Nannaria equalis Chamberlin, 1949

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

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Nannaria equalis Chamberlin, 1949


Nannaria equalis Chamberlin, 1949   Fig. 54 Vernacular name: "The Knoxville Twisted-Claw Millipede" View Figure 54

Nannaria equalis   Chamberlin, 1949: 4, fig. 4. Chamberlin and Hoffman 1958: 40. Hoffman 1999: 366. Marek et al. 2014: 37. Means et al. 2021: S69-S70.

Material examined.

United States - Tennessee • 1 ♂; Knox County, site 2 mesic cove hardwoods; from label: 17.233902E 3993447N; 10 May 2005; J. Sevier leg.; VMNH NAN0335. For detailed collection data see Suppl. material 7   .


Adult males of N. equalis   are distinct from other Nannaria   and the nearby N. monsdomia   sp. nov. and N. scutellaria   based on the following combination of characters: Gonopods. Gonopodal acropodite straight before apex, not slightly curving before apex as in N. monsdomia   sp. nov. Distal zone much reduced, rounded, simple, not with small, lobed lateral flange as in N. monsdomia   sp. nov., or bent at 90° with pronounced medial flange as in N. scutellaria   . Acropodite with medial swelling, lacking in N. scutellaria   . Telopodite basal zone height ca. 1/2 length of acropodite, not > 2/3 length as in N. monsdomia   sp. nov. or < 1/3 length as in N. scutellaria   . Prefemur with large, straight prefemoral process (Fig. 54A View Figure 54 , red arrow), not curving medially, crossing acropodite, and expanding before tip, as in N. monsdomia   sp. nov., or curving laterally as in N. scutellaria   . Prefemoral spine sharp, curving laterally (Fig. 54B View Figure 54 , red triangle), not widely separated from prefemoral process and rounded, as in N. monsdomia   sp. nov., or lacking as in N. scutellaria   . Acropodite appearing as straight line in medial view, therefore the medial view was not illustrated. Color. Chamberlin reports that the paranota of Nannaria equalis   are yellow in color, though this would be unique in the Nannaria   and has not been confirmed by the authors. It is likely Chamberlin was referring to a preserved specimen whose color had faded in alcohol.


♂ ( VMNH, NAN0335): BL = 27.2, CW = 4.0, IW = 1.7, ISW = 0.9, B11W = 4.7, B11H = 2.8.


No known variation.


Known only from the type locality (Tennessee: Knox County; Suppl. material 7; Fig. 126 View Figure 126 ). Distribution area: N/A; status: MRE.


Chamberlin (1949) gives no information on the ecology of N. equalis   .


Chamberlin (1949) provided no etymology for the name equalis, though it is reasonable to assume that it is in reference to the prefemoral process nearly equaling in length the acropodite, as Chamberlin mentions in his description.

Type locality.

United States, Tennessee, Knox County, Knoxville.


In the original publication Chamberlin (1949: 4) designated a male holotype and female paratype (originally as an “allotype”).














Nannaria equalis Chamberlin, 1949

Means, Jackson C., Hennen, Derek A. & Marek, Paul E. 2021

Nannaria equalis

Chamberlin 1949