Aclista subtilis, Macek, 2005

Macek, Jan, 2005, Revision of the Central European species of Aclista (Diapriidae, Hymenoptera). Part II. Aclista insolita Nixon, 1957 Aclista dubia Kieffer, 1909 and similar species, Acta Entomologica Musei Nationalis Pragae 45, pp. 183-197 : 93-94

Aclista subtilis sp. nov.

( Figs. 7, 14, 21 , 28 , 35, 42 )

Type locality. Czech Republic, Sokolov district, Svatý Jiří strip mine.

Type material. HOLOTYPE: ♀, ‘ Czech Republic , Sokolov distr. , Starý Jiří (5742), 6.vii.2001, Macek lgt. et det.’ ( NMPC) . ALLOTYPE: J, ‘ Czech Republic, Bohemia cent., Karlštejn (6051), 10.viii.1986, Macek lgt. et det.’ ( NMPC) . PARATYPES: CZECH REPUBLIC: BOHEMIA or., Orlické hory Mts. , NR Kačerov (5764), 28.vi.-16.vii.1994, 2 ♀♀ ; BOHEMIA centr., Klíčava river (5950), 16.vii.1986, 1 J ; Čelákovice (5854), 25.ix.-9.x.1991, 1♀. MORAVIA mer., NP Podyjí, NR Braitava (7160), 8.ix.1993, 1 J, all J. Macek lgt. et det. ( NMPC) . SLOVAKIA centr.: Slovenské Rudohorie Mts., NR Dobročský prales (7284), 30.vii.1986, 1 J ; J. Macek lgt. & det. ( NMPC) . AUSTRIA: STEIERMARK, Mariahof, Vockenberg , 900 m a.s.l., 3.viii.1969, 1 J, M. Fischer lgt., J. Macek det. ( NHMW) . POLAND centr.: Warszawa, NR Park J Sobieskiego , 7.-16.x.1980, 1 J ; 26.viii.1980, 2 JJ, J. Ekipa lgt., J. Macek det. ( NMPC) .

Description. Female (holotype). Length 2.4 mm; colour dark brown; legs, mouthparts and tegulae yellowish; antennae brownish, scape yellowish brown.

Head in dorsal view strongly transverse and not wider than mesosoma, with moderately prominent antennal sockets; toruli distant from each other, with shallow cleft between them; vertex highly convex; temples strongly receding behind; ocelli small, OOL> POL; longest eye diameter longer than malar space; eyes large, pubescent; head in lateral view higher than long, in frontal view suboval with smooth and lustrous face; genae strongly converging towards mouthparts; antennal sockets smooth below; subantennal furrows slightly indicated; epistomal sulcus developed; tentorial pits placed in deep hollows; mouth aperture shorter than malar space; clypeus (in lateral view) convex in middle, finely rugose; mandibles strongly sickle-shaped, overlapping; antennae long, slender, 15-segmented, submoniliform; scape slender, cylindrical, bowed, 2.5 times as long as F1; F1-F5 cylindrical, becoming gradually shorter towards apex of flagellum; F1 four times as long as wide; F6-F12 moniliform, with erect pubescence, F13 subconical, not larger than the preceding segment.

Mesosoma wide, convex, slightly narrower than head, with long decumbent pubescence; pronotal collar indistinct, pronotal shoulders rounded; epomia short; lateral pronotum strongly impressed, smooth and lustrous, not visible from above; sulcus connecting pronotal shoul- der to tegula with indistinct upper rim; mesoscutum convex, wider than long, steeply falling anteriorly; notauli continuous, deep and diverging posteriorly; scutellum convex, smooth, with transverse, rounded anterior fovea; lateral foveae smooth, with a very fine tuft of pubescence; mesopleuron smooth, with large scrobe in middle; dorsellum not prominent medially; sides of metanotum smooth and concave; propodeum transverse with smooth sparsely pubescent dorsal surface; posterior rim of propodeum distinct and raised; medial keel of propodeum simple; inner plicae converging and not projecting posteriorly; space between median keel and inner plicae with short posterior keel.

Wings slightly infumate; marginal vein longer than half of parastigma; radial cell closed, long, twice as long as marginal vein; postmarginal vein far surpassing radial cell; stigmal vein oblique to marginal vein, slightly shorter than marginal vein, straight.

Petiole subcylindrical, smooth, with irregular longitudinal rugosity, twice as long as wide; gaster fusiform, very sparsely pubescent all over dorsal surface; base of macrotergite as wide as petiole; basal sculpture of macrotergite with distinct medial furrow reaching as far as one fourth of macrotergite and with short lateral striation; prepygidial segments ring-like, very narrow, tightly abutting each other; pygidium slightly longer than prepygidium, apical segment slightly upcurved.

Male (allotype). Differs from female as follows: (i) antennae 14-segmented, filiform, with slender flagellar segments; (ii) scape slender, slightly longer than F1; (iii) F1 cylindrical with prominent tyloid on basal third; (iv) pubescence of antennae short, semidecumbent and dense.

Variability. The examined material shows no apparent variation.

Differential diagnosis. Based on the morphological characters, A. subtilis sp. nov. resembles Aclista elegantula sp. nov. (see below) but differs in the following combination of characters: i) habitus stouter, mesosoma as wide as head (dorsal view); (ii) scape shorter and stouter, six times as long as wide; (iii) antennae brownish with a pale scape; (iv) propodeum rectangular with a narrow raised posterior ledge; (vi) petiole stout, at most 2.5 times as long as wide, longitudinally rugose; (vii) scape in males nearly as long as F1; viii) F 3 in males not emarginate and with a long and straight tyloid at the base.

Etymology. Adjective subtilis (Latin) = subtle; referring to the general appearence of the species.

Bionomy. Hosts unknown; flight period from July to October

Distribution. So far known from Austria, the Czech Republic, Poland and Slovakia.

Figures

Figs. 1-7. Head, frontal view (female). 1 – Aclista insolita Nixon, 1957; 2 – A. vernalis sp. nov.; 3 – A. dubia Kieffer, 1909; 4 – A. longistilus Kieffer, 1909; 5 – A. marginalis Kieffer, 1909; 6 – A. elegantula sp. nov.; 7 – A. subtilis sp. nov. Scale = 1 mm.

Figs. 22-28. Antennae, lateral view (female). 22 – Aclista insolita Nixon, 1957; 23 – A. vernalis sp. nov.; 24 – A. dubia Kieffer, 1909; 25 – A. longistilus Kieffer, 1909; 26 – A. marginalis Kieffer, 1909; 27 – A. elegantula sp. nov.; 28 – A. subtilis sp. nov. Scale = 2 mm.

Figs. 29-42. 29-35 – fore wing venation (female). 29 – Aclista insolita Nixon, 1957; 30 – A. vernalis sp. nov.; 31 – A. dubia Kieffer, 1909; 32 – A. longistilus Kieffer, 1909; 33 – A. marginalis Kieffer, 1909; 34 – A. elegantula sp. nov.; 35 – A. subtilis sp. nov. 36-42 – proximal part of antenna, lateral view (male). 36 – A. insolita; 37 – A. vernalis sp. nov.; 38 – A. dubia; 39 – A. longistilus; 40 – A. marginalis; 41 – A. elegantula sp. nov.; 42 – A. subtilis sp. nov. Scale = 1 mm (Figs. 29-35) and 0.7 mm (Figs. 36-42).