.I. Storey & K.H. Halfpapp (25-058256 4 specimens in ANIC)GoogleMaps  ; 7.5 km NNW of Kuranda [16°46’00”S, 145°37’00”E], 20.ii–20.iii.1985, R.I. Storey & K.H. Halfpapp (25-058254 2 specimens in ANIC)GoogleMaps  ; 4 km NNW of Kuranda [16°47’00”S, 145°37’00”E], 1.v–14.vi.1985, R.I. Storey & K.H. Halfpapp (25-058263 1 specimen in ANIC)GoogleMaps  ; 4 km NNW of Kuranda [16°47’00”S, 145°37’00”E], 10.xii.1984 – 15.i.1985, R.I. Storey & K.H. Halfpapp (25-058264 1 specimen in ANIC)GoogleMaps  ; 4 km NNW of Kuranda [16°47’00”S, 145°37’00”E], 20.ix–6.xi.1984, R.I. Storey & K.H. Halfpapp (25-058265 1 specimen in ANIC)GoogleMaps  ; 4 km NNW of Kuranda [16°47’00”S, 145°37’00”E], 6.xi– 10.xii.1984, R.I. Storey & K.H. Halfpapp (25-058261 1 specimen in ANIC)GoogleMaps  ; 4 km NNW of Kuranda [16°47’00”S, 145°37’00”E], 20.ii–20.iii.1985, R.I. Storey & K.H. Halfpapp (25-058262 2 specimens in ANIC)GoogleMaps  ; Kuranda State Forest , 3 km N Kuranda [16°49’00”S, 145°38’00”E], 360 m, 25.vi–3.viii.1982, S. Peck & J. Peck (25-058218 1 specimen in ANIC)GoogleMaps  ; 6 km NW of Kuranda [16°49’00”S, 145°34’00”E], 6.xi–10.xii.1984, R.I. Storey & K.H. Halfpapp (25-058258 1 specimen in ANIC)GoogleMaps  ; Kuranda State Forest [16°49’00”S, 145°38’00”E], 360 m, 27–31.vii.1982, S. Peck & J. Peck (25-058215 1 specimen in ANIC)GoogleMaps  ; Windy Hollow Road [16°51’00”S, 145°36’00”E], 500 m, 19–21.iv.1999, C. Reid & I. Reid (25-058242 1 specimen in ANIC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 10.xii.1984 – 15.i.1985, R.I. Storey & K.H. Halfpapp (25-032656 1 specimen in QDPC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 15.i–20.ii.1985, R.I. Storey & K.H. Halfpapp (25-058257 2 specimens in ANIC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 20.ii–20.iii.1985, R.I. Storey & K.H. Halfpapp (25-058259 1 specimen in ANIC)GoogleMaps  ; 6 km SW of Kuranda [16°52’00”S, 145°37’00”E], 10.xii.1984 – 15.i.1985, R.I. Storey & K.H. Halfpapp (25-058260 3 specimens in ANIC)GoogleMaps  ; Reids Pocket [16°55’00”S, 145°36’00”E], 420 m, 17–19.iv.1999, C. Reid & I. Reid (25-058243 1 specimen in ANIC)GoogleMaps  ; Mount Murray Prior [16°56’00”S, 145°51’00”E], 7–8.xii.1998, G.B. Monteith (25-060907 3 specimens in ANIC)GoogleMaps  ; 3 mi. SW Gordonvale [17°08’00”S, 145°45’43”E], 18.viii.1974, J.G. Brooks (CMNEN-00030216–00030217 2 specimens in CMNC)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 8–9. ii.1998, S. Grove ( T73132View Materials 1 specimen in QM)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 13.iii–8.v.1998, S. Grove ( T88750, T88752,T88754 T88755View Materials 4specimens in QM)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 10.ix–24.x.1998, S. Grove ( T88751, T88753 2 specimens in QM)GoogleMaps  ; Pimm’s Block , 16°11’20”S, 145°24’18”E, 9.ii–13.iii.1998, S. Grove ( T85684View