Aclista elegantula, Macek, 2005

Macek, Jan, 2005, Revision of the Central European species of Aclista (Diapriidae, Hymenoptera). Part II. Aclista insolita Nixon, 1957 Aclista dubia Kieffer, 1909 and similar species, Acta Entomologica Musei Nationalis Pragae 45, pp. 183-197 : 95-96

Aclista elegantula sp. nov.

( Figs. 6, 13 , 20 , 27 , 34 , 41 )

Type locality. Czech Republic, Králický Sněžník Mt., Horní Morava.

Type material. HOLOTYPE: ♀, ‘ Czech Republic, NR Králický Sněžník, Horní Morava (5866), 24.viii.2001, J. Macek lgt. et det.’ ( NMPC) . ALLOTYPE: J, ‘ Czech Republic, NR Králický Sněžník, Mlýnský potok (brook) (5866), 2.viii.2001, J. Macek lgt. et det.’ ( NMPC) . PARATYPES: CZECH REPUBLIC: BOHEMIA or., NR Králický Sněžník, Mlýnský potok brook (5866), 20.viii.2001, 1 ♀ ; same locality, 24.x.2001, 1 ♀ ; Horní Morava (5866), 24.viii.2001, 2 ♀♀ ; BOHEMIA or., Orlické hory Mts., NR Kačerov (5764), 18.vii.-5.viii.1994, 1 ♀ ; Pod Sfingou hill (5764), 19.viii.2003, 2 ♀♀ ; Bedřichovka (5664), 15.-24.ix.1993, 1 J . SLOVAKIA centr.: Nízké Tatry Mts., Štiavnica (7084), 6.viii.1984, 1 ♀ ; Pohorelá (7186), 31.vii.1984, 1 ♀ ; NR Ohniště (7084), 3.viii.1984, 1 J, all J. Macek lgt. & det. ( NMPC) . AUSTRIA: STEIERMARK, Vockenberg, Furtner Teich , 900 m a.s.l., 3.viii.1971, 4 ♀♀ 3 JJ ; KÄRNTEN, St. Georgen, Längsee , 700 m a.s.l., 7.viii.1977, 1 ♀, all M. Fischer lgt., J. Macek det. ( NMPC, NHMW) ; Zell am See , 3.x.1988, 1 ♀, R. Danielsson lgt., J. Macek det. ( MZLU) .

Description. Female (holotype). Length 3.3 mm; colour black; legs, mouthparts and tegulae yellowish; antennae yellowish brown, pedicel brownish.

Head in dorsal view strongly transverse, wider than mesosoma, in lateral view higher than long and in frontal view strongly transverse, with smooth, lustrous face; genae strongly converging towards mouthparts; antennal sockets slightly prominent, smooth below, with toruli separated by shallow cleft; vertex convex; temples strongly receding backwards; ocelli small, OOL> POL; eyes pubescent, in longest diameter longer than malar space; face lustrous; subantennal furrows short and distinct; epistomal sulcus distinct; tentorial pits placed in deep hollows; mouth aperture shorter than malar space; clypeus (in lateral view) moderately convex in middle, rugose; mandibles sickle-shaped and widely overlapping; antennae 15-segmented, long, slender, submoniliform; scape very slender, cylindrical, bowed, three time as long as F1; F1-F8 cylindrical, becoming gradually shorter towards apex; F1 three times as long as wide; F8 one and half times as long as wide, F9-F12 moniliform, with erect short pubescence, F13 subconical, not larger than F12.

Mesosoma wide, convex, slightly narrower than head, with long decumbent pubescence; pronotal collar indistinct, pronotal shoulders rounded; epomia short; lateral pronotum strongly impressed, lustrous, not perceptible from above; rim connecting pronotal spiracle with tegula obliterated; mesoscutum convex, as long as wide, rounded anteriorly; notauli continuous, deep and parallel posteriorly; scutellum convex, smooth, with subrectangular anterior fovea; lateral foveae smooth, with a very fine tuft of pubescence; mesopleura smooth, with large scrobe in middle; dorsellum with weak tubercle medially; sides of metanotum smooth and hollowed; propodeum transverse, with smooth, sparsely pubescent, dorsal surface; posterior rim of propodeum large and raised; medial keel of propodeum simple; inner plicae converging posteriorly and not prominent behind; outer plicae diverging and projecting posteriorly; space between median keel and inner plicae with short posterior keel.