Materials 1 specimen in QM)GoogleMaps  ; Windsor Tableland , NW open forest 1, 16°12’47”S, 144°59’05”E, 9.ii–17.v.1998 ( T70773, T66788 T66799View Materials 13 specimens in QM)GoogleMaps  ; Windsor Tableland , NW open forest 1, 11.4 km past barracks, 16°12’47”S, 144°59’05”E, 24.xi.1997 – 7.ii.1998 ( T57484View Materials T57491, T60059, T60418 10 specimens in QM)GoogleMaps  ; Windsor Tableland , NW open forest 1, 11.4 km past barracks, 16°12’47”S, 144°59’05”E, 7–9.ii.1998 ( T60001View Materials T60009, T60434 T60438, T60053, T60054 16 specimens in QM)GoogleMaps  ; Windsor Tableland , NW open forest 2, 10.9 km past barracks, 16°12’54”S, 144°59’24”E, 24.xi.1997 – 7.ii.1998 ( T60035View Materials T60037View Materials 3 specimens in QM)GoogleMaps  ; Windsor Tableland , NW open forest 2, 10.9 km past barracks, 16°12’54”S, 144°59’24”E 7–9.ii.1998 ( T60419View Materials T60424View Materials 6 specimens in QM)GoogleMaps  ; Windsor Tableland , NW open forest 2, 10.9 km past barracks, 16°12’54”S, 144°59’24”E, 9.ii–17.v.1998 ( T66781View Materials T66786View Materials 6 specimens in QM)GoogleMaps  ; Gillies Road, Heales Monument [17°13’00”S, 145°42’00”E], 7.v.1969, G. Bornemissza & Ferrar ( SAMA25-027759View Materials 1 specimen in SAMA)GoogleMaps  ; Hutchinson Creek , 16°13’04”S, 145°23’57”E, 13.i–11.ii.1998, S. Grove ( T73128View Materials T73131View Materials 4 specimens in QM)GoogleMaps  ; Hutchinson Creek , 16°13’04”S, 145°23’57”E, 12.ix–23.x.1998, S. Grove ( T88758View Materials T88760, T88762 4 specimens in QM)GoogleMaps  ; Hutchinson Creek , 16°13’04”S, 145°23’57”E, 14.iii–8.v.1998, S. Grove ( T88761, T88763 T88766View Materials 5 specimens in QM)GoogleMaps  ; Fairy Creek , 16°14’16”S, 145°24’57”E, 13.iii–8.v.1998, S. Grove ( T88771View Materials T88773View Materials 3 specimens in QM)GoogleMaps  ; Fairy Creek , 16°14’16”S, 145°24’57”E, 11.ii–13.iii.1998, S. Grove ( T85687View Materials 1 specimen in QM)GoogleMaps  ; Buchanan Creek , 16°14’39”S, 145°25’54”E, 12.iii–8.v.1998, S. Grove ( T88769View Materials 1 specimen in QM)GoogleMaps  ; Buchanan Creek , 16°14’39”S, 145°25’54”E, 12.ix– 23.x.1998, S. Grove ( T88770View Materials 1 specimen in QM)GoogleMaps  ; Alexandra Range , road at summit, 16°14’46”S, 145°25’51”E, 13.x.1980 ( T55621View Materials 1 specimen in QM)GoogleMaps  ; Windsor Tableland , barracks, 16°15’46”S, 145°02’31”E, 17–19.xi.2001 (T111985–T111988 4 specimens in QM)GoogleMaps  ; Windsor Tableland , barracks, 16°15’46”S, 145°02’31”E, 23–25.xi.1997 ( T60000View Materials 1 specimen in QM)GoogleMaps  ; Windsor Tableland , barracks, 16°15’46”S, 145°02’31”E, 8–9.ii.1998 ( T60042View Materials 1 specimen in QM)GoogleMaps  ; Bellenden Ker Range, Cableway Base Station [17°16’00”S, 145°54’00”E], 100 m, EarthwatchGoogleMaps  , Queensland Museum (25-032810 1 specimen in QDPC)  ; Bellenden Ker Range, Cableway Base Station [17°16’00”S, 145°54’00”E], 100 m, EarthwatchGoogleMaps  , Queensland Museum , 17.x–9.xi.1981 (25-058224 10 specimens in ANIC, 25- 058225 1 specimen in ANIC, 25-058226 1 specimen in ANIC)  ; Windsor Tableland ANZSES Site 9, 16°16’55”S, 145°03’56”E, 27.xii.1988 – 8.i.1989 ( T55803View Materials 1 specimen in QM)GoogleMaps  ; The Boulders via Babinda [17°20’00”S, 145°51’00”E], 11.iv.1973, R.I. Storey (25-032649 2 specimens in QDPC, CMNEN-00030218 1 specimen in CMNC)GoogleMaps  ; The Boulders via Babinda [17°20’00”S, 145°51’00”E], 4.iv.1968, E. Matthews ( SAMA25-027757View Materials 6 specimens in SAMA)GoogleMaps  ; The Boulders via Babinda [17°20’00”S, 145°51’00”E], 11.iv.1973, R.I. Storey (25-058515 3 specimens in QM)GoogleMaps  ; The Boulders , 7 km W of Babinda [17°20’00”S, 145°51’00”E], 7.vii.1984, R.B. Halliday (25- 058238 1 specimen in ANIC)GoogleMaps  ; Devils Thumb transect Site 4, 16°23’39”S, 145°19’02”E, 26.xii.1989 – 15.i.1990, ( T55467View Materials 1 specimen in QM)GoogleMaps  ; High Falls , 16°24’18”S, 145°18’35”E, 25.i–12.ii.1996, ( T55626View Materials 1 specimen in QM)GoogleMaps  ; Mount Spurgeon open forest, trap 4, 16°27’17”S, 145°11’33”E, 19.xi.1997 – 7.ii.1998, ( T60043View Materials T60044, T60057 3 specimens in QM)GoogleMaps  ; Mount Spurgeon open forest, trap 5, 16°27’25”S, 145°11’37”E, 19.xi.1997 – 7.ii.1998 ( T60030, T60058 2 specimens in QM)GoogleMaps  ; Polly Creek Garradunga, 17°27’22”S, 146°01’12”E, 52 m, 13.i–4.ii.2010, J. Hasenpusch (25-060862 1 specimen in ANIC also as COL1110)GoogleMaps  ; Polly Creek Garradunga, 17°27’32”S, 146°01’08”E, 40 m, 16–17.ix.2010, G.B. Monteith (25-060858 1 specimen in ANIC also as COL1109)GoogleMaps  ; Mount Spurgeon open forest, trap 7, 16°27’37”S, 145°11’45”E, 20.xi.1997 – 7.ii.1998 ( T60039View Materials T60041View Materials 3 specimens in QM)GoogleMaps  ; Mount Spurgeon open forest, trap 8, 16°27’40”S, 145°11’54”E, 20.xi.1997 – 7.ii.1998 ( T60010View Materials T60012View Materials 3 specimens in QM)GoogleMaps  ; Mount Spurgeon open forest, trap 9, 16°27’50”S, 145°11’58”E, 20.xi.1997 – 7.ii.1998 ( T60013View Materials T60016View Materials 4 specimens in QM)GoogleMaps  ; Mount Spurgeon , 3 km S, open forest, trap 10, 16°27’59”S, 145°12’02”E, 20.xi.1997 – 7.ii.1998 ( T60052View Materials 1 specimen in QM)GoogleMaps  ; Mossman Bluff Site 5, 16°28’25”S, 145°17’49”E, 20.xii.1989 – 15.i.1990 ( T55461View Materials T55465View Materials 5 specimens in QM)GoogleMaps  ; Mossman Bluff Site 5, 16°28’25”S, 145°17’49”E, 16–30.xii.1988 ( T55473View Materials T55479View Materials 7 specimens in QM)GoogleMaps  ; Mossman Bluff Site 4, 16°28’29”S, 145°18’09”E, 20.xii.1989 – 15.i.1990 ( T55460View Materials 1 specimen in QM)GoogleMaps  ; Mossman Bluff Site 4, 16°28’29”S, 145°18’09”E, 