Wings infumate; marginal vein longer than half of parastigma; radial cell twice as long as marginal vein; postmarginal vein far surpassing radial cell; stigmal vein oblique to marginal vein, slightly shorter than marginal vein, slightly curved.

Petiole subcylindrical, longitudinally ribbed, three times as long as wide; gaster fusiform, sparsely pubescent all over dorsal surface; base of macrotergite as wide as petiole; basal sculpture of macrotergite with distinct medial furrow extending as far as one third of macrotergite and with short lateral striation; prepygidial segments annular, very narrow, tightly abutting each other; pygidium as long as prepygidium, apical segment straight.

Male (allotype). Differs from female as follows: (i) antennae 14-segmented, filiform, with slender flagellar segments; (ii) scape slender, twice as long as F1; (iii) F1 emarginate on basal half; (iv) pubescence of antennae as long as flagellomeres wide, semidecumbent and dense. Differential diagnosis. Based on the morphological characters, A. elegantula sp. nov. is similar to A. subtilis sp. nov. but differs in the following combination of characters (compare also the differential diagnosis of the latter species): (i) habitus slender, mesosoma narrower than head (dorsal view); (ii) scape very slender, seven times as long as wide; (iii) antennae blackish with black scape; (iv) propodeum qudrate with large raised posterior ledge; (vi) petiole long, three times as long as wide, longitudinally ribbed; (vii) scape in males twice as long as F1; and (viii) F 3 in males emarginate with a short tyloid at the base.

Variability. The examined material shows no apparent variation.

Etymology. Adjective elegantulus (Latin) = elegant; referring to the general appearence of the species.

Bionomy. Hosts unknown; flight period from July to October.

Distribution. So far known from Austria, the Czech Republic and Slovakia.

Figures

Figs. 1-7. Head, frontal view (female). 1 – Aclista insolita Nixon, 1957; 2 – A. vernalis sp. nov.; 3 – A. dubia Kieffer, 1909; 4 – A. longistilus Kieffer, 1909; 5 – A. marginalis Kieffer, 1909; 6 – A. elegantula sp. nov.; 7 – A. subtilis sp. nov. Scale = 1 mm.

Figs. 15-21. Metasoma, dorsal view (female). 15 – Aclista insolita Nixon, 1957; 16 – A. vernalis sp. nov.; 17 – A. dubia Kieffer, 1909; 18 – A. longistilus Kieffer, 1909; 19 – A. marginalis Kieffer, 1909; 20 – A. elegantula sp. nov.; 21 – A. subtilis sp. nov. Scale = 2 mm.

Figs. 22-28. Antennae, lateral view (female). 22 – Aclista insolita Nixon, 1957; 23 – A. vernalis sp. nov.; 24 – A. dubia Kieffer, 1909; 25 – A. longistilus Kieffer, 1909; 26 – A. marginalis Kieffer, 1909; 27 – A. elegantula sp. nov.; 28 – A. subtilis sp. nov. Scale = 2 mm.

Figs. 29-42. 29-35 – fore wing venation (female). 29 – Aclista insolita Nixon, 1957; 30 – A. vernalis sp. nov.; 31 – A. dubia Kieffer, 1909; 32 – A. longistilus Kieffer, 1909; 33 – A. marginalis Kieffer, 1909; 34 – A. elegantula sp. nov.; 35 – A. subtilis sp. nov. 36-42 – proximal part of antenna, lateral view (male). 36 – A. insolita; 37 – A. vernalis sp. nov.; 38 – A. dubia; 39 – A. longistilus; 40 – A. marginalis; 41 – A. elegantula sp. nov.; 42 – A. subtilis sp. nov. Scale = 1 mm (Figs. 29-35) and 0.7 mm (Figs. 36-42).