1–16.i.1989 ( T55480View Materials T55482View Materials 3 specimens in QM)GoogleMaps  ; Rex Range , summit, 16°32’12”S, 145°22’44”E, 35930 ( T66787View Materials 1 specimen in QM)GoogleMaps  ; Kingfisher Park 16°35’31”S, 145°20’27”E, 6–9.ii.1998 ( T57492View Materials T57494, T60045 T60051View Materials 10 specimens in QM)GoogleMaps  ; Black Mountain , lower slopes, 16°39’52”S, 145°30’04”E, 2.xii.1997 – 6.ii.1998 ( T60031View Materials T60034View Materials 4 specimens in QM)GoogleMaps  ; Mount Formartine South , 16°43’48”S, 145°36’45”E, 23–24.xi.1990 ( T55345View Materials T55346View Materials 2 specimens in QM)GoogleMaps  ; Saddle Mountain , summit, 16°49’11”S, 145°39’42”E, 10.xii.1995 – 2.ii.1996 ( T55348View Materials T55367View Materials 20 specimens in QM)GoogleMaps  ; Saddle Mountain , summit, 16°49’11”S, 145°39’42”E, 3.xi–10.xii.1995 ( T55368View Materials T55377View Materials 10 specimens in QM)GoogleMaps  ; Mount Murray Prior , 16°55’59”S, 145°50’59”E, 7.xii.1995 ( T55436View Materials 1 specimen in QM)GoogleMaps  ; Mount Murray Prior , 16°55’59”S, 145°50’59”E, 31.x–8.xii.1995 ( T55437View Materials T55445, T70911 T70923View Materials 22 specimens in QM)GoogleMaps  ; Pine Creek Tower, 16°59’36”S, 145°50’07”E, 12.ix–20.x.1991 ( T55446View Materials T55447View Materials 2 specimens in QM)GoogleMaps  ; Bones Knob , 3 km W, 17°13’55”S, 145°25’05”E ( T55347View Materials 1 specimen in QM)GoogleMaps  ; Bellenden Ker Base Station , 17°16’36”S, 145°53’54”E, 30.xi–2.xii.1998 ( T70878View Materials T70885View Materials 8 specimens in QM)GoogleMaps  ; Graham Range , 350 m, 17°16’37”S, 145°57’48”E, 1.xi–8.xii.1995 ( T55310View Materials T55313View Materials 4 specimens in QM)GoogleMaps  ; Graham Range, W. Base , 17°17’13”S, 145°57’47”E, 9.iv.1979 ( T55314View Materials 1 specimen in QM)GoogleMaps  ; Stone Creek , 17°27’40”S, 146°01’04”E, 1.xi.1995 – 6.ii.1966, J. Hasenpusch ( T55315View Materials 1 specimen in QM)GoogleMaps  ; Stone creek , 17°27’40”S, 146°01’04”E, 1.x–1.xi.1995, J. Hasenpusch ( T55829View Materials 1 specimen in QM)GoogleMaps  ; Bramston Beach , 5 km SW, 17°22’57”S, 145°59’06”E, 5–11.ii.1998 ( T60027View Materials T60029View Materials 3 specimens in QM)GoogleMaps  ; Henrietta Creek, Palmerstone National Park [17°34’00”S, 145°45’00”E], 22–24.iv.1973, R.I. Storey (25-032811 1 specimen in QDPC)GoogleMaps  ; South Johnstone Research Station [17°36’00”S, 145°58’00”E], 29.xi.1989 – 14.i.1990, K.H. Halfpapp (25- 032645 1 specimen in QDPC)GoogleMaps  ; South Johnstone Research Station [17°36’00”S, 145°58’00”E], vi–xi.1990, K.H. Halfpapp (25-032646 1 specimen in QDPC)GoogleMaps  ; South Johnstone Research Station [17°36’00”S, 145°58’00”E], 12.ix– 11.x.1990, K.H. Halfpapp (25-032647 3 specimens, 25-032648 1 specimen in QDPC)GoogleMaps  ; Wooroonooran National Park, Patterson Sector, Henrietta Creek Campground, 17°35’56”S, 145°45’32”E, 550 m, 22.i.2000, R.A. Anderson (CMNEN-00030212 1 specimen in CMNC)GoogleMaps  ; Wooroonooran National Park , 17°36’17”S, 145°45’38”E, 400 m, 8– 9.ix.2004, L. Masner (CMNEN-00030214 1 specimen in CMNC)GoogleMaps  ; Palmerston National Park , E margin, 17°36’40”S, 145°45’48”E, 2.xi–10.xii.1995 ( T55187View Materials T55188View Materials 2 specimens in QM)GoogleMaps  ; Palmerston National Park , E margin, 17°36’40”S, 145°45’48”E, 10.xii.1995 – 7.ii.1996 ( T55189View Materials T55209View Materials 21 specimens in QM)GoogleMaps  ; Palmerston National Park , E margin, 17°36’40”S, 145°45’48”E, 1.xii.1993 – 25.ii.1994 ( T55210View Materials T55219View Materials 10 specimens in QM)GoogleMaps  ; Wooroonooran National Park, Patterson Sector, Crawfords Lookout , 17°36’46”S, 145°46’36”E, 550 m, 22.i.2000, R.A. Anderson (CMNEN-00030211 1 specimen in CMNC)GoogleMaps  ; Palmerston National Park [17°37’00”S, 145°47’00”E], 1.iv.1968, E. Matthews ( SAMA25-027760View Materials 1 specimen in SAMA)GoogleMaps  ; McNamee National Park [17°39’00”S, 145°51’00”E], 250 m, 16–18.iv.1999, C. Reid & I. Reid (25-058244 1 specimen in ANIC)GoogleMaps  ; Laceys Creek, via Mission Beach [17°49’00”S, 146°04’00”E], 14.v.1975, R.I. Storey (25-058516 3 specimens in QM)GoogleMaps  ; S3 Mission Beach 17°50’00”S, 146°06’00”E], 10 m, 4.iii–1.iv.1996, M. Cermak (25-058245 1 specimen in ANIC)GoogleMaps  ; S3View Materials Mission Beach [17°50’00”S, 146°06’00”E], 10 m, 1–29.iv.1996, M. Cermak (25-058246 1 specimen in ANIC)GoogleMaps  ; Lacey’s Creek, via El Arish , 17°51’10”S, 146°03’17”E, 19–20.vi.1974 ( T55294View Materials T55295, T60082 T60083View Materials 4 specimens in QM)GoogleMaps  ; Mission Beach , [17°52’00”S, 146°06’00”E], 9–10.v.1974, R.I. Storey (25-032651 1 specimen in QDPC)GoogleMaps  ; S1View Materials Mission Beach , [17°52’00”S, 146°04’00”E], 40 m, 29.i–4.iii.1996, M. Cermak (25-058237 1 specimen, 25-058249 2 specimens in ANIC)GoogleMaps  ; S1View Materials Mission Beach [17°52’00”S, 146°04’00”E], 40 m, 2–29.i.1996, M. Cermak (25-058247 1 specimen in ANIC)GoogleMaps  ; S1View Materials Mission Beach [17°52’00”S, 146°04’00”E], 40 m, 4.xii.1995 – 2.i.1996, M. Cermak (25- 058248 1, 25-058250 1 specimen in ANIC)GoogleMaps  ; S1View Materials Mission Beach [17°52’00”S, 146°04’00”E], 40 m, 4.iii–1.iv.1996, M. Cermak (25-058251 1 specimen in ANIC)GoogleMaps  ; Laceys Creek , 10 km SE El Arish [17°53’00”S, 146°04’00”E], 40 m, 23.vi–5.viii.1982, S. Peck & J. Peck (25-058217 4 specimens in ANIC)GoogleMaps  ; 7 km ENE Tully [17°55’00”S, 145°59’00”E], 50 m, 16–25.iv.1999, C. Reid & I. Reid (25-058240 2 specimens in ANIC)GoogleMaps  ; Cardwell Gap , 18°32’0”S, 146°11’08”E, 4–12.ii.1998 ( T60038View Materials 1 specimen in ANIC)GoogleMaps  .

Description. Holotype uniformly light brown in colour. Other specimens range from light brown to dark brown verging on black; sometimes with paler humeri. Antennal clubs with apical antennomere (3rd) yellowish white and basal 2 antennomeres (1 st, 2 nd) grey.

Total length: 3.6–4.4 mm. Holotype measurements: total length 4.2 mm elytral width 2.8 mm.

Male. Head: Width to length ratio 36: 25. Almost V–shape between clypeal teeth. Surface finely punctate with very fine setae. Dorsal part of eyes moderate, separated by an interocular space approximately 13 times eye width (26: 2).

Prothorax: Pronotum smooth, nitid, with very fine punctation towards the anterior and finely punctate posteriorly. Hypomeral surface finely reticulate. Hypomeral stria absent. Pronotum width to length ratio 54: 38.

Elytra: Smooth and nitid. Intervals finely punctate and with extremely fine setae. Ratio of length of the elytra along suture to maximum elytral width 70: 75.

Legs: As per L. nitidus  species group. Ridge on ventral side of protibia apically more-or-less joined to the second tooth by a fine carina.

Abdomen: Rest of pygidium surrounding central disc is finely punctate and setose with fine reticulations basally.

Pterothorax: Lateral lobe of metaventrite finely reticulate and punctate, inner border beside mesocoxa anteriorly widened. Medial lobe of metaventrite very finely punctate. Metaventrite with very small tubercle located approximately halfway between mesocoxae and metacoxae (slightly more anterior than in L. dichrous  (see Fig. 13View FIGURE 13 I–J)).

Male genitalia: Aedeagus with left paramere excised apically. As in Fig. 16DView FIGURE 16

Female. Metaventral tubercle absent.

Distribution. Known from rainforest in Wet Tropics (WET) IBRA bioregion [Zones 1, 2, 3, 4, 6, 7, 8, 9, 10, 11, 12, and 13] from Rossville to Cardwell ( Fig. 18AView FIGURE 18). Almost all specimens have been collected at elevations below 500 m. Species distribution estimates from BIOCLIM models shown in Fig. 18CView FIGURE 18.

Comments. Lepanus nitidus  is a generalist species and has been collected at pitfall traps baited with macropod, cow or human feces, mushrooms, vertebrate carrion, and invertebrate carrion (shrimp, earthworm, cricket and mealworm). It has also been collected using a variety of other methods including flight intercept and malaise traps and in berlesed leaf and log litter.

A specimen of L. nitidus  from Shiptons Flat (25-053899) was included in the phylogeny of Gunter et al. (2019). Here, additional specimens from this locality, Rex Range and Polly Creek were sequenced (see Fig. 1View FIGURE 1). In both analyses, L. nitidus  was recovered as sister to L. kulki  within a larger clade containing all five species within the Lepanus nitidus  species group. This molecular relationship is strongly supported and congruent with the morphological similarity between the two species.

In the altitudinal transect study of Monteith (1985a, 1985b) along 11 sites from sea level to 780 m at Cape Tribulation L. nitidus  was taken only at sites below 400 m. Matthews (1974) lists specimens from high elevations (Mount Haig, 1220 m (25-058517, 25-058518) and Tully Falls, 760 m (25-058505–08)), however these are the morphologically similar species Lepanus vangerweni  and L. dichrous  respectively. The elevational survey of Aristophanous (2014) collected 557 speciemens in the L. nitidus  species group within the Atherton, Carbine and Windsor transects, however did not identify them beyond “ Lepanus nitidus  / dichrous  ”. Given the broad and often overlapping distribution of species within the group, we refrain from making inferences from this data.

ANIC

Australian National Insect Collection

QM

Queensland Museum

QDPC

Queensland Primary Industries Insect Collection

SAMA

South Australia Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus

Loc

Lepanus nitidus Matthews, 1974

Gunter, Nicole L. & Weir, Thomas A. 2021
2021
Loc

Lepanus nitidus Matthews, 1974: 110

Matthews, E. G. 1974: 110
1974